18(1) - Andean Botanical Information System
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18(1) - Andean Botanical Information System
ARNALDOA Revista del Museo de Historia Natural UNIVERSIDAD PRIVADA ANTENOR ORREGO El Museo de Historia Natural, es la unidad que conserva, educa, investiga y difunde los conocimientos que generan ȱ ȱ Çęȱ ȱ ȱ ȱ àȱ ¢ȱ àȱ ȱ ȱ ȱ àȱ ¢ȱ ȱ ȱ ȱÇǰȱȱȱȱȱȱȱǯ ISSN : 1815-‐‑8242 TRUJILLO REPÚBLICA DEL PERÚ 18(1) Enero -‐‑ Junio 2011 1 ,661 $UQDOGRD Floristic Checklist of the Peruvian Lomas Formations &DWiORJRÀRUtVWLFRGHODVLomas peruanas Michael O. Dillon Botany Department, The Field Museum, 1400 South Lake Shore Drive, Chicago, IL 60605, USA. PGLOORQ#¿HOGPXVHXPRUJ Segundo Leiva González & Mario Zapata Cruz Museo de Historia Natural, Universidad Privada Antenor Orrego, Casilla Postal 1075, Trujillo, PERÚ. VHJXQGRBOHLYD#KRWPDLOFRPP]DSDWDF#XSDRHGXSH Pedro Lezama Asencio Departamento Académico de Ciencias, Universidad Privada Antenor Orrego, Trujillo, PERU. SOH]DPDD# JPDLOFRP Victor Quipuscoa Silvestre Departamento Académico de Biología, Universidad Nacional de San Agustín, Arequipa, PERU. YTXLSXVFRDV#KRWPDLOFRP Abstract $ÀRULVWLFFKHFNOLVWRIWKH3HUXYLDQlomasIRUPDWLRQVLVSUHVHQWHGFRQVLVWLQJRIRQH*QHWRSK\WDJ\PQRVSHUP IDPLOLHVJHQHUDDQGVSHFLHVRI/LOLRSVLGDPRQRFRW\OHGRQVDQGIDPLOLHVJHQHUDDQGVSHFLHVRI 0DJQROLRSVLGDGLFRW\OHGRQVIRUDJUDQGWRWDORIIDPLOLHVJHQHUDDQGVSHFLHV7KHFKHFNOLVWLVEXWDVQDSVKRW RIWKLVÀRUDRXUVRXUFHVRIUHFRUGVLQFOXGHVPDQ\\HDUVRI¿HOGFROOHFWLQJUHFRUGLQJlomasSODQWVKHUEDULDLQ3HUXWKH 8QLWHG6WDWHVDQG(XURSH5HFRUGVDUHVXSSRUWHGZLWKYRXFKHUVRIZKLFKWKHPDMRULW\DUHDFFHVVLEOHYLDDVHDUFKDEOH GDWDEDVHDQGDVVRFLDWHGZLWKGLJLWL]HGKHUEDULXPLPDJHV$EULHIGHVFULSWLRQRIWKH3HUXYLDQlomasIRUPDWLRQVDUHYLHZ RIKLVWRULFDOFROOHFWLQJHIIRUWVDQGDGLVFXVVLRQRIIDFWRUVLQÀXHQFLQJlomasG\QDPLFVDUHSUHVHQWHG/RFDOLW\PDSVDQG DOLVWLQJRIlomasIRUPDWLRQVZLWKDSSUR[LPDWHFRRUGLQDWHVDUHSURYLGHG .H\ZRUGV3HUXFKHFNOLVWlomas formation. Resumen 6HSUHVHQWDXQFDWiORJRÀRULVWLFRGHODVlomasFRVWHUDVGHO3HU~TXHFRQVLVWHGHXQD*QHWRSK\WDJLPQRVSHUPD IDPLOLDVJHQHURV\HVSHFLHVGH/LOLRSVLGDPRQRFRWLOHGRQHDV\IDPLOLDVJHQHURV\HVSHFLHVGH 0DJQROLRSVLGDGLFRWLOHGRQHDVKDFLHQGRXQWRWDOGHIDPLOLDVJHQHURV\HVSHFLHV(OFDWiORJRFRQVWLWX\HXQ FRQVROLGDGRGHODÀRUDORPDOFX\DVIXHQWHVGHUHJLVWURVLQFOX\HQPXFKRVDxRVGHUHFROHFFLRQGHPXHVWUDVHQHOFDPSR DVLFRPRFRQVXOWDVHQKHUEDULRVGH3HU~(VWDGRV8QLGRV\(XURSD/RVUHJLVWURVHVWDQUHVSDOGDGRVSRUFROHFFLRQHVGH KHUEDULRGHODVFXDOHVODPD\RUtDVRQDFFHVLEOHVYLDEDVHVGHGDWRV\DVRFLDGDVFRQLPDJHQHVGHKHUEDULRGLJLWDOL]DGDV 6HSUHVHQWDWDPELHQXQDEUHYHGHVFULSFLRQGHODVLomasSHUXDQDVXQDUHYLVLRQKLVWyULFDGHORVHVIXHU]RVGHUHFROHFFLyQ \XQDGLVFXVLRQGHORVIDFWRUHVTXHLQÀX\HQHQODGLQiPLFDlomal$VLPLVPRVHSURSRUFLRQDQPDSDVGHORFDOLGDGHV\ XQDOLVWDGHIRUPDFLRQHVGHlomasFRQFRRUGHQDGDVDSUR[LPDGDV 3DODEUDVFODYH3HU~FDWiORJRIRUPDFLyQ lomal. Introduction )RU QHDUO\ NP DORQJ WKH ZHVWHUQ FRDVW RI 6RXWK$PHULFD >6 ODWLWXGH@ WKH 3HUXYLDQ DQG $WDFDPD GHVHUWV IRUP D FRQWLQXRXV K\SHUDULG EHOW EURNHQ RQO\ E\ RFFDVLRQDO ULYHUV YDOOH\V IURP WKH $QGHDQ &RUGLOOHUD 1DWLYH YHJHWDWLRQ RI WKH GHVHUWV FRQVLVWV RI RYHU VSHFLHV PDQ\ KLJKO\ HQGHPLF DQGODUJHO\UHVWULFWHGWRWKHIRJ]RQHORFDWLRQVRUlomas IRUPDWLRQV³VPDOOKLOOV´7KHÀRULVWLFFRPPXQLWLHVRI the lomas formations essentially function as terrestrial LVODQGVVHSDUDWHGE\K\SHDULGKDELWDWZKHUHYLUWXDOO\ QR SODQWV H[LVW ,Q 3HUX QR IHZHU WKDQ ORFDOLWLHV KDYHEHHQUHFRJQL]HGDVXQLTXHDVVHPEODJHVDOWKRXJK VRPHDUHQRZFRPSOHWHO\GHVWUR\HGGXHWRWKHDFWLRQV 7 Dillon ȱǯDZȱȱȱȱȱȱȱ 80W 75W 70W 5`S Desierto De Sechura Bayóvar 0 Chiclayo 100 200 300 Cerro Reque Cerro Cabezón Cerro Chiputur Cerro Virú Cerro Campana Trujillo BRAZIL Cerro Chimbote Casma 10`S Lupín Pativilca Barranca Lachay Iguanil Isla San Lorenzo Pasamayo Lima Lurin Atocongo Asia Isla San Gallán Amara 13`S PACIFIC OCEAN PERÚ Cañete Pisco Ica Jajuay San Nicolás Cháparra San Juan Arequipa Atico Lomas Ocoña Atiquipa Cachendo Chala Carranza Sama Grande Mollendo Mejía Ilo Ite Tacna Arica BOLIVIA )LJ/RFDOLWLHVRIFRDVWDO Lomas )RUPDWLRQVLQ3HUX$GDSWHGIURP'LOORQ RI H[SDQGLQJ KXPDQ SRSXODWLRQV 7KH SODQWV ZLWKLQ the lomas IRUPDWLRQV KDYH GLYHUVH RULJLQV LQFOXGLQJ DPSKLWURSLFGLVMXQFWVVHPLDULG(FXDGRULDQDQGFHQWUDO &KLOHDQVSHFLHVPRQWDQH$QGHDQGLVMXQFWVDQGPDQ\ lomasHQGHPLFV 7KH SK\WRJHRJUDSK\ DQG HFRORJ\ RI WKH GHVHUWV RI ZHVWHUQ 6RXWK$PHULFD KDYH EHHQ UHYLHZHG LQ GHWDLO5XQGHOHWDO:KLOHWKHGHVHUWLV FRQWLQXRXVIURP3HUXWR&KLOHWKHWRSRJUDSK\FOLPDWH DQG YHJHWDWLRQ RI HDFK GHVHUW LV GLVWLQFW$ULGLW\ LV FRQWUROOHG E\ WKUHH FOLPDWLF DQRPDOLHV 7KH ¿UVW DQ DEUXSWFOLPDWLFWUDQVLWLRQERWKWRWKHQRUWKDQGVRXWK UHVXOWLQJ LQ D SRRUO\ GHYHORSHG VWHSSH FOLPDWH DORQJ WKH PDUJLQV VHFRQG EULHI SHULRGV RI KHDY\ UDLQIDOO DQG UHODWLYHO\ KLJK WHPSHUDWXUHV DVVRFLDWHG ZLWK UDUH EXW UHFXUUHQW (O 1LxR HYHQWV VHH 'LOORQ 'LOORQ 5XQGHO RFFDVLRQDOO\ DIIHFW SDUWV RI WKH GHVHUW EULQJLQJ ZHW WURSLFDO FRQGLWLRQV DQG WKH WKLUGWKHUHPDUNDEOHWHPSHUDWXUHKRPRJHQHLW\DORQJ WKHHQWLUHODWLWXGLQDOH[WHQWRIWKHGHVHUWV7KLVSDWWHUQ 8 RI WHPSHUDWXUH VWDELOLW\ UHVXOWV IURP WKH LQÀXHQFH RI FRROVHDVXUIDFHWHPSHUDWXUHVDVVRFLDWHGZLWKWKHVRXWK WR QRUWK ÀRZ RI WKH +XPEROGW RU 3HUXYLDQ &XUUHQW $OVRLPSRUWDQWLVWKHLQÀXHQFHRIVWURQJDWPRVSKHULF VXEVLGHQFH DVVRFLDWHG ZLWK D SRVLWLRQDOO\ VWDEOH VXEWURSLFDODQWLF\FORQH7KHUHVXOWLVDPLOGXQLIRUP FRDVWDO FOLPDWH ZLWK WKH UHJXODU IRUPDWLRQ RI WKLFN VWUDWXV FORXG EDQNV EHORZ P GXULQJ WKH ZLQWHU PRQWKV :KHUH FRDVWDO WRSRJUDSK\ LV ORZ DQG ÀDW WKLVVWUDWXVOD\HUGLVVLSDWHVLQODQGZLWKOLWWOHELRORJLFDO LPSDFWEXWZKHUHLVRODWHGPRXQWDLQVRUVWHHSFRDVWDO VORSHVLQWHUFHSWWKHFORXGVDIRJ]RQHGHYHORSVZLWKD VWUDWXVOD\HUFRQFHQWUDWHGDJDLQVWWKHKLOOVLGHV7KHVH IRJV WHUPHG ³JDU~D´ DUH WKH NH\ WR WKH H[WHQW DQG GLYHUVLW\ RI YHJHWDWLRQ WKURXJKRXW WKH GHVHUWV RI WKH ZHVWHUQFRDVW:KLOHWKHH[WHQWRIWKH3HUXYLDQGHVHUW DFWXDOO\FRYHUVQHDUO\NPVLQFRDVWOLQHWKHDUHD FRYHUHGE\YHJHWDWLRQHYHQGXULQJH[FHSWLRQDO\HDUV LVOHVVWKDQVTNP $UQDOGRD Dillon ȱǯDZȱȱȱȱȱȱȱ no vegetation Tillandsia spp 1000 800 600 woody plants herbaceous perennials cryptogams no vegetation Tillandsia spp 400 Elevation(m) PERU 200 0 Fig. 2. Diagram illustrating interaction of fog and vegetation in the Lomas Formations. 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A. Lomas IRUPDWLRQV DW 3DFKDFDPDF 'HSDUWPHQW RI /LPD % Lomas Formations near Arequipa, Department of Arequipa. Fog clouds or “garua” can be seen laying on hillsides in the morning. $UQDOGRD 13 Dillon ȱǯDZȱȱȱȱȱȱȱ ,QWKHVWURQJ(O1LxR\HDURIERWK)UDQFLV: 3HQQHOO EG DQG ,YDQ 0XUUD\ -RKQVWRQ EG FROOHFWHG LQ WKH lomas formations of VRXWKHUQ 3HUX (ULFK :HUGHUPDQQ EG D ERWDQLVWZLWKWKH%HUOLQ'DKOP%RWDQLFDO0XVHXPDOVR FROOHFWHGLQVRXWKHUQ3HUXLQ$QDPDWHXUERWDQLVW IURP(QJODQG'RUD%6WDIIRUGPDGHDVHULHVRIFROOHFWLRQV IURP GHSDUWPHQWV LQ VRXWKHUQ 3HUX EHWZHHQ DQG FI 6WDIIRUG +HU HIIRUWV \LHOGHG PDQ\ collections from the lomas IRUPDWLRQVHVSHFLDOO\LQWKH 0ROOHQGR DUHD7 +DUSHU *RRGVSHHG EG IURPWKH8QLYHUVLW\RI&DOLIRUQLDDW%HUNHOH\LQLWLDWHGD VHULHVRIH[SHGLWLRQVWR3HUX*RRGVSHHGSXEOLVKHG DQDFFRXQWRIWKHH[SHGLWLRQVLQKLVERRN3ODQW+XQWHUV LQWKH$QGHV'XULQJDVWURQJ(O1LxR\HDURIKH YLVLWHGWKHLomasRI/DFKD\DQG/XUtQZLWK:HEHUEDXHU DQGGHVFULEHGWKHlomasIRUPDWLRQVDV³0HDGRZVRQWKH 'HVHUW´$GGLWLRQDOFROOHFWRUVLQKLVJURXSLQFOXGHG$$ %HHWOHEG+(6WRUNEG2 %+RUWRQ&5:RUWKE-/0RUULVRQE 5'0HWFDOI3&+XWFKLVRQEG-:HVW EGDQG-.:ULJKW 5DPyQ )HUUH\UD +XHUWD EG ZDV D VWXGHQWRI:HEHUEDXHUDQGSXEOLVKHGWKH¿UVWV\VWHPDWLF FRPSLODWLRQRISODQWVIRXQGZLWKLQWKHlomas formations )HUUH\UD1LFROiV$QJXOR(VSLQR $UQDOGR /ySH] 0LUDQGD EG DQG $EXQGLR 6DJiVWHJXL$OYD DOO KDYH FROOHFWHG ZLWKLQ the lomas formations of Peru. The latter collector PDGH H[WHQVLYH FROOHFWLRQV RQ &HUUR &DPSDQD DQG VXUURXQGLQJDUHDVGXULQJWKHVWURQJ(O1LxR\HDUVRI 7KRVHFROOHFWLRQVSURYLGHGWKHEDVLVIRUDQ LQYHQWRU\RIWKHUHJLRQVlomasIRUPDWLRQV6DJiVWHJXL HW DO 0LNLR 2QR RI WKH7RN\R 0HWURSROLWDQ 8QLYHUVLW\ -DSDQ GLUHFWHG D VHULHV RI H[SHGLWLRQV WR FRDVWDO 3HUX DQG &KLOH LQ WKH ¶V +H DQG KLV FROOHDJXHVFRQGXFWHGDZLGHYDULHW\RILQYHVWLJDWLRQV ZLWKLQ WKH lomas IRUPDWLRQV LQFOXGLQJ FRXQWLQJ chromosomes of lomas VSHFLHV FDOFXODWLQJ ELRPDVV DQGGRFXPHQWLQJVHHGEDQNV2KJD 2ND2JDZD2QR %HJLQQLQJLQWKHVWURQJ(O1LxR\HDURI02 'LOORQ EHJDQ FROOHFWLQJ ZLWKLQ WKH lomas formations ZKLFKFXOPLQDWHGLQVHYHUDOWKRXVDQGVRIDFFHVVLRQVDQG DVHULHVRISDSHUVGHVFULELQJWKHYHJHWDWLRQRIFRDVWDO3HUX DQG&KLOHVHH'LOORQ'LOORQ )LJLomas formation near Atiquipa, Department of Arequipa. 14 $UQDOGRD Dillon ȱǯDZȱȱȱȱȱȱȱ 5XQGHO5XQGHOHWDO$ÀRULVWLF GDWDEDVHFRYHULQJWKHHQWLUHDUHDLVDYDLODEOHVHH'LOORQ &ROOHFWLQJH[SHGLWLRQVZHUHFRQGXFWHGGXULQJDQG DIWHUVWURQJ(O1LxR\HDUVRI DQG7KHVHHIIRUWVKDYH\LHOGHGVHYHUDOWD[DQHZ to science (Dillon HWDO(OLVHQV*yPH]6RVD .UDSRYLFNDV5RELQVRQ0RRUH ELRJHRJUDSKLFDOTXHVWLRQVLQYROYLQJSODQWVRIWKHlomas IRUPDWLRQ IURP DQ H[SOLFLW SK\ORJHQHWLF IUDPHZRUN (Dillon HW DO *HQJOHU1RZDN +XHUWDV HW al/XHEHUW:HQ0RRUH-DQVHQ 6LPSVRQHWDO 6HYHUDO VWXGLHV KDYH FRQWULEXWHG WR WKH JHQHUDO NQRZOHGJHDQGGHVFULSWLRQRIWKH3HUXYLDQ'HVHUWDQG lomasIRUPDWLRQVHJ'LOORQ)HUUH\UD /HLYDHWDO5XQGHOHWDODQG VRPHKDYHDGGUHVVHGELRJHRJUDSKLFTXHVWLRQVIURPWKH SRLQWRIYLHZRILWVÀRUD*DOiQGH0HUDHWDO /H]DPD'LOORQLQSUHS0DVX]DZD5XQGHO 'LOORQ 'LOORQ 3LQWR /XHEHUW 2WKHUDXWKRUVKDYHVWXGLHVWKHFRPPXQLW\HFRORJ\DQG SURGXFWLYLW\RIUHJLRQDOIRUPDWLRQVFI-LPpQH]HWDO. 3pIDXU$IHZVWXGLHVKDYHDSSURDFKHG LomasFRPPXQLWLHVOLNHDQ\RWKHUSODQWIRUPDWLRQ DUH QRW VWDWLF UDWKHU WKHLU FRPSRVLWLRQ LV LQ FRQVWDQW FKDQJHZKLOHVRPHVSHFLHVDUHFRQVWDQWRWKHUVFRPH DQG JR ZLWK HDFK VHDVRQ 'XULQJ DQG DIWHU VWURQJ (O 1LxRHYHQWVUDUHVSHFLHVDUHUHFRUGHGWKDWPD\QRWEH HQFRXQWHUHGLQLQWHUYHQLQJ\HDUV7KHOLVWLQJRISODQWV ZHSUHVHQWLVDORQJWHUPFRPSLODWLRQGUDZQIURP\HDUV RIREVHUYDWLRQDQGKHUEDULXPVWXG\&XUUHQWHVWLPDWHV IRUWKHWRWDOQXPEHURIVSHFLHVUHSUHVHQWHGZLWKLQWKH Peruvian lomasIRUPDWLRQVFRQVLVWVRIIDPLOLHV JHQHUDDQGFDVSHFLHV Overall Diversity Table 1. Families represented within the Lomas Formations of Peru with high diversity at the generic or species level. Families %URPHOLDFHDH 3RDFHDH Asteraceae %RUDJLQDFHDH %UDVVLFDFHDH &DFWDFHDH )DEDFHDH 0DOYDFHDH 6RODQDFHDH Calceolariaceae &\SHUDFHDH $PDUDQWKDFHDH 2[DOLGDFHDH 3LSHUDFHDH $L]RDFHDH &RQYROYXODFHDH Santalaceae Genera >JHQHUDVSS@7LOODQGVLDVSS >JHQHUDVSS@(UDJURVWLVVSS3DVSDOXPVSS >JHQHUDVSS@2SK\URVSRUXVVSS6HQHFLRVSS >JHQHUDVSS@+HOLRWURSLXPVSS7LTXLOLDVSS >JHQHUDVSS@ >JHQHUDVSS@1HRSRUWHULDVSS >JHQHUDVSS@ >JHQHUDVSS@3DODXDVSS >JHQHUDVSS@1RODQDVSS6RODQXPVSS &DOFHRODULDVSS >JHQHUDVSS@&\SHUXVVSS >JHQHUDVSS@$OWHUQDQWKHUDVSS 2[DOLVVSS 3HSHURPLDVSS >JHQHUDVSS@7HWUDJRQLDVSS >JHQHUDVSS@,SRPRHDVSS 4XLQFKDPDOLXPVSS Acknowledgements :H ZLVK WR WKDQN WKH &XUDWRUV DQG &ROOHFWLRQ 0DQDJHUV DW DOO KHUEDULD YLVLWHG LQFOXGLQJ EXW QRW UHVWULFWHG WR % %0 &*( &21& &8= ) ), $UQDOGRD *+ +$2 +86$ +87 . 0 0$ 0$. 02 1< 6*2 8& 86 DQG 860 7KH FROOHFWLRQ RI VSHFLPHQODEHOGDWDZDVVXSSRUWHGLQSDUWE\JUDQWV WR02'IURPWKH1DWLRQDO6FLHQFH)RXQGDWLRQ'(% '(%%65DQG1DWLRQDO 15 Dillon ȱǯDZȱȱȱȱȱȱȱ *HRJUDSKLF 6RFLHW\ 7KH SURIHVVLRQDO VWDII DW WKH )LHOG0XVHXP¶V%RWDQ\'HSDUWPHQWLVDFNQRZOHGJHG IRUWKHLUVXSSRUWRYHUWKHODVW\HDUVDQGLQFOXGHV &KULVWLQH1LH]JRGD/XFLD.DZDVDNL6XVDQ+DPQLN $QQD %DOOD $UOHQH 6LVRQ .U\VWD 'XUDQ 0DU\ *XVWDIVRQ -RDQQD 0F&DIIUH\ :DOW 5HLVOLQJ DQG 1RUWKZHVWHUQ 8QLYHUVLW\ LQWHUQ (OL]DEHWK$OOHQ :H WKDQN'DUOHQH'RZG\3ULWFKHWWIRUWKHH[FHOOHQWMRERI PRXQWLQJDOOWKH)PDWHULDOFROOHFWHGGXULQJPDQ\¿HOG VHDVRQV0DUN$OYH\%RWDQ\'HSDUWPHQW2SHUDWLRQDO 0DQDJHU LV DFNQRZOHGJHG IRU EULGJLQJ EXUHDXFUDWLF VXSSRUWDQGIXQGLQJGRFXPHQWDWLRQ(GXDUGR7LQD 0DUWHOO DUH WKDQNHG IRU ORJLVWLFDO DQG WUDQVSRUWDWLRQ VXSSRUWGXULQJVHYHUDOH[SHGLWLRQV'LDQH2¶'RQQHOO 'LOORQLVDFNQRZOHGJHGIRUKHUKHOSGXULQJWKH ¿HOG VHDVRQ JDWKHULQJ YRXFKHUV DQG SROOLQDWRUV 7KH IROORZLQJLQGLYLGXDOVKDYHKHOSHGLQYDULRXVDVSHFWVRI RXUVWXGLHV3DXO=DKQOH$OEHUWR%HQDYLGHV*DQR]D 0DUXFKD%HQDYLGHVGH7VFKXGL0DXULFLRGH5RPDxD (GXDUGR 0HMLD 3LQWR (GJDUGR 2UWL] 9DOHQFLD -XQ :HQ )UHG %DUULH 7RP 'XQFDQ )HGHULFR /XHEHUW 2OLYHU:KDOOH\0LNLR2QR0L\XNL1DND]DZD)XMLWD ,VLGRUR6iQFKH]9HJD-RVp6DQWLVWHEDQ&DVWLOOR&DPLOR 'tD] 6DQWLVWHEDQ (OLGD &DUULOOR )XHQWHV 0DJGD &KDQFR (VWHOD 5DPyQ )HUUH\UD +XHUWD (OR\ /ySH]/DXUD&iFHUHV0DVVLHO&RUUDOHV*LQD&DVWLOOR 'DQLHO5DPRV0DUFR&XHYD.iURO'XUDQG1DWKDOLH &DVWUR -XDQ *RQ]DOHV (ULF 5RGULJXH] 5RGULJXH] $EXQGLR6DJiVWHJXL$OYDDQGWKH:RPHQ¶V%RDUG ³)LHOG 'UHDPV´ JLIW SURJUDP 6/* ZLVKHV WR WKDQN 83$2 IRU SHUPLVVLRQ FRQGXFW ¿HOG VWXGLHV DQG IRU FRQWLQXLQJVXSSRUWIRU$UQDOGRD946ZLVKHVWRWKDQN WKH'HSDUWDPHQWRGH%LRORJtD816$IRUSHUPLVVLRQ WR FRQGXFW ¿HOG VWXGLHV DQG WKH VWXGHQW PHPEHUV RI ',%,26 *UXSR GH ,QYHVWLJDFLyQ GH OD 'LYHUVLGDG %LROyJLFDGHO6XUIRUWKHLU¿HOGDQGKHUEDULXPZRUN :HDOVRZLVKWRDFNQRZOHGJHDQGWKDQNWKH%LRGLYHUVLW\ +HULWDJH /LEUDU\ DQG WKH 0LVVRXUL %RWDQLFDO *DUGHQ IRUSURYLGLQJGLJLWDODFFHVVWR5XL]3DYyQ¶V)ORUD 3HUXYLDQD HW &KLOHQVLV ZZZELRGLYHUVLW\OLEUDU\ RUJ/DVWEXWFHUWDLQO\QRWOHDVWZHWKDQNWKHPDQ\ WD[RQRPLFH[SHUWVWKDWSURYLGHGGHWHUPLQDWLRQVRYHUWKH ODVWIHZGHFDGHV'R]HQVRIVFLHQWLVWVKDYHKHOSHGZLWK LGHQWL¿FDWLRQVEXW1DQF\+HQVROGGHVHUYHVDVSHFLDO Fig. 6. Lomas formations near the Lagunas de Mejia in 1983, Department of Arequipa. 16 $UQDOGRD Dillon ȱǯDZȱȱȱȱȱȱȱ PHQWLRQ IRU KHU HIIRUWV DW LGHQWL¿QJ DQG DQQRWDWLQJ FROOHFWLRQV 7KH DXWKRUV DFFHSW DQ\ DQG DOO HUURUV RPLVVLRQVRUPLVDSSOLFDWLRQRIQDPHVLQWKHOLVW Literature cited Anderson, E. F. 7KHFDFWXVIDPLO\7LPEHU3UHVV3RUWODQG Oregon, USA. Barthlott , W. , and D. R. Hunt. 1993 . Cactaceae. ,Q K. Kubitzki [ed.], The families and genera of vascular plants. Springer-Verlag, New York, New York, USA. Beresford-Jones, D.G.&KDSWHU7KH*HQXV3URVRSLV RQWKH6RXWK&RDVW3S,Q Pre- Hispanic 3URVRSLV- Human Relationships on the South Coast of Peru: Riparian Forests in the Context of Environmental and Cultural Trajectories of the Lower Ica Valley. PhD Dissertation, Magdalene College, University of Cambridge, UK. Brako, L. & J. L. Zarucchi.&DWDORJXHRIWKHÀRZHULQJ plants & gymnosperms of Peru. (Monographs in Systematic %RWDQ\ 9RO 0LVVRXUL %RWDQLFDO *DUGHQ 6W /RXLV 02 1286 pp. Dillon, M. O.7KH%RWDQLFDO5HVSRQVHRIWKH$QGHDQ'HVHUW Lomas Formations to the 1982-83 El Niño Event. Abstract Amer. -%RW Dillon, M. O. 1989. Origins and diversity of the lomas formations in the Atacama and Peruvian Deserts of western South America. $EVWUDFW$PHU-%RW Dillon, M. O. 1994. Andean Botanical Information System/Sistema de Información Botánica Andina. URL:http://www.sacha.org/, DFFHVVHG1RYHPEHU Dillon, M.O., T. Tu, A. Soejima, T. Yi, Z. Nie, A. Tye, & J. Wen.3K\ORJHQ\RI1RODQD (Nolaneae, Solanoideae, Solanaceae) as inferred from granule-bound starch synthase I *%66,VHTXHQFHV7D[RQ Dillon, M.O., T. Tu, L. Xie, V. Quipuscoa S., & J. Wen. %LRJHRJUDSKLFGLYHUVL¿FDWLRQLQ1RODQD (Solanaceae), a ubiquitous member of the Atacama and Peruvian Deserts along the western coast of South America. Journal of Systematics & (YROXWLRQ Duncan, T. & M.O. Dillon. 1991. Numerical analysis of the ÀRULVWLF UHODWLRQVKLSV RI WKH ORPDV RI 3HUX DQG &KLOH$EVWU Abstract Amer. J. Bot. 78: 183. Elisens, W.J. *HQHWLF GLYHUJHQFH LQ *DOYH]LD (Scrophulariaceae): evolutionary and biogeographic UHODWLRQVKLSVDPRQJ6RXWK$PHULFDQDQG*DOiSDJRVVSHFLHV $PHULFDQ-RXUQDORI%RWDQ\ Ferreyra, R. &RPXQLGDGHV YHJHWDOHV GH DOJXQDV ORPDV FRVWDQHUDVGHO3HU~(VWDF([S$JULFROD/D0ROLQD%RO Ferreyra, R. 1961. Las Lomas Costaneras del extremo sur del 3HU~%ROHWtQGHOD6RFLHGDG$UJHQWLQDGH%RWiQLFD Ferreyra, R. 1983. Los tipos de vegetación de la costa peruana. $QDOHV-DUG%RW0DGULG Fryxell, P. A. 1996. )XHUWHVLPDOYD, a new genus of neotropical Malvaceae. Sida 17(1): 69–76. Galán de Mera, A., J.A. Vicente-Orellana, J.A. Lucas García & A. Probanza Lobo. 1997. Phytogeographical VHFWRULQJRIWKH3HUXYLDQFRDVW*OREDO(FRO%LRJHRJU/HWW *HQJOHU1RZDN . 0 5HFRQVWUXFWLRQ RI WKH biogeographical history of Malesherbiaceae. Botanical Review 68(1): 171–188. Dillon, M.O./RPDV)RUPDWLRQV3HUXSS,Q S. D Davis, V. H. Heywood, O. Herrera- McBryde, J. Villa-Lobos DQG$&+DPLOWRQHGV&HQWUHVRI3ODQW'LYHUVLW\$*XLGH and Strategy for their Conservation. WWF, Information Press, Oxford, U.K. Gómez-Sosa, E. 1986. $VWUDJDOXVQHREDUQHE\DQXV (Leguminosae): A New Species from Peru. Brittonia 38(4): 427- 429. Dillon, M.O. 6RODQDFHDH RI WKH /RPDV IRUPDWLRQV RI &RDVWDO3HUXDQG&KLOH3S,Q: Hollowell, V., Keating, T., Lewis, W. & Croat, T. (eds.), $)HVWVFKULIWIRU:LOOLDP* '¶$UF\7KH/HJDF\RID7D[RQRPLVW. Monogr. Syst. Bot. $QQ0LVVRXUL%RW*DUG Huertas, M. L., J. V. Schneider, & G. Zizka. Phylogenetic Analysis of 3DODXD (Malveae, Malvaceae) based RQ3ODVWLGDQG1XFOHDU6HTXHQFHV6\V%RW Dillon, M. O.)ORUDRIWKH/RPDV)RUPDWLRQV85/KWWS HPXZHE¿HOGPXVHXPRUJERWDQ\VHDUFKBORPDVSKSDFFHVVHG -DQ Jiménez, P., F. Villasante, C. Talavera, L. Villegas, E. Huaman, & A. Ortega. 1998. Southern Peru Loma’s )ORUD3S,Q: Schemenauer, R. (ed). Proceedings of the First International Conference on Fog and Fog Collection, 19-24 July 1998, Vancouver, Canada. pp. 492. Dillon, M.O. & P. W. Rundel. 7KHERWDQLFDOUHVSRQVHRI the Atacama and Peruvian Desert Flora to the 1982-83 El Niño HYHQW3S,Q3:*O\QQHG*OREDO(FRORJLFDO Consequences of the 1982-83 El Niño- Southern Oscillation, Elsevier Science Publishers, New York. Dillon, M.O. , S. Leiva, & V. Quipuscoa. )LYHQHZ species of 1RODQD (Solanaceae-Nolaneae) from Peru and notes RQWKHFODVVL¿FDWLRQIRUDGGLWLRQDOWD[D$UQDOGRD $UQDOGRD Goodspeed, T.H. 1961 [1941]. Plant Hunting In The Andes. Berkeley: University of California Press, 429 pp. Hunt , D. , N. P. Taylor , & G. Charles7KHQHZ cactus lexicon. dh Books, Milborne Port, UK. Jiménez, P.M., C. Talavera, L. Villegas, F. Villasante, A. Ortega, & M. Jiménez. %,$WLTXLSD ,VOD de Biodiversidad en el Desierto Costero Perúano. II Congreso ,QWHUQDFLRQDOGH&LHQWt¿FRV3HUXDQRV/LPD$EVWUDFWS Krapovickas, A. 1996. Sinopsis del género *D\D (Malvaceae). %RQSODQGLD 17 Dillon ȱǯDZȱȱȱȱȱȱȱ Lanning, E. P.(DUO\0DQLQ3HUX6FLHQWL¿F$PHULFDQ of Southern Peru. Vegetatio 49: 163 - 171. León B., A. Cano, & K. R. Young./RVKHOHFKRVGHODV lomas costeras del Perú / Ferns of the fog vegetation of coastal Peru. Arnaldoa 9(2): 7-41. Pinto, R., & F. Luebert. 'DWRVVREUHODÀRUDYDVFXODU del desierto costero de Arica y Tarapacá, Chile, y sus relaciones ¿WRJHRJUi¿FDVFRQHOVXUGH3HU~*D\DQD%RWiQLFD± Leiva-G.,S., M. Zapata C., G. Gayoso B., P. Lezama A., V. Quipuscoa S., & M.O. Dillon.'LYHUVLGDG ÀRUtVWLFDGHOD/RPD0RQJyQ3URYLQFLD&DVPD'HSDUWDPHQWR Ancash, Perú - Floristic diversity of Loma Mongon, Casma 3URYLQFH$QFDVK'HSDUWPHQW3HUX$UQDOGRD Price, R. A. 6\VWHPDWLFV RI WKH *QHWDOHV D UHYLHZ RI morphological and molecular evidence. International Journal of Plant 6FLHQFHV6XSS%LRORJ\DQG(YROXWLRQRI*QHWDOHV66 Lezama-A, P. & M.O. Dillon.1XPHULFDO$QDO\VLVRIWKH Peruvian Lomas Formations. Arnaldoa: In Press. Luebert, F. & J. Wen. 3K\ORJHQHWLF$QDO\VLV DQG (YROXWLRQDU\'LYHUVL¿FDWLRQRI+HOLRWURSLXP Sect. &RFKUDQHD (Heliotropiaceae) in the Atacama Desert. Systematic Botany ± Masuzawa, T. 1986. Structure of 7LOODQGVLD lomas community in Peru coast. In: Taxonomic and ecological studies on the lomas YHJHWDWLRQ LQ WKH 3DFL¿F FRDVW RI 3HUX *HRJUDSKLFDO Reports of Tokyo Metropolitan University. Tokyo. No. 19. Moore M. J. & R. K. Jansen.0ROHFXODUHYLGHQFHIRU the age, origin, and evolutionary history of the American desert plant genus 7LTXLOLD (Boraginaceae) Molecular Phylogenetics and Evolution. 39: 668-687. Ohga, N. 1986. Dynamics of the buried seed population in soil, and the mechanisms of maintenance of the herbaceous lomas vegetation in the coastal desert of central Peru. ,Q Ono, M. (ed.), Taxonomic and ecological studies on the lomas vegetation in WKH3DFL¿FFRDVWRI3HUX0DNLQR+HUEDULXP7RN\R0HWURSRO 8QLYHUVLW\7RN\R3S Ohga, N. 1991. Distribution Patterns of Buried Seeds in the Herbaceous Lomas Community over the Entire Plateau on Loma Ancon in the Coastal Desert of Central Peru. Journal of $ULG/DQG6WXGLHV Ohga, N. 1992. Buried seed population in the herbaceous lomas on Loma Ancon in the coastal desert of central Peru. Ecological 5HVHDUFK Oka, S. & H. Ogawa. 1984. The distribution of lomas vegetation DQGLWVFOLPDWLFHQYLURQPHQWVDORQJWKHSDFL¿FFRDVWRI3HU~ ,Q *HRJUDSKLFDO 5HSRUWV RI 7RN\R 0HWURSROLWDQ University. Tokyo. No. 19. Olmstead, R.G, C.W. Depamphilis, A.D. Wolfe, N.D. Young, W.J. Elisons, & P. A. Reeves.'LVLQWHJUDWLRQ of the Scrophulariaceae. Amer. J. Bot. 88(2): 348–361. Ono, M. 1982. A Preliminary Report of Taxonomic and Ecological 6WXGLHVRQWKH/RPDV9HJHWDWLRQLQWKH3DFL¿F&RDVWRI3HUX In: M. Ono (ed.). Makino Herbarium. Tokyo Metropolitan University. Ono, M. 1986. Taxonomic and Ecological Studies on the Lomas 9HJHWDWLRQLQWKH3DFL¿F&RDVWRI3HUX3S,Q M. Ono (ed.). Makino Herbarium. Tokyo Metropolitan University. Péfaur, J. E. 1982. Dynamics of plant communities in the Lomas 18 Raimondi, A.(O3HU~LWLQHUDULRGHYLDMHVYHUVLyQOLWHUDOGH las libretas originales) Lima. Imprenta Torres Aguirre. Robinson, H. & A. J. Moore.1HZVSHFLHVDQGQHZ combinations in 5K\VROHSLV (Heliantheae: Asteraceae) Proc. Biol. Soc. Wash. 117(3): 423-446. Ruiz-López, H. & J. Pavón. 1799. Flora Peruviana, et Chilensis, sive descriptiones, et icones Plantarum Peruvianarum, et Chilensium, secundum systema Linnaeanum digestae, cum characteribus plurium generum evulgatorum reformatis. v. 2: 6-8, plates CXII, CXIII. Rundel, P.W., M.O. Dillon, H. A. Mooney, S.L. Gulmon, & J.R. Ehleringer. 1991. The phytogeography and ecology RIWKHFRDVWDO$WDFDPDDQG3HUXYLDQ'HVHUWV$OLVR Rundel, P. & M.O. Dillon. 1998. Ecological patterns in the Bromeliaceae of the lomas formations of Coastal Chile and Peru. Plant Syst. Evo. 212: 261-278. Rundel , P.W., P.E Villagra, M.O. Dillon, S. Roig- Juñent and G. Debandi.&KDSWHU$ULGDQGVHPL DULGHFRV\VWHPV3S,Q Veblen, T.T., K. R. Young & $52UPHHGV7KH3K\VLFDO*HRJUDSK\RI6RXWK$PHULFD Oxford University Press: Oxford, UK, 368 pp. Sagástegui, A., J. Mostacero, & S. López. 1988. Fitoecología del Cerro Campana. Bol. Soc. Bot. La Libertad 14: 1-47. Simpson, B. B., Tate, J. A. and Weeks, A. The biogeography of +RIIPDQQVHJJLD (Leguminosae, Caesalpinioideae, Caesalpinieae): a tale of many travels. -RXUQDORI%LRJHRJUDSK\± Steele, A. R. 1982. Flowers for the King: The expedition of Ruiz and Pavón and the Flora of Peru. Stafford, D. 1939. On the Flora of Southern Peru. Proc. Linnean 6RF/RQGRQ Tryon, R.7KH(FRORJ\RI3HUXYLDQ)HUQV$PHULFDQ)HUQ -RXUQDO Vermeij, G.J. $Q HFRORJLFDO FULVLV LQ DQ HYROXWLRQDU\ FRQWH[W(O1LxRLQWKHHDVWHUQ3DFL¿F3S±,Q P.W. *O\QQHG*OREDO(FRORJLFDO&RQVHTXHQFHVRIWKH± El Niño- Southern Oscillation. Amsterdam: Elsevier. Weberbauer, A. /D LQÀXHQFLD GH FDPELRV FOLPiWLFRV \ geológicos sobre la vegetación de la costa peruana. Academia 1DFLRQDOGH&LHQFLDV([DFWDV)tVLFDV\1DWXUDOHV Weberbauer, A. (O 0XQGR 9HJHWDO GH ORV $QGHV 3HUXDQRV (VWXGLR )LWRJHRJUi¿FR (VWDFLyQ ([SHULPHQWDO Agrícola de La Molina. Ministerio de Agricultura, Lima. 776 pp. $UQDOGRD Dillon ȱǯDZȱȱȱȱȱȱȱ A B Fig. 7. Lomas Formations, A. 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