Catalogue of the spiders (Arachnida Araneae

Transcription

Biodiversity Journal, 2013, 4 (1): 3-104
M ONOGRAPH
Catalogue of the spiders (Arachnida Araneae) of Sardinia
Paolo Pantini1*, Antonio Sassu2 & Giuseppe Serra3
Museo civico di Scienze Naturali “E. Caffi”. Piazza Cittadella, 10 - 24129, Bergamo (BG), Italy; e-mail: ppantini@comune.bg.it
Via Europa, 59 - 09070, Bonarcado (OR), Italy; e-mail: a.sassu77@gmail.com
3
CNR, Istituto per lo Studio degli Ecosistemi - sede di Sassari. Traversa La Crucca, 3 - 07100, Sassari-Li Punti (SS), Italy; e-mail:
g.serra@ise.cnr.it
*
Corresponding author
1
2
ABSTRACT
KEY WORDS
This catalogue summarizes and critically reviews araneological knowledge about Sardinian
fauna. 140 publications from 1868 to 2012 were considered and about 4800 specimens belonging to 254 species were directly examined. The list report 495 species belonging to 229
genera divided into 43 families. For each species, literature, chorotype, regional distribution,
and, where possible, habitats are indicated. Among the listed species Philodromus bosmansi
Muster et Thaler, 2004 (Philodromidae) is new for Europe and the family of Prodidomidae
with the species Zimirina brevipes Pérez et Blasco, 1986 is new for Italy. Also Setaphis parvula (Lucas, 1846) (Gnaphosidae), Centromerus succinus (Simon, 1884) and Hybocoptus
corrugis (O. P.-Cambridge, 1875) (Linyphiidae) are new records for the Italian fauna and
further 37 species are new for Sardinia. The genitalia of Phrurolithus corsicus (Simon, 1878),
Zimirina brevipes, Drassodes luteomicans (Simon, 1878) and Zelotes dentatidens Simon,
1914 are illustrated.
biodiversity; distribution; endemic species.
Received 05.04.2012; accepted 30.11.2012; printed 30.03.2013
INTRODUCTION
The knowledge of biodiversity of an area means
to how much and what taxa live in it. This information is crucial in order to realize an appropriate protection and management of the territory or also
further in-depth ecological and biological research.
Italy was the first country in Europe to organize
a census of its national fauna publishing a Checklist
between 1993 and 1995 (Minelli, Ruffo & La Posta,
1993-1995). With the aim of better quantifying the
faunistic resources of our country and to check their
distribution throughout the territory, between 2001
and 2005 following the CKmap project in which the
specialists compiled three tables for each species:
1) a species-index card including taxonomic, chorological and ecological data such as habitat, food,
conservation status and indicator value; 2) a distribution-table including all species records with coordinates added by a simplified software method,
localities were automatically geo-referenced in the
UTM grid (10 x 10 km grid cells); 3) a referencetable storing the list of literature sources and collections examined. (Ruffo & Stoch, 2005; Letardi
& Pantaleoni, 2007).
Despite this effort, our knowledge of Italian
fauna is still far from complete, especially if we
consider invertebrates (Ruffo & Vigna Taglianti,
2002). Among these, spiders are a particularly interesting group, rich in species, many of which are
endemic. Information about the ecology and distribution of Araneae in Italy is still sketchy and fragmentary, often limited to some families, spread
across a large number of publications, few of which
4
P. Pantini, A. Sassu & G. Serra
give us a unified picture of the situation in smaller
or larger areas of our peninsula. A recent analysis
of the state of knowledge on Italian araneofauna
(Pantini, in press) demonstrated how much effort is
still needed to reach a satisfactory level. From 1995
to 2012, 238 scientific publications with data regarding Italian spider fauna was published. Overall
1200 species are mentioned, 229 of which were new
for Italy. This means an increase of 17,4% compared
to the Checklist (Pesarini, 1994), bringing the number of Italian spiders above 1600. In this period of
time, 61 new species have been described from
specimens coming from Italy, among which, as far
as we know, 52 are endemic or sub-endemic. The
publications examined can be classified into 5 different groups: faunal (40%), taxonomic (26%), ecological (17%), bio-speleological (10%) and various
(7%). Regarding faunal aspects, some works
must be mentioned which analyze Italian spider
fauna, both as a whole (Trotta, 2005; Pantini & Isaia,
2008) and in some regions such as Alto Adige (Noflatscher, 1996), Valle d’Aosta (Isaia, 2005), Piedmont and Lombardy (Isaia et al., 2007) and Veneto
(Ballarin et al., 2011). However, a great part of the
southern regions of our country are still almost unexplored, or at least little is known about them.
With the publication of this catalogue, we
summarize and critically review the araneological
knowledge regarding Sardinia, an extremely interesting Italian region from a faunal and bio-geographical point of view, grouping into one text a large
amount of data from literature and from an analysis
of a vast quantity of new material.
ARANEOLOGICAL RESEARCH IN SARDINIA
In 1868, Giovanni Canestrini and Pietro Pavesi
published “Araneidi italiani”, the first, and even
now, only catalogue of Italian spiders. The two authors, who we can consider the fathers of Italian
arachnology, introduce the work with a long and learned analysis of the literature and authors who,
from Pliny the Elder onwards, had anything to do
with Italian araneofauna. There then follows the actual catalogue which reports for every species observed its regional distribution (based mostly on
material collected or studied by the authors) and its
bibliographical reference. The catalogue reports
only three species in Sardinia: Argiope bruennichi
(Scopoli, 1772), Latrodectus tredecimguttatus
(Rossi, 1790), and Eresus ruficapillus C.L. Koch,
1846. Even in the updated version in 1870, the data
about Sardinia does not change, while on the whole,
data referring to 41 families and 327 species of spider was reported. The records, as we can see from
the map in figure 1, were concentrated mostly in the
north of Italy where the two authors lived and worked. At the time Canestrini (born and raised in Val
di Non, Trento) was a Professor at the University
of Modena while Pavesi (born in Pavia) was a teacher at Lugano High School. Sardinia and most of
southern Italy were much less known except Campania (where Prof. Panceri worked) and Sicily.
However, the first studies on Sardinian araneofauna, with the description of Zelotes sardus (sub
Prothesima sarda) a new species of Gnaphosidae,
were made by Canestrini (1873; 1876). Later Pavesi
(1876; 1878) published two papers on the collections deriving from the cruises of the “Violante”, he
listed a total of 22 species mostly from the small islands surrounding Sardinia. Successively Magretti
(1878; 1880) published accounts of two zoological
excursions in Sardinia, in which he cites 18 species
of spiders (Table 1) while De Carlini (1885) in his “Artropodi dell’Isola di San Pietro” reports 19 species.
But the first consistent set of data is due to the naturalistic Sardinian explorations of Achille Costa
(1882; 1983; 1984; 1985; 1986) who reports 135
species in his “Notizie ed osservazioni sulla geofauna sarda” (Table 1).
In the first years of the 20th century the most
complete contribution on Sardinian spiders was
published by Garneri (1902). In his “Contribuzione
alla fauna sarda”, the author reports a long list of
species, as many as 235, arising partly from bibliographical research and partly from material from Sardinia preserved in the University of Pavia. Fifty years
follow with a complete absence of publications about
Sardinian fauna, only some works by Gozo (1908)
and by Simon (1914; 1926; 1929; 1932) report sparse
data. In the second half of the 20th century there was
new interest: Roewer (1953, 1956) published on Sardinian cave-dwelling arachnids and Krauss (1955)
on the spiders of Corsica, Sardinia and Elba. The latter reports 51 species, of which 16 are new to the island, while Dresco (1956) described Tegenaria
henroti. The work of Paolo Marcello Brignoli deserves, as usual, special attention. He published as
many as 22 works in which he reports a large
5
Figure 1. Number of spider species reported in Italy by Canestrini & Pavesi (1870) divided into regions.
amount of original data about Sardinia. In particular, we remember his contributions on Onoopidae
(Brignoli, 1967a), on Leptonetidae (Brignoli,
1967b; 1969) and on cave dwellers (Brignoli,
1974), dedicated specifically to Sardinian fauna.
From the 1970s on, bio-speleological research carried out on the island also led to the publication of
several works. In particular, regarding spider fauna,
we should mention the contributions of Thermes
(1972) and Puddu & Pirotta (1974) with the “Catalogo sistematico ragionato della fauna cavernicola
della Sardegna”. Another important contribution is
that of Giuseppe Grafitti, a tireless researcher and
great expert on the hypogean fauna of the island
(see Table 1).
Papers on Sardinian spiders over the last twenty
years are relatively scarce and is almost exclusively
the result of faunal or taxonomic contributions:
6
Citations
1
Canestrini & Pavesi, 1868
2b
Canestrini, 1873
2a
2c
3
4
5
6
7a
7b
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
Species New
cited species
(n.)
(n.)
3
3
3
2
Pavesi, 1876
16
15
Pavesi, 1878
9
7
Canestrini & Pavesi, 1870
Canestrini, 1876
Magretti, 1878
3
5
1
2
Magretti, 1880
17
11
Costa, 1883
1
1
Costa, 1882
8
3
Costa, 1884
56
38
Costa, 1885
73
65
Costa, 1886
3
3
Simon, 1884
de Carlini, 1885
2
18
2
2
Garneri, 1902
229
107
Simon, 1914
4
2
Gozo, 1908
Simon, 1926
Simon, 1929
Simon, 1932
4
1
2
2
5
3
Krauss, 1955
50
16
Roewer, 1956
5
2
Roewer, 1953
Dresco, 1956
Grasshoff, 1959
Lugetti & Tongiorgi, 1965
Alicata, 1966
Tongiorgi, 1966
Brignoli, 1967a
Brignoli, 1967b
Altara, 1968
9
1
3
3
2
3
5
2
16
Brignoli, 1969b
1
Brignoli, 1969c
Dresco & Hubert, 1969
Helsdingen, 1969
1
2
2
2
2
5
6
1
1
38
Puddu, 1970
6
37
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
57
58
59
60
61
62
63
64
65
66
67
68
69
70
72
73
3
2
Lugetti & Tongiorgi, 1969
71
1
Cerruti, 1968
Brignoli, 1969a
6
36
74
2
1
Argiolas et al., 1970
Brignoli, 1971a
Brignoli, 1971b
6
1
5
1
4
Brignoli, 1971c
11
Bartolo et al., 1972
1
Puddu,1971
Casale, 1972
Sanna et al., 1972
3
2
1
3
2
3
Thermes, 1972
16
3
Brignoli, 1973
1
1
Bartolo et al., 1973
Wunderlich, 1973
Baum, 1974
Brignoli, 1974
2
1
1
6
Puddu & Pirodda, 1974
11
Brignoli, 1975
4
Tongiorgi, 1974
Brignoli, 1976a
Brignoli, 1976b
Brignoli, 1977
Brignoli & Gaddini, 1979
Brignoli, 1979a
Brignoli, 1979b
1
2
1
1
1
2
1
Grafitti, 1979
1
Puddu,1980
Brignoli, 1982
Cannas, 1972 [1982]
Grasshoff, 1983
Loerbroks, 1983
Puddu,1983
Brignoli & Murphy, 1984
Brignoli, 1984
1
1
14
Brignoli, 1980
1
7
Brignoli, 1979c
Brignoli, 1979d
1
1
1
15
6
2
2
3
1
2
1
1
1
1
2
1
continued
Table 1. List of publications where data on Sardinian araneofauna is present. For each article, the number of species
mentioned and the new reports for Sardinia are indicated.
7
Citations
75
Grafitti, 1984
77
Grimm, 1986
76
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
98
99
100
101
102
103
104
105
106
107
108
109
110
111
112
Grimm, 1985
Hansen, 1986
Deeleman-Reinhold &
Deeleman, 1988
Tongiorgi & Sola, 1991
Platnick & Di Franco, 1992
Grafitti, 1993
Pons, 1993
Pesarini, 1994
Buchar & Thaler, 1995
Wunderlich, 1995a
Species New
cited species
(n.)
(n.)
1
4
1
2
1
2
1
1
1
1
12
1
1
1
225
1
5
1
Wunderlich, 1995b
13
10
Wunderlich, 1995d
2
1
2
1
Wunderlich, 1995c
Wunderlich, 1995e
Fontana et al., 1996
Knoflach, 1996
Bosmans, 1997
1
1
1
4
1
1
Arnò et al., 1998
32
10
Bosmans & Van Keer, 1999
7
4
Thaler & Knoflach, 1998
Gasparo, 1999
Knoflach, 1999
Metzner, 1999
Knoflach & Thaler, 2000
Pesarini, 2000
Grafitti, 2001
Muccedda et al.,2002
Grafitti, 2003
Casale et al., 2004
Knoflach & Pfaller, 2004
Knoflach, 2004
Logunov, 2004
Muster & Thaler, 2004
Senglet, 2004
Thaler & Knoflach, 2004
1
1
1
3
1
1
1
1
1
2
2
3
1
3
4
3
115
Hansen, 2005
114
116
117
118
119
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
1
3
2
1
Grill et al., 2005
21
3
Helsdingen, 2005
24
10
Buchar et al.,2006
Burger et al., 2006
Lana et al., 2006
Logunov & Azarkina, 2006
Grafitti, 2007a
Grafitti, 2007b
Muster et al., 2007
Bolzern et al., 2008
Bosmans, 2008
Grafitti & Merella, 2008
19
3
1
3
1
2
1
4
2
3
5
1
4
1
1
Loru et al., 2008
71
19
Colombo & Manunza, 2009
1
1
Arnedo et al., 2009
Knoflach et al., 2009
Muster, 2009
2
4
2
2
Pantini & Sassu, 2009
64
8
Marras et al., 2011
1
1
Trotta, 2009
Tanasevitch, 2011
Wunderlich, 2011
Senglet, 2011
30
4
1
3
3
Kulczycki et al., 2012
1
Total
Collection of Bergamo
Museum (MBG)
Collection of Cagliari
University (UCA)
Total
1
1
76
Trotta, 2011
4
2
Di Pompeo et al., 2011
Collection of CNR-ISE
Sassari (ISE)
2
1
Thaler et al., 2004
120
1
1
113
7
1
1417
453
144
21
216
21
9
369
42
Table 1 (continued). List of publications where data on Sardinian araneofauna is present. For each article, the number
of species mentioned and the new reports for Sardinia are indicated.
8
Tongiorgi & Sola (1991) report the presence of
Pardosa gefsana in Sardinia, Wunderlich (1995a,
1995b) describes a new species of Thomisidae and
carries out a bio-geographical analysis on SardoCorsican araneofauna. Arnò et al. (1998) analyze
the spiders on lime trees of a small tree-lined town
in Sassari. Grill et al. (2005) publish an ecological
work on butterflies, spiders and plants in different
environments, van Helsdingen (2005) reports data
on some collections he made on the island, Bolzern
et al. (2008) report interesting data about Tegenaria
and Malthonica, Loru et al. (2008) list the species
of spider collected in Sardinian hazelnut orchards,
Arnedo et al. (2009) describe two new species of
Dysdera, Colombo & Manunza (2009) report the
first sighting of Cyrtarachne ixoides, Pantini &
Sassu (2009) and Trotta (2009) report data on the
spiders of the islands surrounding Sardinia, Di
Pompeo et al. (2011) report the presence of Argiope
trifasciata in Sardinia and Marras et al. (2011) analyze the influence of the exclusion of ants on the
quantity of four groups of generalist predators, including spiders, in a citrus orchard. Trotta (2011)
supplies some faunal data on Sardinian spiders and
Kulczycki et al. (2012) report the first sighting of
Steatoda nobilis. In figure 2 the growth in aranelogical knowledge in Sardinia from 1868 to today
is reported.
Finally, we must point out that in the checklist
of the species of Italian fauna, each species indicates whether it is present in Sardinia. Pesarini
(1994) reported 229 species on the island, some of
which were very probably based on new material
as they had not been cited previously. A final
source that reports general data on the araneofauna
of Sardinia is “Fauna Europaea” (Helsdingen,
2012) which reports 353 species for Sardinia.
Research on araneofauna in Sardinia over the
last 20 years does not stop with that listed above.
In fact, from the early 1990s a working group from
the Institute of Ecosystem Study (ISE-CNR) in
Sassari has been involved in faunal studies of various natural and anthropized ecosystems (Fresi et
al, 2002). This activity has allowed them to collect
a considerable quantity of spiders. The collected
material, only a small part of which has been
published, is the principal source of unpublished
material presented in this catalogue.
MATERIALS AND METHODS
In order to create the catalogue which covers a
time period from 1868 to 2012, 140 publications
were examined. The starting point is in 1868 when
Canestrini and Pavesi published “Araneidi italiani”,
the first catalogue of Italian spiders. The catalogue
was then updated in 1870 by the same authors. For
the bibliography included between 1870 and 1939
the first volume of the “Bibliographia Araneorum”
by Pierre Bonnet (1945) was consulted, while for
later publications, accurate research was carried out
into their bibliographical sources. The complete list
of publications consulted is in table 1. The works are
listed in chronological order indicating the number
of the species and the new sightings for the region.
A fair number of species are reported in the catalogue exclusively thanks to sightings made over a
century ago and which therefore require further
confirmation. However, this problem is often overcome for the most common species through a series
of successive citations.
To avoid an excessive amount of data, each
time that it was possible the citations of other citations were excluded, reporting only records based
on the direct examination of the specimens except
for the data from the checklist by Pesarini (1994)
and from Garneri’s catalogue (1902). Excluded
from the catalogue are: nomina dubia, species inquirendae, citations where the author expresses
doubt about the identification and the few species
whose nomenclature was not easily identifiable
under current nomenclature. When adequate confirmation was missing, the subspecies were
grouped with the nominal species. The taxa or the
citations excluded from the catalogue are still commented on in the notes relating to their respective
families. Nomenclature follows the “The World
Spider Catalog, Version 12.5” by Norman Platnick
(2012), which was referred to also for synonymy.
An asterisk next to the name indicates that the
species was reported for the first time in Sardinia.
Our research included data from literature and reports from approximately 1900 notifications regarding 254 species deriving from the study of new
material kept in the collections of the ISE-CNR at
its Sassari headquarters, at the Civic Natural
Science Museum “E. Caffi” of Bergamo (MBG)
and the University of Cagliari (UCA).
9
Figure 2. The growth in aranelogical knowledge in Sardinia from 1869 to today.
Most of this new material was collected over 20
years of faunal studies carried out by the ISE-CNR
in Sardinia. First of all they were concerned with the
arthropod fauna of the olive grove ecosystem in
1992-1994 and in vineyards in 1994-1996, both in
North Sardinia. Studies continued on urban tree-lined
avenues of the city of Sassari between 1993 and 1996
(Arnò et al., 1998). Other collections/observations
were carried out in mixed fruit orchards (pomaceous,
drupaceous) of the Province of Sassari in 1997-1998
and in a forest of cork oak in North-East Sardinia in
1999-2000. In 2003-2004 research was carried out
on the araneofauna of the island of Asinara Pantini
& Sassu (2009) and continued in Central Sardinia on
the peaks of the Gennargentu Range (2006-2007)
and in different mountainous fruit orchard ecosystems such as the hazelnut in 2004-2006 (Loru et al.,
2008) and the cherry and walnut in 2007-2008. Finally, recently systematic sampling was carried out
in the disused mining area of Montevecchio-Ingurtosu in South-West Sardinia (2009). The analysis of
this material (over 7000 adult specimens) has allowed us to confirm the presence of a large quantity
of species, to widen the area of their distribution and
to identify new species for Sardinia.
For each species examined, the following information is supplied:
i) Chorotype. Referring to the works of Vigna
Taglianti et al. (1992, 1999) each species is assigned
a chorotype with the purpose of precisely defining
its distribution. For the assigning of chorological
categories, several specialist works were consulted
relating to the genera or to the single species in
question. In addition, the following websites were
used: Fauna Europaea [http://www.faunaeur.org/]
(Helsdingen, 2011), Araneae Spiders of Europe
[http://www.araneae.unibe.ch] (Nentwig et al., 2011),
Biodiversity of Russia [http://www.zin.ru/BioDiv/aranei.asp]. Sometimes limited knowledge about the
distribution of some species made it difficult to attribute a chorotype. In this case, the chorotype is
followed by a question mark. For subspecies, the
chorotype of the nominal species is indicated in
square brackets.
ii) References. In this field the publications in
which the species has been cited for Sardinia are
listed. A reference number has been assigned to
each work in increasing order starting from the oldest ones (Tab. 1). Works containing citations of data
which was previously published have been excluded while specimens seen again personally by
the authors in revision works were considered. Next
to the publication, the original identification of the
species is indicated if different from the present one.
10
iii) Province. In order to have an idea of the distribution of the araneological collections in Sardinia, the province where each item was found is
included. In cases where the bibliography indicates
only the region, a definite province has not been attributed and this data is shown by a hyphen. If the
area indicated is on the border between provinces,
these are indicated and separated by a bar. The current eight Sardinian provinces have been considered: Cagliari, Carbonia-Iglesias, Medio Campidano,
Nuoro, Ogliastra, Olbia-Tempio, Oristano and Sassari (Fig. 3). The list of boroughs (APPENDIX 1)
belonging to the new provinces is available on-line
at the link: http://www.biodiversityjournal.com/pdf/
4%281%29_3-104_appendix.pdf and for each of
these the original province it belonged to is mentioned (Cagliari Oristano, Nuoro and Sassari).
iv) Habitat. When indicated in literature, for each
species the environment where specimens were collected is reported. This information is listed in alphabetical order including in brackets the number
of the publication it was taken from.
v) Examined material. The following is indicated
for the new checked material: province, borough (in
underlined characters) in which the specimens were
sampled, number and sex of each specimen. Where
possible we also indicated: location, altitude (m asl),
habitat, date of sampling and the capturing technique
(pitfall traps etc.), if different from direct collection.
Figure 3. Number of species reported for
each province.
LIST OF SPECIES
Family CTENIzIDAE
Cteniza brevidens (Doleschall, 1871)
CHOROTyPE. Tyrrhenian endemic.
REFERENCES. (10) Costa, 1885 (sub Aepicephalus brevidens); (13) Garneri, 1902 (sub Aepicephalus brevidens); (84) Pesarini, 1994 (sub
Aepycephalus brevidens).
PROVINCES. (84); Oristano (10, 13).
REMARKS. Cited only for Sicily (Ausserer, 1871)
and Sardinia. Species insufficently described, most
probably, should be named Aepicephalus sp. (fam.
Cyrtaucheniidae). Mediterranean Cytaucheniidae
are easily confused with Cteniza (all large stocky
black spiders). Aepychephalus has been incorrectly
synonymized with Cyrtauchenius by Raven 1985.
Aepicephalus is common in SW Italy (Campania,
Basilicata, Calabria) and in both Sicily and Sardinia. Probably there are several different species of
Aepicephalus in Italy and the Sardinian species then
would be a new and as yet undescribed species;
hence Aepicephalus sp. (Decae A. pers. comm.).
Cteniza moggridgei O.P.-Cambridge, 1874
CHOROTyPE. Mediterranean.
REFERENCES. (139) Trotta, 2011.
PROVINCE. Carbonia-Iglesias (139).
Cteniza sauvagesi (Rossi, 1788)
REFERENCES. (3) Pavesi, 1876 (sub C. Sauvagii);
(6) Magretti, 1880 (sub C. sauvagei); (7a) Costa,
1882 (sub Migale fodiens); (8) Costa, 1884 (sub
Mygale fodiens); (13) Garneri, 1902 (sub Nemesia
fodiens).
PROVINCE. (8); Cagliari (6, 7a); Carbonia-Iglesias (3, 6); Oristano/Nuoro (13).
Family NEMESIIDAE
Nemesia fertoni Simon, 1914
CHOROTyPE. Sardo-Corsican endemic.
11
REFERENCES. (15) Simon, 1914; (84) Pesarini,
1994; (132) Pantini & Sassu, 2009.
PROVINCES. (84); Sassari (15, 132).
Nemesia kahmanni Kraus, 1955
CHOROTyPE. Sardinian endemic.
REFERENCES. (20) Krauss, 1955; (84) Pesarini,
1994.
PROVINCES. (84); Sassari (20).
Nemesia maculatipes Ausserer, 1871
CHOROTyPE. W-Mediterranean.
REFERENCES. (13) Garneri, 1902; (15) Simon,
1914; (132) Pantini & Sassu, 2009.
PROVINCES. (15); Carbonia-Iglesias (13); Sassari
(132).
REMARKS. This probably is a good species and
endemic to Sardinia. The idea that it is widely
spread in the western Mediterranean is incorrect
(Decae & Di Franco 2005).
Nemesia meridionalis (Costa, 1835)
REFERENCES. (13) Garneri, 1902; (84) Pesarini,
1994 (sub N. badia).
PROVINCE. (84); Carbonia-Iglesias (13).
REMARKS ON FAMILy NEMESIIDAE. A revision of
the Sardinian Nemesia fauna (and also the Corsican
Nemesia fauna) is badly needed. A check of type material, and the analysis of topotypical material is necessary. Regarding our poor state of knowledge of
the genus Nemesia in the Mediterranean area see
Decae & Di Franco (2005: 133). Nemesia africana
was cited by Garneri (1902) and Pesarini (1994). It
is very unlikely that this species really occurs in Sardinia. The species itself is insufficiently described to
make a diagnosis without checking possible type material (Decae A. pers. comm.). It is not reported in
the catalogue. The citations of N. caementaria (Magretti, 1880; Garneri, 1902 (sub N. coementaria); Pesarini, 1994), N. cellicola (Pesarini, 1994) and N.
manderstjernae (Garneri, 1902; Pesarini, 1994) are
based on misidentifications (Decae A. pers. comm.)
and so they are not reported in the catalogue.
12
Family FILISTATIDAE
Filistata insidiatrix (Forskål, 1775)
REFERENCES. (6) Magretti, 1880 (sub F. testacea); (10) Costa, 1885 (sub F. testacea); (13) Garneri, 1902 (sub F. testacea); (42) Puddu,1971 (sub
Filistrata insidiatrix); (45) Sanna et al., 1972; (68)
Brignoli, 1982; (83) Pons, 1993; (84) Pesarini,
1994; (94) Arnò et al., 1998; (133) Trotta, 2009.
PROVINCE. (13, 84); Cagliari (42); CarboniaIglesias (6, 45, 133); Nuoro (68, 83); Olbia-Tempio
(68); Sassari (10, 94, ined.).
HABITAT. Avenue of lime trees (94); hypogean
habitat (42, 45, 83).
EXAMINED MATERIAL. OLBIA-TEMPIO: Olbia,
Isola Tavolara, 1 juv. 24.II.1966, 1 juv. 25.IV.1966
leg. Gruppo Isole C.N.R. (MBG).
*Pritha nana (Simon, 1868)
HABITAT. Garrigue (ined.); mining landfills covered by pioneer plants (ined.).
EXAMINED MATERIAL. MEDIO CAMPIDANO:
Arbus, Ingurtosu, Naracauli, m 170, mining landfills covered by pioneer vegetation 1 male 519.V.2009, 3 males 19.V-4.VI.2009, 3 males, 1
female 29.VI-14.VII.2009, 3 males 4-16.VI.2009,
2 males 16-29.VI.2009, 1 female 14-28.VII.2009
(ISE); m 200, garrigue, 3 males 19.V-4.VI.2009, 2
males 29.VI-14.VII.2009, 2 males 16-29.VI.2009,
2 males 4-16.VI.2009 (ISE), 2 males 4-16.VI.2009,
1 female 14-28.VII.2009 (MBG), 1 male 1428.VII.2009 (ISE) (all pitfall traps and leg. Sassu
A., Verdinelli M.). Guspini, Montevecchio, Piccalinna, m 235, garrigue, pitfall traps, 1 male 1530.VI.2009 leg. Sassu A., Verdinelli M. (ISE).
Family SICARIDAE
Loxosceles rufescens (Dufour, 1820)
CHOROTyPE. Cosmopolitan.
REFERENCES. (6) Magretti, 1880 (sub L. erythrocephala); (10) Costa, 1885; (13) Garneri, 1902 (sub
L. erytrocephala); (31) Brignoli, 1969a; (32) Bri-
gnoli, 1969b; (34) Dresco & Hubert, 1969; (46)
Thermes, 1972; (57) Brignoli, 1976a; (84) Pesarini,
1994; (126) Grafitti & Merella, 2008; (133) Trotta,
2009; (139) Trotta, 2011.
PROVINCE. (84); Cagliari (10, 13, 31, 32, 46, 57,
139); Carbonia-Iglesias (6, 13, 46, 126, 133, 139);
Medio Campidano (139, ined.); Nuoro (31, 34);
Olbia-Tempio (31, 32, 133); Oristano (ined.); Sassari (6, 13, 57, 133, ined.).
HABITAT. Hypogean habitat (31, 34, 46, 126);
orchard (ined.); garrigue (ined.); inside house
(ined.); mediterranean maquis (ined.); mining landfills covered by pioneer plants (ined.).
Arbus, Ingurtosu, Naracauli, m 120, mediterranean maquis, 1 male and 1 female 19.V4.VI.2009, 1 male 16-29.VI.2009, 1 female 1428.VII.2009 (ISE); m 170, mining landfills
covered by pioneer plants, 1 female 5-19.V.2009,
1 male 19.V-4.VI.2009, 1 male and 1 female 416.VI.2009, 2 males 16-29.VI.2009, 6 males and
1 female 29.VI-14.VII.2009 (ISE), 2 males 416.VI.2009 (MBG), 1 male, 2 females and 1 juv.
14-28.VII.2009, 2 males 28.VII-10.VIII.2009, 2
males and 1 female 6-20.X.2009 (ISE); m 200,
garrigue, 2 males and 3 females 19.V-4.VI.2009
(MBG), 1 male and 1 female 19.V-4.VI.2009, 1
male 4-16.VI.2009, 3 males and 1 female 1428.VII.2009 (ISE), 1 male 28.VII-10.VIII.2009
(MBG), 1 male 28.VII-10.VIII.2009, 1 male
25.VIII-8.IX.2009, 1 male 10-25.VIII.2009 (ISE)
(all pitfall traps and leg. Sassu A., Verdinelli M.).
Guspini, Montevecchio, Piccalinna, m 225, mining landfills covered by pioneer plants, 1 female
5-18.V.2009, 1 male 30.VI-13.VII.2009, 2 males
3-15.VI.2009 (ISE); m 235, garrigue, 1 male
18.V-3.VI.2009 (ISE), 1 female 15-30.VI.2009
(MBG), 1 male 15-30.VI.2009, 1 male 1327.VII.2009 (ISE), 1 female 13-27.VII.2009
(MBG), 1 male 6-20.X.2009 (ISE) (all pitfall
traps and leg. Sassu A., Verdinelli M.).
OLBIA-TEMPIO: Olbia, Isola Molara, 1 female
25.II.1966; near little church, 1 female 3.XI.1965;
Isola Tavolara, 24.II.1966 (MBG) (all leg. Gruppo
Isole C.N.R.).
ORISTANO: Bonarcado, inside house, 1 juv
5.I.2009 leg. Sassu A. (ISE).
SASSARI: Sorso, Pianu Mannu, Nali farm, orchard, pitfall traps, 1 male 24-31.VII.1997 leg. Ist.
Ent. SS (ISE).
Family SCyTODIDAE
Scytodes thoracica (Latreille, 1802)
CHOROTyPE. Holarctic.
REFERENCES. (13) Garneri, 1902; (31) Brignoli,
1969a; (84) Pesarini, 1994; (94) Arnò et al., 1998.
PROVINCE. (84); Cagliari (ined.); Carbonia-Iglesias (13, 31); Olbia-Tempio (31); Oristano (ined.);
Sassari (94, ined.).
HABITAT. Avenue of lime trees (94); inside house
(ined.); pinewood (ined.).
EXAMINED MATERIAL. CAGLIARI: Pula, Pixina
Manna, m 140, pinewood, emergence cages, 1 female 25.V.2009 leg. Ist. Ent. SS (ISE).
ORISTANO: Bonarcado, inside house, 1 female
5.I.2009 leg. Sassu A. (ISE).
OLBIA-TEMPIO: Olbia, Isola Molara, 2 females 25.II.1966 leg. Valle A., Bianchi R. (MBG).
Scytodes velutina Heineken et Lowe, 1832
REFERENCES. (13) Garneri, 1902 (sub S. delicatula); (31) Brignoli, 1969a (sub S. delicatula); (57)
Brignoli, 1976a; (133) Trotta, 2009.
PROVINCE. Cagliari (13); Carbonia-Iglesias (31);
Medio Campidano (ined.); Nuoro (31, 57); OlbiaTempio (133); Sassari (ined.).
HABITAT. Garrigue (ined.); mediterranean maquis
(ined.); mining landfills covered by pioneer plants
(ined.); vineyard (ined.).
Arbus, Ingurtosu, Naracauli, m 120, mediterranean
maquis, 1 female 28.VII-10.VIII.2009, 2 males 1025.VIII.2009 (MBG); m 170, mining landfills covered by pioneer plants, 1 male and 1 juv.
5-19.V.2009, 6 males 19.V-4.VI.2009, 2 males and
1 juv. 16-29.VI.2009, 4 males 4-16.VI.2009, 1 male
29.VI-14.VII.2009, 2 males and 3 females 1428.VII.2009, 2 males and 1 females 28.VII10.VIII.2009, 2 females 8-22.IX.2009, 2 males
6-20.X.2009 (ISE); m 200, garrigue, 4 males 19.V4.VI.2009, 1 male 5-19.V.2009, 1 male 1629.VI.2009, 2 males 29.VI-14.VII.2009, 1 male
14-28.VII.2009 (ISE) (all pitfall traps and leg. Sassu
A., Verdinelli M.). Guspini, Montevecchio, Piccalinna, m 225, mining landfills covered by pioneer
plants, 2 males 15-30.VI.2009, 6 males and 1 female
13
3-15.VI.2009, 1 male 30.VI-13.VII.2009, 1 male and
1 female 13-27.VII.2009, 2 males and 1 female
27.VII-10.VIII.2009, 1 female 10-25.VIII.2009
(ISE); m 235, garrigue, 1 male 18.V-3.VI.2009, 1
male 3-15.VI.2009 (ISE), 1 male and 1 female
30.VI-13.VII.2009 (MBG), 1 female 15-30.VI.2009,
2 males 27.VII-10.VIII.2009 (ISE), 1 female 1327.VII.2009 (MBG), 1 male 10-25.VIII.2009 (ISE);
m 260, mediterranean maquis, 1 male and 1 juv. 518.V.2009, 4 males, 1 female and 1 juv. 18.V3.VI.2009, 3 males and 1 female, 30.VI-13.VII.2009,
2 males and 1 female 15-30.VI.2009, 5 males and 1
female 3-15.VI.2009, 1 male 27.VII-10.VIII.2009, 1
male, 2 females and 1 juv 13-27.VII.2009, 1 male
and 1 female 13-27.VII.2009, 1 male 25.VIII8.IX.2009, 2 females 10-25.VIII.2009 (ISE), 1 male
and 1 female 22.IX-6.X.2009 (MBG), 1 male and 1
female 22.IX-6.X.2009 (ISE) (all pitfall traps and
leg. Sassu A., Verdinelli M.).
SASSARI: Torralba, vineyard, 1 male 1.VII.1995,
leg. Ist. Ent. SS (ISE).
Family LEPTONETIDAE
Leptoneta insularis Roewer, 1953
REFERENCES. (19) Roewer, 1953; (28) Brignoli,
1967b; (30) Cerruti, 1968; (84) Pesarini, 1994.
PROVINCE. (84); Sassari (19, 28, 30).
HABITAT. Hypogean habitat (19, 28, 30).
Leptoneta patrizii Roewer, 1953
REFERENCES. (19) Roewer, 1953; (30) Cerruti,
1968; (41) Brignoli, 1971c; (51) Brignoli, 1974;
(84) Pesarini, 1994.
PROVINCE. (84); Carbonia-Iglesias (19, 30, 41, 51).
HABITAT. Hypogean habitat (19, 30, 41, 51).
Leptoneta serbariuana Roewer, 1953
REFERENCES. (19) Roewer, 1953; (29) Altara,
1968; (30) Cerruti, 1968; (33) Brignoli, 1969c; (38)
Puddu, 1970; (52) Puddu & Pirodda, 1974; (84)
Pesarini, 1994.
14
PROVINCE. (84); Carbonia-Iglesias (19, 30);
Ogliastra (29, 33, 38, 52).
HABITAT. Hypogean habitat (19, 29, 30, 33,
38, 52).
Leptoneta taramellii Roewer, 1956
REFERENCES. (22) Roewer, 1956; (84) Pesarini,
1994.
PROVINCE. (84); Nuoro (22).
HABITAT. Hypogean habitat (22).
Paraleptoneta spinimana (Simon, 1884)
REFERENCES. (19) Roewer, 1953 (sub P. fagei);
(28) Brignoli, 1967b (sub P. pasquinii); (30) Cerruti, 1968 (sub P. fagei); (33) Brignoli, 1969c (sub
P. fagei); (41) Brignoli, 1971c (sub P. spinimana
fagei); (52) Puddu & Pirodda, 1974 (sub Segrea
sardiniensis); (64) Brignoli, 1979d; (67)
Puddu,1980; (84) Pesarini, 1994; (121) Grafitti,
2007a (sub Segrea sardiniensis).
PROVINCE. (84); Cagliari (33, 41); Cagliari/Nuoro
(67); Carbonia-Iglesias (28, 33, 41, 52, 64); Ogliastra (64); Sassari (19, 30, 121).
HABITAT. Hypogean habitat (19, 28, 30, 33, 41,
52, 64, 67, 121).
Family PHOLCIDAE
Holocnemus pluchei (Scopoli, 1763)
REFERENCES. (3) Pavesi, 1876 (sub Pholcus rivulatus); (13) Garneri, 1902 (sub H. rivulatus); (20)
Krauss, 1955 (sub H. pulchii); (40) Brignoli, 1971b;
(54) Brignoli, 1975; (84) Pesarini, 1994.
PROVINCE. (20, 84); Cagliari (13, 54); CarboniaIglesias (3, 13, 40, 54); Nuoro (40); Olbia-Tempio
(54); Sassari (40, 54).
Pholcus opilionoides (Schrank, 1781)
1994.
PROVINCE. (84); Cagliari (13).
Pholcus phalangioides (Fuesslin, 1775)
REFERENCES. (5) Pavesi, 1878; (6) Magretti,
1880; (10) Costa, 1885; (13) Garneri, 1902; (14)
Gozo, 1908; (19) Roewer, 1953; (22) Roewer,
1956; (30) Cerruti, 1968; (34) Dresco & Hubert,
1969; (37) Argiolas et al., 1970; (38) Puddu, 1970;
(40) Brignoli, 1971b; (41) Brignoli, 1971c; (44) Casale, 1972; (46) Thermes, 1972; (47) Bartolo et al.,
1973; (51) Brignoli, 1974; (64) Brignoli, 1979d;
(83) Pons, 1993; (84) Pesarini, 1994; (105) Grafitti,
2003; (121) Grafitti, 2007a; (122) Grafitti, 2007b;
(126) Grafitti & Merella, 2008.
PROVINCE. (84); Cagliari (6, 10, 13, 14, 37, 40,
46, 64); Carbonia-Iglesias (14, 19, 30, 40, 41, 46,
47, 64, 83, 105, 126); Nuoro (22, 34, 41, 44, 51, 83);
Ogliastra (38, 64); Olbia-Tempio (5, 13, 40, ined.);
Oristano (64, 122); Sassari (51, 64, 83, 121, ined.).
37, 38, 40, 41, 44, 46, 47, 51, 64, 83, 105, 121, 126,
ined.); inside house (ined.).
Isola Molara, 3 females 25.II.1966 leg. Valle A.
(MBG); Isola Tavolara, Grotta del Papa, 1 female
26.II.1966 leg. Gruppo Isole C.N.R. (MBG).
SASSARI: Sassari, inside house, 1 female
2.X.2007 leg. Sassu A. (ISE).
Psilochorus simoni (Berland, 1911)
CHOROTyPE. European.
REFERENCES. (64) Brignoli, 1979d.
PROVINCE. Cagliari (64).
Spermophora senoculata (Dugès, 1836)
REFERENCES. (34) Dresco & Hubert, 1969; (37)
Argiolas et al., 1970; (40) Brignoli, 1971b; (45) Sanna
et al., 1972; (64) Brignoli, 1979d; (84) Pesarini, 1994.
PROVINCE. (84); Cagliari (37); Carbonia-Iglesias
(45, 64); Nuoro (34, 40).
Spermophorides baunei Wunderlich, 1995
REFERENCES. (87) Wunderlich, 1995b.
PROVINCE. Nuoro/Ogliastra (87); Ogliastra (87).
HABITAT. Under stones (87).
Spermophorides elevata (Simon, 1873)
REFERENCES. (40) Brignoli, 1971b (sub Spermophora elevata).
Medio Campidano (40); Nuoro (40); Olbia-Tempio
(40, ined.); Sassari (40).
Isola Molarotto, 1 juv. 25.II.1966 (MBG); Isola Tavolara, 1 male 5.XI.1965 (MBG) (all leg. Gruppo
Isole C.N.R.).
Family SEGESTRIDAE
Ariadna brignolii Wunderlich, 2011
REFERENCES. (136) Wunderlich, 2011.
PROVINCE. Medio-Campidano (136).
REMARKS. The citation is based only on female
specimens and requires further confirmation.
Segestria bavarica C.L. Koch, 1843
CHOROTyPE. Palearctic.
REFERENCES. (127) Loru et al., 2008.
PROVINCE. (127); Nuoro (ined.).
HABITAT. Hazelnut-orchard (127, ined.).
EXAMINED MATERIAL. NUORO: Tiana, m 522,
hazelnut-orchard, bark traps, 1 juv 19-27.VII.2005
(ISE), 1 juv 31.VIII-7.IX.2005 (MBG) (all leg.
I.S.E.).
Segestria florentina (Rossi, 1790)
REFERENCES. (8) Costa, 1884; (13) Garneri,
1902; (58) Brignoli, 1976b; (84) Pesarini, 1994;
(94) Arnò et al., 1998; (132) Pantini & Sassu, 2009.
(58); Nuoro (8, 13); Olbia-Tempio (8, 13); Sassari
(8, 94, 132, ined.).
15
HABITAT. Avenue of lime trees (94).
EXAMINED MATERIAL. SASSARI: Sassari,
Giordano, m 40, 1 female 6.III.2009, leg. Molinu
A. (ISE).
Segestria senoculata (Linnaeus, 1758)
REFERENCES. (13) Garneri, 1902; (94) Arnò et
al., 1998; (127) Loru et al., 2008.
PROVINCE. (127); Cagliari (13); Nuoro (ined.);
Sassari (94).
HABITAT. Avenue of lime trees (94); hazelnutorchard (127, ined.).
EXAMINED MATERIAL. NUORO: Belvì, Su Ajone,
m 585, hazelnut-orchard, bark traps, 1 juv. 18.VI.2005 (ISE), 1 juv. 1-8.VI.2005 (MBG), 1 juv.
14-19.VII.2005 (MBG), 1 juv. 19-27.VII.2005
(MBG), 1 juv. 10-17.VIII.2005 (MBG), 1 juv. 1017.VIII.2005 (ISE), 1 male 13-20.X.2004 (MBG)
(all leg. I.S.E.).
REMARKS ON FAMILy SEGESTRIDAE. Wunderlich
(2011) illustrates the complex nomenclature and taxonomic issues to do with the genus Ariadna Audouin,
1826 in the Mediterranean. He claims that A. insidiatrix is a species limitated to North Africa, therfore the citation of this species in Sardinia by Trotta
(2009) refers very probably to another taxon,
maybe A. brignolii. While awaiting further data
about the distribution of this genus, it was thought
better not to include A. insidiatrix in the catalogue.
Family DySDERIDAE
Dysdera ancora Grasshoff, 1959
REFERENCES. (23) Grasshoff, 1959; (84) Pesarini, 1994.
PROVINCE. (84); Cagliari (23); Medio Campidano (23); Oristano (23).
REMARKS. It would be necessary to re-examine
Grasshoff’s material in order to define the real identity of the species. (Gasparo F. pers. comm.).
16
Dysdera crocata C.L. Koch, 1838
REFERENCES. (3) Pavesi, 1876 (sub D. crocota);
(4) Magretti, 1878 (sub D. crocota); (6) Magretti,
1880 (sub D. crocota); (10) Costa, 1885; (13) Garneri, 1902 (sub D. crocota); (20) Krauss, 1955; (23)
Grasshoff, 1959; (79) Deeleman-Reinhold & Deeleman, 1988; (84) Pesarini, 1994.
PROVINCE. (13, 20, 23, 84); Cagliari (6, 13, 23,
79); Carbonia-Iglesias (3, 6, 13, 23); Medio-Campidano (ined.); Nuoro (ined.); Oristano (4, 6, 13,
23); Sassari (6, 10, 13, ined.).
HABITAT. Cherry-orchard (ined.); orchard
(ined.); river (23); mediterranean maquis (ined.);
hazelnut-orchard (ined.); near a cave (23); under Juniperus sp. (ined.).
maquis, pitfall traps, 1 female 29.VI-14.VII.2009
leg. Sassu A., Verdinelli M. (ISE). Guspini, Montevecchio, Piccalinna, m 260, mediterranean maquis, pitfall traps, 1 female 30.VI-13.VII.2009 leg.
Sassu A., Verdinelli M. (ISE).
NUORO : Belvì, Antonitzò, m 683, cherryorchard, 2 males 11-18.IV.2007, 2 males 1826.IV.2007, 4 males 16-23.V.2007, 3 males and 3
females 21-28.VI.2007, 1 male and 1 female 1724.VII.2007, 1 female 24-30.VIII.2007 (ISE); Su
Enazzu, m 550, hazelnut-orchard, 4 males and 2 females 27.VII-8.VIII.2006, 1 male and 1 female 1327.VII.2006 (MBG), 3 males 13-27.VII.2006 (ISE),
1 male 8-24.VIII.2006, 1 male and 2 females 21.IX5.X.2006, 1 female 31.X-21.XI.2006, 1 male 516.X.2006, 1 female 21.XI.2006-8.I.2007, 1 male
21.II-19.III.2007 (MBG), 4 males 19.III-26.IV.2007
(MBG), 1 female 26.IV-30.V.2007 (ISE), 1 male
and 1 juv. 30.V-4.VII.2007 (MBG) (all pitfall traps
and leg. I.S.E.).
SASSARI: Sorso, Pianu Mannu, Nali farm, orchard, pitfall traps, 1 male and 1 female 2431.VII.1997 leg. Ist. Ent. SS (ISE).
Dysdera jana Gasparo et Arnedo, 2009
REFERENCES. (128) Arnedo et al., 2009.
PROVINCE. Nuoro (128).
HABITAT. Under Pinus (128); under Quercus
ilex (128).
Dysdera shardana Opatova et Arnedo, 2009
REFERENCES. (23) Grasshoff, 1959 (sub Dysdera
kollari misidentification) (128) Arnedo et al., 2009.
PROVINCE. Cagliari (128); Nuoro (128, ined.);
Oristano (23, 128); Sassari (23).
HABITAT. Castaneus and Quercus suber forest
(127); dry, open Quercus forest (127); mixed Pinus
and Quercus forest (127); old crops and open forest
Quercus and Rubus (127); open Quercus forest
w/mosses (127); Quercus pubescens forest (127);
under Quercus ilex (127); walnut-orchard (ined.).
EXAMINED MATERIAL. NUORO: Tonara, Su Azzime, m 682, walnut-orchard, pitfall traps, 1 male
11-18.IV.2007, 1 male 16-23.V.2007 (MBG) (all
leg. I.S.E.).
Dysdera ventricosa Grasshoff, 1959
REFERENCES. (23) Grasshoff, 1959; (84) Pesarini, 1994.
REMARKS. A species to re-examine. From Grasshoff’s drawing it is unclear how the “bulb” is
formed, which seems to lack all the apical structures
typical of the genus Dysdera. The holotype could
be a defective specimen (Gasparo F. pers. comm.).
Harpactea alicatai Brignoli, 1979
REFERENCES. (62) Brignoli, 1979b; (84) Pesarini, 1994.
PROVINCE. (84); Sassari (62).
Harpactea gennargentu Wunderlich, 1995
PROVINCE. Nuoro (ined.); Nuoro/Ogliastra (87).
HABITAT. Hazelnut-orchard (ined.); alder grove
(ined.); under Juniperus sp. (ined.); under stones in
a forest (87).
EXAMINED MATERIAL. NUORO: Belvì, Su
Enazzu, m 550, hazelnut-orchard, pitfall traps, 1
male 30.V-4.VII.2007 (ISE), 6 males 30.V4.VII.2007 (MBG) (all leg. I.S.E.). Desulo, Gennargentu, mountain refuge, m 1500, under
Juniperus sp., 1 female 11-25.VII.2006 (ISE),
alder-grove, 1 female 23.VIII-5.IX.2006, 1 female
19.IX-3.X.2006, 3 males and 1 female 1024.V.2007 (ISE) (all pitfall traps and leg. Fiori F.).
Fonni, Gennargentu, mountain refuge, m 1550,
under Juniperus sp., pitfall traps, 1 male 519.VI.2007 leg. Fiori F. (ISE).
Harpactea nuragica Alicata, 1966
REFERENCES. (25) Alicata, 1966; (84) Pesarini,
1994.
PROVINCE. (84); Medio-Campidano (ined.);
Nuoro (25).
HABITAT. Quercus ilex forest, under stones (25);
mediterranean maquis (ined.).
maquis, pitfall traps, 2 males 7-21.IV.2009, 2 males
19.V-4.VI.2009, 1 male 5-19.V.2009 leg. Sassu A.,
Verdinelli M. (ISE). Guspini, Montevecchio, Piccalinna, m 260, mediterranean maquis, pitfall traps,
2 males 21.IV-5.V.2009, 2 males 7-21.IV.2009 leg.
Harpactea sardoa Alicata, 1966
REFERENCES. (25) Alicata, 1966; (84) Pesarini,
1994.
PROVINCE. (84); Nuoro (25, ined.).
HABITAT. Cherry-orchard (ined.); Quercus ilex
forest (25); walnut-orchard (ined.).
EXAMINED MATERIAL. NUORO: Belvì, Antonitzò, m 683, cherry-orchard, 1 male 18-26.IV.2007
(ISE); Canale e' Figu, m 576, walnut-orchard, 1
male 16-23.V.2007 (ISE); Mannaritzò, m 706,
cherry-orchard, 1 male 11-18.IV.2007, 3 males 1826.IV.2007 (ISE) (all pitfall traps and leg. I.S.E.).
Tonara, Su Azzime, m 682, walnut-orchard, pitfall
traps, 1 male 18-26.IV.2007, 1 male 16-23.V.2007
leg. I.S.E. (ISE).
17
Parachtes inaequipes (Simon, 1882)
REFERENCES. (54) Brignoli, 1975; (84) Pesarini,
1994.
PROVINCES. (84); Olbia-Tempio (54); Sassari
(54).
Parachtes limbarae (Kraus, 1955)
REFERENCES. (20) Krauss, 1955 (sub Harpactocrates limbarae); (84) Pesarini, 1994.
PROVINCE. (84); Medio Campidano (ined.);
Nuoro (ined.); Sassari (20).
HABITAT. Cherry-orchard (ined.); mediterranean
maquis (ined.); hazelnut-orchard (ined.); aldergrove (ined.).
Guspini, Montevecchio, Piccalinna, m 260, mediterranean maquis, pitfall traps, 1 male 5-18.V.2009
(MBG), 2 males 5-18.V.2009, 1 male 1327.VII.2009 (ISE) (all leg. Sassu A., Verdinelli M.).
NUORO: Belvì, Antonitzò, m 683, cherry-orchard, 1 male 11-18.I.2008 (ISE); Su Enazzu, m
550, hazelnut-orchard, 1 male 21.II-19.III.2007
(ISE) (all pitfall traps and leg. I.S.E.). Desulo, Gennargentu, mountain refuge, m 1500, alder-grove,
pitfall traps, 1 female and 3 males 10-24.V.2007 leg.
Fiori F. (ISE). Fonni, Gennargentu, mountain refuge, m 1550, alder-grove, pitfall traps, 2 males 519.VI.2007 leg. Fiori F. (ISE).
Sardostalita patrizii (Roewer, 1956)
REFERENCES. (22) Roewer, 1956 (sub Stalita patrizii); (30) Cerruti, 1968 (sub Stalita patrizii); (84)
Pesarini, 1994; (98) Gasparo, 1999; (103) Grafitti,
2001; (106) Casale et al., 2004.
PROVINCE. (84); Nuoro (22, 30, 98); Ogliastra
(98, 103, 106).
HABITAT. Hypogean habitat (22, 30, 98, 103, 106).
REMARKS ON FAMILy DySDERIDAE. Citations of
Dysdera erythrina (Costa, 1885; Garneri, 1902) and
D. nicaeensis (Roewer, 1953; Cerruti, 1968) are
18
considered unreliable (Gasparo F. pers. comm.) and
therefore excluded from the catalogue.
Family OONOPIDAE
Oonops placidus corsicus Dalmas, 1916
CHOROTyPE. W-Mediterranean (?).
REFERENCES. (27) Brignoli, 1967a; (54) Brignoli, 1975; (84) Pesarini, 1994 (sub O. placidus).
PROVINCES. (84); Nuoro (27); Oristano (27, 54);
Sassari (27).
REMARKS. Pesarini (1994) cites only O. placidus
in the checklist without indicating the subspecies.
In Sardinia only the subspecies corsicus is cited.
Orchestina pavesii (Simon, 1873)
REFERENCES. (27) Brignoli, 1967a; (54) Brignoli, 1975; (84) Pesarini, 1994.
PROVINCE. (84); Sassari (27, 54, ined.).
HABITAT. Olive-grove (ined.).
EXAMINED MATERIAL. SASSARI: Sassari, Lu
Buttangaru, olive-grove, chemical knockdown, 1
male 20.X.2008 leg. Ist. Ent. SS (MBG).
Orchestina setosa Dalmas, 1916
REFERENCES. (27) Brignoli, 1967a.
PROVINCE. Sassari (27).
Orchestina simoni Dalmas, 1916
CHOROTyPE. S-European.
Silhouettella loricatula (Roewer, 1942)
CHOROTyPE. Turano-Europeo-Mediterranean.
REFERENCES. (27) Brignoli, 1967a (sub Dysderina loricatula); (84) Pesarini, 1994 (sub Dysderina loricata); (118) Burger et al., 2006; (133)
Trotta, 2009.
PROVINCE: (84); Cagliari (133); CarboniaIglesias (133); Medio Campidano (27, ined.);
Nuoro (27, 118); Olbia-Tempio (133); Sassari (27,
ined.).
maquis, 1 female 16-29.VI.2009, 3 females 1428.VII.2009, 1 female 8-22.IX.2009 (ISE); m 170,
mining landfills covered by pioneer plants, 1 female
5-19.V.2009 (ISE) (all pitfall traps and leg. Sassu A.,
Verdinelli M.). Guspini, Montevecchio, Piccalinna,
m 235, garrigue, 1 female 13-27.VII.2009 (ISE); m
260, mediterranean maquis, 1 female 13-27.VII.2009
(MBG), 1 female 22.IX-6.X.2009 (ISE) (all pitfall
SASSARI: Torralba, vineyard, 1 female
29.IV.1995 leg. Ist. Ent. SS (ISE).
Family PALPIMANIDAE
Palpimanus gibbulus Dufour, 1820
1902; (20) Krauss, 1955; (84) Pesarini, 1994; (133)
Trotta, 2009; (139) Trotta, 2011.
PROVINCE. (84); Cagliari (8, 13, ined.); Carbonia-Iglesias (13, 133); Medio Campidano (139,
ined.); Sassari (20).
HABITAT. Garrigue (ined.); pinewood (ined.);
mining landfills covered by pioneer plants (ined.).
Manna, m 140, pinewood, emergence cages, 1 female 29.IV.2009, 1 male, 2 females, 1 juv
25.V.2009 leg. Ist. Ent. SS (ISE).
MEDIO CAMPIDANO: Arbus, Ingurtosu, Naracauli, m 170, mining landfills covered by pioneer
plants, 1 male 21.IV-5.V.2009, 1 male 4-16.VI.2009
(ISE); m 200, garrigue, 1 male 19.V-4.VI.2009
(MBG), 1 female 25.VIII-8.IX.2009 (ISE) (all pitfall traps and leg. Sassu A., Verdinelli M.). Guspini,
Montevecchio, Piccalinna, m 235, garrigue, pitfall
traps, 1 female 18.V-3.VI.2009 (MBG), 1 male
18.V-3.VI.2009, 1 male 3-15.VI.2009 (ISE) (all leg.
Sassu A., Verdinelli M.).
Family MIMETIDAE
Ero aphana (Walckenaer, 1802)
REFERENCES. (13) Garneri, 1902; (112) Thaler
& Knoflach, 2004; (113) Thaler et al., 2004.
PROVINCE. Cagliari (13); Ogliastra (112, 113);
Oristano (ined.); Sassari (ined.).
HABITAT. Orchard (ined.); olive-grove (ined.);
vineyard (ined.).
EXAMINED MATERIAL. ORISTANO: Cabras,
Sinis, at sea level, 1 male 20.IV.2003 leg. Pantini
P., Magnati F. (MBG).
SASSARI: Sorso, Pianu Mannu, Nali farm,
olive-grove, 1 female 25.VI.1993 (ISE); orchard, 1
female 26.VI.1997 (ISE) (all beating tray and leg. Ist.
Ent. SS). Torralba, vineyard, 1 female 17.VI.1995
Ero tuberculata (De Geer, 1778)
1902; (127) Loru et al., 2008.
PROVINCE. (13, 127); Cagliari (ined.); Medio
Campidano (12); Nuoro (ined.); Sassari (ined.).
HABITAT. Citrus-grove (ined.); marshland (12);
hazelnut-orchard (127, ined.); vineyard (ined.).
EXAMINED MATERIAL. CAGLIARI: Uta, S'Appassiu, Agris farm, citrus-grove, beating tray, 1 female
23.X.2000 leg. CRAS (ISE).
NUORO: Belvì, Su Ajone, m 585, hazelnutorchard, bark traps, 1 female 21-29.IX.2005 leg.
I.S.E. (ISE). Tiana, m 522, hazelnut-orchard, beating tray, 1 female 21.IX.2005 leg. I.S.E. (ISE).
15.X.1994 leg. Ist. Ent. SS (ISE).
Mimetus laevigatus (Keyserling, 1863)
CHOROTyPE. Centralasiatic-Mediterranean.
REFERENCES. (13) Garneri, 1902 (sub M. interfector Simon, 1881 misidentification).
Family ERESIDAE
Eresus kollari Rossi, 1846
CHOROTyPE. Sibero-European.
19
REFERENCES. (7a) Costa, 1882 (sub E. quatuorguttatus); (8) Costa, 1884 (sub E. quatuorguttatus);
(13) Garneri, 1902 (sub Eresus cinnaberinus) (Eresus cinnaberinus).
PROVINCE. Nuoro/Ogliastra (7a, 8, 13); Ogliastra
(ined.).
EXAMINED MATERIAL. OGLIASTRA: Arzana,
near Punta La Marmora, 1 male 1.X.2007 leg. Marrosu M. (ISE).
Eresus ruficapillus C.L. Koch, 1846
REFERENCES. (1) Canestrini & Pavesi, 1868; (2a)
Canestrini & Pavesi, 1870; (8) Costa, 1884; (13)
Garneri, 1902.
PROVINCES. (1, 2a); Nuoro/Ogliastra (8); Ogliastra (13).
Family OECOBIIDAE
Oecobius maculatus Simon, 1870
CHOROTyPE. Turano-Mediterranean.
REFERENCES. (90) Wunderlich, 1995e; (116)
Helsdingen, 2005.
PROVINCE. Medio Campidano (ined.); Nuoro
(90, 116); Ogliastra (116); Sassari (90).
HABITAT. Garrigue (ined.); inside house (116);
under stones (90); mining landfills covered by pioneer plants (ined.).
Arbus, Ingurtosu, Naracauli, m 170, mining landfills covered by pioneer plants, 4 males and 1 female 19.V-4.VI.2009, 4 males 4-16.VI.2009, 1
female 16-29.VI.2009, 3 males and 1 female
28.VII-10.VIII.2009, 4 males and 1 female 1025.VIII.2009 (ISE), 1 female 25.VIII-8.IX.2009
(MBG); m 200, garrigue, 1 female 4-16.VI.2009, 1
male 28.VII-10.VIII.2009 (ISE) (all pitfall traps and
leg. Sassu A., Verdinelli M.). Guspini, Montevecchio, Piccalinna, m 225, mining landfills covered
by pioneer plants, pitfall traps, 3 males 518.V.2009, 12 males and 1 female 18.V-3.VI.2009,
3 males and 2 females 15-30.VI.2009 (ISE), 6
males 3-15.VI.2009 (MBG), 1 male and 1 female
30.VI-13.VII.2009, 10 males, 2 females and 1 juv.
3-15.VI.2009, 6 males, 2 females and 3 juv. 1327.VII.2009, 14 males and 5 females 27.VII-
20
10.VIII.2009, 2 males and 1 female 1025.VIII.2009, 2 females 25.VIII-8.IX.2009 (ISE),
1 male and 4 females 10-25.VIII.2009 (MBG) (all
Oecobius navus Blackwall, 1859
REFERENCES. (37) Argiolas et al., 1970 (sub O. annulipes); (46) Thermes, 1972 (sub O. annulipes); (50)
Baum, 1974 (sub O. annulipes); (52) Puddu & Pirodda, 1974 (sub O. annulipes); (94) Arnò et al., 1998
(sub O. annulipes); (132) Pantini & Sassu, 2009.
PROVINCES. Cagliari (37, 46, 52); Medio Campidano (ined.); Nuoro (50); Sassari (94, 132).
habitat (37, 46, 52); garrigue (ined.); mediterranean maquis (ined.).
Arbus, Ingurtosu, Naracauli, m 120, mediterranean maquis, pitfall traps, 1 male 19.V4.VI.2009 (ISE), 1 male and 1 female, 1629.VI.2009 (MBG), 4 males 29.VI-14.VII.2009,
6 males 4-16.VI.2009, 2 males 16-29.VI.2009
(ISE), 1 male 14-28.VII.2009 (MBG), 1 female
14-28.VII.2009 (ISE) (all leg. Sassu A., Verdinelli
M.). Guspini, Montevecchio, Piccalinna, m 235,
garrigue, pitfall traps, 1 male 18.V-3.VI.2009 leg.
Uroctea durandi (Latreille, 1809)
REFERENCES. (13) Garneri, 1902.
PROVINCE. Nuoro/Ogliastra (13).
Family ULOBORIDAE
Hyptiotes paradoxus (C.L. Koch, 1834)
PROVINCE. (127); Nuoro (ined.); Sassari (ined.).
HABITAT. Hazelnut-orchard (127, ined.); olivegrove (ined.); vineyard (ined.).
hazelnut-orchard, beating tray, 1 male 7.IX.2005
leg. I.S.E. (ISE).
SASSARI: Alghero, Mamuntanas, olive-grove,
chemical knockdown, 1 male 13.X.2009, 1 female
20.X.2009 leg. Ist. Ent. SS (ISE). Torralba, vineyard, beating tray, 1 female 30.IX.1995 leg. Ist.
Ent. SS (ISE).
Polenecia producta (Simon, 1873)
PROVINCES. Olbia-Tempio (61).
Uloborus plumipes Lucas, 1846
REFERENCES. (46) Thermes, 1972; (52) Puddu
& Pirodda, 1974; (83) Pons, 1993.
PROVINCE. Cagliari (46, 52, ined.); Nuoro (83);
EXAMINED MATERIAL. CAGLIARI: Domus de
Maria, Punta su Pisu, m 14, 2 males, 2 females,
XII.2006 leg. Ruzzante (MBG).
SASSARI: Sorso, Platamona, at sea level, 1 female 4.VIII.1994 leg. Ist. Ent. SS (ISE).
Uloborus walckenaerius Latreille, 1806
1902; (127) Loru et al., 2008.
PROVINCE. (127); Nuoro (ined.); Olbia-Tempio
(8, 13); Sassari (ined.).
HABITAT. Hazelnut-orchard (127, ined.); vineyard (ined.).
m 585, hazelnut-orchard, beating tray, 1 female
10.VIII.2005 leg. I.S.E. (ISE).
SASSARI: Sassari, Platamona, at sea level, 1
male, 22.V.2008 leg. Sassu A. (ISE). Torralba, vineyard, beating tray, 1 female 6.VIII.1994 leg. Ist.
Ent. SS (ISE).
Family NESTICIDAE
Nesticus eremita Simon, 1879
REFERENCES. (38) Puddu, 1970; (64) Brignoli,
1979d; (83) Pons, 1993; (104) Muccedda et al.,
2002; (139) Trotta, 2011.
PROVINCE. Cagliari (64); Ogliastra (38); Sassari
(83, 104, 139, ined.).
HABITAT. Hypogean habitat (38, 64, 83, 104,
ined.).
EXAMINED MATERIAL. SASSARI: Cossoine, cave
Sa Ucca e Mammuscone, 1 female 22.VII.1993 leg.
Giachino P.M. (MBG).
Family THERIDIIDAE
Anelosimus pulchellus (Walckenaer, 1802)
REFERENCES. (10) Costa, 1885 (sub Theridion
pulchellum).
PROVINCE. Cagliari (10); Sassari (ined.).
EXAMINED MATERIAL. SASSARI: Sassari, Platamona, at sea level, 1 male and 1 female 22.V.2008
leg. Sassu A. (ISE).
Anelosimus vittatus (C.L. Koch, 1836)
vittatum); (67) Puddu,1980; (116) Helsdingen,
2005; (127) Loru et al., 2008.
PROVINCE. (127); Cagliari/Nuoro (67); Nuoro
(116, ined.); Olbia-Tempio (ined.); Oristano (10);
Sassari (ined.).
HABITAT. Hypogean habitat (67); orchard (ined.);
hazelnut-orchard (127, ined.); olive-grove (ined.);
grassland (ined.); forest of Quercus suber (ined.).
m 585, hazelnut-orchard, beating tray, 1 male
1.VI.2005 leg. I.S.E. (ISE). Bolotana, Badde e'Salighes, m 850, grassland, 2 males 27.V.2006 leg.
Sassu A. (ISE). Tiana, m 522, hazelnut-orchard,
beating tray, 1 female 19.V.2005 leg. I.S.E. (ISE).
OLBIA-TEMPIO: Buddusò, near Nuraghe Loelle, forest of Quercus suber, beating tray, 1 male
and 1 female 24.V.1999 leg. Ist. Ent. SS (ISE).
olive-grove, 1 male and 1 female 21.V.1993 (ISE);
olive-grove, beating tray, 1 female 9.VI.1993, 1 fe-
21
male 18.VI.1993 (ISE); orchard, beating tray, 1
male 27.V.1998 (ISE) (all leg. Ist. Ent. SS).
Argyrodes argyrodes (Walckenaer, 1842)
REFERENCES. (3) Pavesi, 1876 (sub A. gibbosa);
(11) de Carlini, 1885 (sub A. gibbosus); (13) Garneri, 1902; (84) Pesarini, 1994.
PROVINCE. (84); Cagliari (3, 13); Carbonia-Iglesias (3, 11, 13); Sassari (ined.).
EXAMINED MATERIAL. SASSARI: Alghero, Mamuntanas, olive-grove, chemical knockdown, 1
male 7.IX.2009 leg. Ist. Ent. SS (ISE).
Asagena phalerata (Panzer, 1801)
REFERENCES. (13) Garneri, 1902; (92) Knoflach,
1996 (sub Steatoda phalerata).
PROVINCE. (13); Nuoro/Ogliastra (13); Ogliastra
(92).
Crustulina scabripes Simon, 1881
PROVINCE. Olbia-Tempio (133).
Cryptachaea riparia (Blackwall, 1834)
REFERENCES. (10) Costa, 1885 (sub Theridion riparium).
*Dipoena coracina (C.L. Koch, 1837)
HABITAT. Vineyard (ined.).
EXAMINED MATERIAL. SASSARI: Torralba, vineyard, 1 female 28.X.1995 leg. Ist. Ent. SS (ISE).
Dipoena melanogaster (C.L. Koch, 1837)
22
hazelnut-orchard, beating tray, 1 male and 1 female
19.V.2005 leg. I.S.E. (ISE).
Dipoena torva (Thorell, 1875)
REFERENCES. (94) Arnò et al., 1998.
Enoplognatha afrodite Hippa et Oksala, 1983
REFERENCES. (96) Bosmans & Van Keer, 1999;
(102) Pesarini, 2000; (127) Loru et al., 2008.
PROVINCE. (127); Carbonia-Iglesias (102);
Nuoro (ined.); Olbia-Tempio (96).
HABITAT. Hazelnut-orchard (127, ined.); park
area (96); Quercus suber forest (96).
16.VI.2005 leg. I.S.E. (MBG).
Enoplognatha gemina Bosmans et Van Keer, 1999
REFERENCES. (96) Bosmans & Van Keer, 1999.
PROVINCE. Ogliastra (96); Olbia-Tempio (96);
Oristano (96); Nuoro (ined.).
HABITAT. Stones (96); stones in grassland (96);
under stones (96).
EXAMINED MATERIAL. NUORO: Gavoi, S'eremu
e sa mela, m 1000, 1 male 30.XII.2006 leg. Mulas
A. (MBG).
Enoplognatha latimana Hippa et Oksala, 1982
REFERENCES. (108) Knoflach, 2004.
PROVINCES. Ogliastra (108).
Enoplognatha mandibularis (Lucas, 1846)
1902; (88) Wunderlich, 1995c (sub E. nigrocincta); (96) Bosmans & Van Keer, 1999; (132) Pantini & Sassu, 2009; (133) Trotta, 2009; (139)
Trotta, 2011.
PROVINCES. Cagliari (13); Carbonia-Iglesias
(139); Nuoro (96); Ogliastra (88, 96); Olbia-Tempio
(133); Oristano (10, 13); Sassari (132).
HABITAT. In a stream (88); under stones (96).
Enoplognatha nigromarginata (Lucas, 1846)
REFERENCES. (13) Garneri, 1902; (96) Bosmans
& Van Keer, 1999.
PROVINCE. Cagliari (13); Ogliastra (96).
HABITAT. Stones along rivulet (96).
Enoplognatha ovata (Clerck, 1757)
REFERENCES. (8) Costa, 1884 (sub Theridion lineatum); (13) Garneri, 1902 (sub Theridion lineatum); (84) Pesarini, 1994; (127) Loru et al., 2008.
PROVINCES. - (84, 127); Cagliari (13); Nuoro (8,
13, ined.); Ogliastra (13).
HABITAT. Hazelnut-orchard (127, ined.); walnutorchard (ined.).
m 585, hazelnut-orchard, beating tray, 1 male and 3
juv. 16.VI.2005, 6 males, 4 females and 3 juv.
29.VI.2005 (MBG), 2 males and 1 female
13.VII.2005 (ISE), 2 males, 1 female and 1 juv.
13.VII.2005, 2 males and 8 juv. 16.VI.2005, 2 males,
3 females and 2 juv. 29.VI.2005 (MBG), 1 male and
2 females 29.VI.2005 (ISE), 1 female 7.IX.2005, 1
female 21.IX.2005 (MBG) (all leg. I.S.E.). Tonara,
Su Azzime, m 682, walnut-orchard, pitfall traps, 1
female 17-24.VIII.2007 leg. I.S.E. (ISE).
Enoplognatha testacea Simon, 1884
REFERENCES. (66) Brignoli, 1980 (sub Robertus
arganoi n.sp.); (84) Pesarini, 1994 (sub Robertus
arganoi); (96) Bosmans & Van Keer, 1999.
PROVINCE. (84); Ogliastra (66); Olbia-Tempio
(96); Oristano (96).
HABITAT. Hypogean habitat (66); stones along
cliff (96).
Enoplognatha thoracica (Hahn, 1833)
REFERENCES. (96) Bosmans & Van Keer, 1999.
PROVINCE. Nuoro (96, ined.); Ogliastra (96);
Olbia-Tempio (96).
HABITAT. Along a river (96); cherry-orchard
(ined.); in Quercus suber forest (96); walnutorchard (ined.); under stones (96).
EXAMINED MATERIAL. NUORO: Belvì, Antonitzò,
m 683, cherry-orchard, 1 male 21-28.VI.2007 (ISE);
Canale e' Figu, m 576, walnut-orchard, 1 female 2430.VIII.2007 (ISE) (all pitfall traps and leg. I.S.E.).
traps, 1 female 17-24.VII.2007 leg. I.S.E. (ISE).
Enoplognatha verae Bosmans et Van Keer, 1999
REFERENCES. (96) Bosmans & Van Keer, 1999;
(133) Trotta, 2009.
PROVINCE. Carbonia-Iglesias (133); Medio
Campidano (96); Oristano (96).
HABITAT. Under stone near cost (96).
Episinus algiricus Lucas, 1846
REFERENCES. (101) Knoflach & Thaler, 2000;
(130) Knoflach et al., 2009.
PROVINCE. Carbonia-Iglesias (101); Ogliastra
(130); Sassari (ined.).
HABITAT. From Quercus (130); inside house
(ined.); in oak forest with rocky, bare ground (130);
pasture plateau, beating from Cistus (130).
EXAMINED MATERIAL. SASSARI: Sassari, Li
Punti, inside house, 1 male 31.V.2010 (MBG).
Episinus maculipes Cavanna, 1876
REFERENCES. (108) Knoflach, 2004; (130) Knoflach et al., 2009; (132) Pantini & Sassu, 2009.
PROVINCES. Ogliastra (108, 130); Sassari (132).
23
HABITAT. From scrub and oak (130).
Episinus theridioides Simon, 1873
REFERENCES. (15) Simon, 1914 (sub Plocamis
theridioides).
Episinus truncatus Latreille, 1809
1902; (84) Pesarini, 1994.
PROVINCE. (84); Cagliari (13); Nuoro (8, 13);
Ogliastra (13); Oristano (8, 13).
Euryopis episinoides (Walckenaer, 1847)
REFERENCES. (2c) Canestrini, 1876 (sub Theridium acuminatum); (10) Costa, 1885 (sub E. acuminata); (13) Garneri, 1902 (sub E. acuminata);
(84) Pesarini, 1994 (sub E. acuminata); (94) Arnò
et al., 1998 (sub E. tarsalis); (132) Pantini & Sassu,
2009; (133) Trotta, 2009.
PROVINCE. (2c, 84); Cagliari (ined.); Nuoro (10,
ined.); Olbia-Tempio (133); Oristano (13); Sassari
(10, 94, 132, ined.).
HABITAT. citrus-grove (ined.); avenue of lime trees
(94); cherry-orchard (ined.); orchard (ined.); walnutorchard (ined.); olive-grove (ined.); pinewood (ined.);
vineyard (ined.).
Manna, m 140, pinewood, emergence cages, 1 female, 21.IV.2009, leg. Ist. Ent. SS (ISE). Uta, S’Appassiu, Agris farm, citrus-grove, beating tray, 1
female 23.XI.2000 leg. CRAS (ISE).
NUORO: Belvì, Antonitzò, m 683, cherry-orchard, bark traps, 1 female 24-30.VIII.2007 (ISE);
Canale e' Figu, m 576, walnut-orchard, bark traps, 1
female 27.IX-4.X.2007 (ISE); Mannaritzò, m 706,
cherry-orchard, pitfall traps, 1 female 16-23.V.2007,
1 female 17-24.VII.2007 (ISE); bark traps, 2 females
24-30.VIII.2007 (ISE) (all leg. I.S.E.).
chemical knockdown,1 female 21.VIII.2009, 3
males and 3 females 6.VIII.2009, 1 male
24
30.IX.2009, 1 female, 13.X.2009 leg. Ist. Ent. SS
(ISE). Sassari, Gioscari, olive-grove, chemical
knockdown, 9 males and 2 females 6.VIII.2008, 1
female 20.VIII.2008, 1 male 4.IX.2008 (ISE); Lu
Buttangaru, olive-grove, 1 male 30.IX.2008, 3 males
13.X.2008, 1 male 29.IX.2009 leg. Ist. Ent. SS
(ISE); Li Punti, Baldinca, m 110, 1 male
14.VII.2008 leg. Pantaleoni R.A. (ISE). Sorso,
Pianu Mannu, Nali farm, orchard, pitfall traps, 1 female 26.VI-2.VII.1997; orchard, bark traps, 1 male
24-31.VII.1997 leg. Ist. Ent. SS (ISE). Torralba,
vineyard, 1 female 23.IV.1994, 1 male 21.V.1994,
1 male 25.VI.1994, 1 male 15.VII.1995, 1 female
19.VIII.1995, 1 female 12.VIII.1995 leg. Ist. Ent.
SS (ISE).
Kochiura aulica (C.L. Koch, 1838)
aulicum); (11) de Carlini, 1885 (sub Theridion aulicum); (13) Garneri, 1902 (sub Theridion aulicum);
(84) Pesarini, 1994 (sub Anelosimus aulicus); (94)
Arnò et al., 1998 (sub Anelosimus aulicus); (114)
Grill et al., 2005 (sub Anelosimus aulicus); (133)
Trotta, 2009; (134) Marras et al., 2011.
PROVINCES. (84, 114, 134); Cagliari (13, ined.);
Carbonia-Iglesias (11, 13); Nuoro/Ogliastra (13);
Ogliastra (13); Olbia-Tempio (10, 133); Oristano
(13); Sassari (94, ined.).
HABITAT. Citrus-grove (ined.); avenue of lime
trees (94); orchard (134); olive-grove (ined.); shurblands and trees, agricultural land (114); vineyard
(ined.).
EXAMINED MATERIAL. CAGLIARI: Uta, S’Appassiu, Agris farm, citrus-grove, beating tray, 1
male 19.VI.2000 leg. CRAS (ISE).
OLBIA-TEMPIO: Olbia, Isola Tavolara - penisolotto, 23.IV.1966 leg. Gruppo Isole C.N.R.
(MBG).
chemical knockdown, 1 female 7.IX.2009, 1 male
27.X.2009, 1 male and 1 female 7.X.2009, 2 males
6.X.2009 (ISE). Sassari, Gioscari, olive-grove, 1 female 21.VIII.2009, Lu Buttangaru, olive-grove,
chemical knockdown, 1 male 20.X.2008 leg. Ist.
Ent. SS. Sorso, Pianu Mannu, Nali farm, olivegrove, 1 female 21.IX.1992, leg. Ist. Ent. SS (ISE).
Torralba, vineyard, 1 male 16.VII.1994 leg. Ist. Ent.
SS (ISE).
Latrodectus tredecimguttatus (Rossi, 1790)
REFERENCES. (1) Canestrini & Pavesi, 1868 (sub
Latrodectes malmignathus); (2a) Canestrini & Pavesi, 1870 (sub Latrodectes malmignathus); (6) Magretti, 1880 (sub L. 13 guttatus); (8) Costa, 1884 (L.
tredecimguttatus, Ross var. lugubris, L.); (13) Garneri, 1902; (84) Pesarini, 1994; (133) Trotta, 2009.
PROVINCE. (1, 2a, 84); Cagliari (8, 13); Carbonia-Iglesias (13); Medio Campidano (6); OlbiaTempio (8, 13); Sassari (8, 13, 133, ined.).
EXAMINED MATERIAL. SASSARI: Ozieri, Chilivani, 1 female 18.XI.1981 leg. Meloni (MBG).
Neottiura bimaculata (Linnaeus, 1767)
bimaculatum Var. pellucidum); (84) Pesarini, 1994.
PROVINCE. (84); Olbia-Tempio (10).
Neottiura herbigrada (Simon, 1873)
REFERENCES. (116) Helsdingen, 2005.
PROVINCE. Ogliastra (116).
HABITAT. Wasteland plot (116).
Neottiura uncinata (Lucas, 1846)
uncinatum); (99) Knoflach, 1999.
PROVINCE. Cagliari (12); Medio Campidano
(12); Oristano (99); Sassari (ined.).
HABITAT. Wetland (12); in grassland near lagoon
(99); pinewood (ined.); vineyard (ined.).
E XAMINED MATERIAL . SASSARI : Alghero,
Pinewood Maria Pia, pinewood, 1 male 4.V.2008
leg. Cesaroni C. (ISE). Sassari, Li Punti, Baldinca,
m 110, 1 male 23.V.2008 (MBG). Torralba, vineyard, 1 female 18.VI.1994 leg. Ist. Ent. SS (ISE).
Paidiscura pallens (Blackwall, 1834)
PROVINCE. (127); Nuoro (ined.); Sassari
(ined.).
HABITAT. Hazelnut-orchard (127, ined.); olivegrove (ined.).
m 585, hazelnut-orchard, beating tray, 8 females
19.V.2005, 4 males and 14 females 4.V.2005, 1 female 29.VI.2005 (MBG), 5 females 1.VI.2005
(ISE), 2 females 1.VI.2005, 2 females 16.VI.2005,
3 males and 9 females 4.V.2005, 1 female
16.VI.2005, 2 females 1.VI.2005, 1 female
27.VII.2005 (MBG) (all leg. I.S.E.). Tiana, m 522,
hazelnut-orchard, beating tray, 1 female 4.V.2005
(MBG), 1 male and 1 female 19.V.2005 (ISE), 1
male and 1 female 19.V.2005, 1 male 16.VI.2005
(MBG) (all leg. I.S.E.).
SASSARI: Sorso, Pianu Mannu, Nali farm, olivegrove, 1 male and 2 females 6.V.1993, 1 male and 1
female 18.III.1994, 2 males and 1 juv. 4.III.1994;
olive-grove, beating tray, 2 males 17.III.1993, 12
males, 3 females and 3 juv. 29.III.1993, 14 males, 14
females and 6 juv. 6.IV.1993, 1 female 9.VII.1993
(ISE) (all leg. Ist. Ent. SS).
Parasteatoda lunata (Clerck, 1757)
REFERENCES. (107) Knoflach & Pfaller, 2004
(sub Achaearanea lunata); (127) Loru et al., 2008
(sub Achaearanea lunata).
PROVINCES. (127); Nuoro (107, ined.).
m 585, hazelnut-orchard, beating tray, 2 females,
16.VI.2005 leg. I.S.E. (ISE).
Parasteatoda tepidariorum (C.L. Koch, 1841)
REFERENCES. (19) Roewer, 1953 (sub Theridion
tepidariorum); (30) Cerruti, 1968 (sub Theridion tepidariorum).
PROVINCES. Sassari (19, 30).
HABITAT. Hypogean habitat (19, 30).
Pholcomma gibbum (Westring, 1851)
25
1974.
PROVINCE. Cagliari (13); Carbonia-Iglesias (51).
Phylloneta impressa (L. Koch, 1881)
REFERENCES.(108) Knoflach, 2004 (sub Theridion impressum); (127) Loru et al., 2008 (sub Theridion impressum); (139) Trotta, 2011.
PROVINCE. (127); Carbonia-Iglesias (139); Nuoro
(ined.); Ogliastra (108); Olbia-Tempio (ined.).
HABITAT. Hazelnut-orchard (127, ined.); forest
of Quercus suber (ined.).
Enazzu, m 550, hazelnut-orchard, beating tray, 1 female 29.VI.2005 leg. I.S.E. (ISE).
28.VI.1999 leg. Ist. Ent. SS (ISE).
Phylloneta sisyphia (Clerck, 1757)
sisyphium); (13) Garneri, 1902 (sub Theridion sisyphium).
PROVINCES. Cagliari (13); Nuoro (8, 13); Oristano (13).
Platnickia tincta (Walckenaer, 1802)
tincutm); (13) Garneri, 1902 (sub Theridion tinctum); (127) Loru et al., 2008 (sub Keijia tincta);
(132) Pantini & Sassu, 2009 (sub Keijia tincta);
(139) Trotta, 2011.
PROVINCE. (127); Cagliari (13); Carbonia-Iglesias (139); Nuoro (ined.); Olbia-Tempio (ined.);
Oristano (10); Sassari (132, ined.).
HABITAT. Hazelnut-orchard (127, ined.); olivegrove (ined.); forest of Quercus suber (ined.).
Enazzu, m 550, hazelnut-orchard, beating tray, 1
male 16.VI.2005, 1 female 29.VI.2005 leg. I.S.E.
26
(MBG). Tiana, m 522, hazelnut-orchard, beating tray,
1 male and 2 females, 16.VI.2005 (MBG), 1 male
1.VI.2005, 1 female 10.VIII.2005 leg. I.S.E. (ISE).
OLBIA-TEMPIO: Buddusò, near Nuraghe Loelle,
forest of Quercus suber, beating tray, 1 female
24.V.1999, 1 male 28.VI.1999, leg. Ist. Ent. SS (ISE).
chemical knockdown, 1 male 6.VIII.2009 leg. Ist.
Ent. SS (ISE). Sassari, Platamona, at sea level, 1
male and 1 female, 22.V.2008 leg. Sassu A. (ISE).
Sorso, Pianu Mannu, Nali farm, olive-grove, 1 male
and 1 female 6.V.1993, 1 male 9.VII.1993 (ISE);
olive-grove, beating tray, 1 male, 1 female and 2
juv. 15.III.1993, 2 females and 1 juv. 29.IV.1993, 2
males, 3 females and 11 juv. 25.VI.1993, 5 males
and 4 females 18.VI.1993, 2 females and 2 juv.
22.VII.1993, 3 females and 18 juv. 1.VII.1993, 2
males, 5 females and 3 juv. 9.VII.1993, 2 males
30.VII.1993, 1 male 12.VIII.1993, 1 female
26.VIII.1993 (ISE) (all leg. Ist. Ent. SS).
*Rhomphaea nasica (Simon, 1873)
HABITAT. Citrus-grove (ined.); olive-grove (ined.)
EXAMINED MATERIAL. CAGLIARI: Uta, S’Appassiu, Agris farm, citrus-grove, beating tray, 1 male
31.VII.2000, 1 juv. 21.IX.2000 leg. CRAS (ISE).
olive-grove, beating tray, 1 male 15.VII.1993 leg.
Ist. Ent. SS (ISE).
Robertus arundineti (O.P.-Cambridge, 1871)
REFERENCES. (66) Brignoli, 1980; (84) Pesarini, 1994.
PROVINCE. (84); Ogliastra (66).
Ruborridion musivum (Simon, 1873)
musivum); (107) Knoflach & Pfaller, 2004 (sub
Theridion musivum); (127) Loru et al., 2008 (sub
Theridion musivum); (130) Knoflach et al., 2009
(sub Theridion musivum).
PROVINCE. (127); Cagliari (130); Carbonia-Igle-
sias (130); Nuoro (ined.); Ogliastra (107, 130); Oristano (130); Sassari (10).
HABITAT. Hazelnut-orchard (127, ined.); pasture
plateau, beating from Cistus sp. (130).
16.VI.2005 leg. I.S.E. (ISE).
Sardinion blackwalli (O.P.-Cambridge, 1871)
REFERENCES. (86) Wunderlich, 1995a (sub S.
perplexum); (87) Wunderlich, 1995b (sub Sardinidion perplexum).
PROVINCE. (87); Ogliastra (86); Sassari (ined.).
HABITAT. On a tree (86); orchard (ined.).
EXAMINED MATERIAL. SASSARI: Sorso, Pianu
Mannu, Nali farm, orchard, beating tray, 1 female
31.VII.1997, 1 male 2.VII.1997 leg. Ist. Ent. SS
(ISE).
Simitidion simile (C.L. Koch, 1836)
REFERENCES. (13) Garneri, 1902 (sub Therodion
simile); (114) Grill et al., 2005; (116) Helsdingen,
2005; (127) Loru et al., 2008.
PROVINCE. (114, 127); Cagliari (ined.); Nuoro
(116, ined.); Ogliastra (13); Olbia-Tempio (ined.);
Sassari (ined.).
HABITAT. Citrus-grove (ined.); low shurbs, shurblands and trees, agricultural land (114); hazelnutorchard (127, ined.); pinewood (ined.); forest of
Quercus suber (ined.).
EXAMINED MATERIAL. CAGLIARI: Uta, S’Appassiu, Agris farm, citrus-grove, beating tray, 1 female 19.VI.2000 leg. CRAS (ISE).
NUORO: Belvì, Su Ajone, m 585, hazelnut-orchard, 1 male 19.V.2005, 1 female 1.VI.2005 (ISE);
Su Enazzu, m 550, hazelnut-orchard, 1 male
1.VI.2005 (MBG) (all beating tray and leg. I.S.E.).
OLBIA-TEMPIO: Buddusò, near Nuraghe Loelle, forest of Quercus suber, beating tray, 1 female
31.V.1999 leg. Ist. Ent. SS (ISE). Tempio, Curadoreddu, m 580, pinewood, emergence cages, 1 female 15.V.2009 leg. Ist. Ent. SS (ISE).
SASSARI: Bonnannaro, Cannisones, 1 male
28.V.2002 leg. Nuvoli T. (ISE).
Steatoda albomaculata (De Geer, 1778)
REFERENCES. (8) Costa, 1884 (sub Lithyphantes
corollatus); (13) Garneri, 1902 (sub Lithyphantes
corollatus); (84) Pesarini, 1994.
PROVINCE. (13, 84); Cagliari (13); Nuoro/Ogliastra
(8, 13).
Steatoda grossa (C.L. Koch, 1838)
REFERENCES. (13) Garneri, 1902 (sub Teutana
grossa); (34) Dresco & Hubert, 1969 (sub Teutana
grossa); (42) Puddu,1971 (sub Teutana grossa);
(51) Brignoli, 1974; (64) Brignoli, 1979d; (83)
Pons, 1993; (133) Trotta, 2009; (139) Trotta, 2011.
PROVINCE. Cagliari (13, 42); Carbonia-Iglesias
(64, 133, 139); Nuoro (34, 83); Olbia-Tempio
(133); Oristano (133); Sassari (51, 83, ined.).
HABITAT. Hypogean habitat (34, 42, 51, 64, 83);
inside house (ined.).
EXAMINED MATERIAL. SASSARI: Alghero, inside
house, 1 female 18.I.2000, 1 male 25.III.2000 leg.
Pantaleoni R.A. (ISE). Sassari, inside house, 1 female 23.I.2007 (ISE).
Steatoda nobilis (Thorell, 1875)
27
lianus); (84) Pesarini, 1994; (132) Pantini & Sassu,
2009; (133) Trotta, 2009; (139) Trotta, 2011.
PROVINCE. (13, 84); Cagliari (6, 13); CarboniaIglesias (6, 13, 139); Medio Campidano (6); Ogliastra (13); Olbia-Tempio (5, 7a, 8, 13); Oristano (6,
13); Oristano/Nuoro (6); Sassari (6, 13, 132, 133).
Steatoda triangulosa (Walckenaer, 1802)
REFERENCES. (10) Costa, 1885 (sub Teutana
triangulosa); (13) Garneri, 1902 (sub Teutana triangulosa); (37) Argiolas et al., 1970 (sub Teutana
triangulosa).
PROVINCE. Cagliari (13, 37); Nuoro/Ogliastra
(13); Oristano (10, 13); Sassari (ined.).
HABITAT. Hypogean habitat (37); in house (ined.).
EXAMINED MATERIAL. SASSARI: Sassari, in
house, Dept. of Plant Protection, 1 female
4.IV.2004 leg. Sassu A. (ISE).
Theridion familiare O.P.-Cambridge, 1871
1994.
Theridion genistae Simon, 1873
REFERENCES. (140) Kulczycki et al., 2012.
PROVINCE. Cagliari (140); Olbia-Tempio (140);
Sassari (140, ined).
HABITAT. Inside house (ined.).
EXAMINED MATERIAL. SASSARI: Alghero, inside
house, 1 male 6.I.2009 leg. Pantaleoni R.A. (ISE).
Sassari, Giordano, m 40, 1 female 21.IV.2004, 1 female 15.II.2009 leg. Molinu A. (ISE).
REFERENCES. (127) Loru et al., 2008; (130) Knoflach et al., 2009.
PROVINCE. (127); Nuoro (ined.); Ogliastra (130).
HABITAT. In oak forest (130); hazelnut-orchard
(127, ined.).
male 1.VI.2005 leg. I.S.E. (ISE).
CHOROTyPE. Centralasiatic-European.
REFERENCES. (5) Pavesi, 1878 (sub Lithyphantes
paykullianus); (6) Magretti, 1880 (sub Lityphantes
paykullianus); (7a) Costa, 1882 (sub Phrurolithus
hamatus); (8) Costa, 1884 (sub Phrurolitus hamatus); (13) Garneri, 1902 (sub Lithyphantes paykul-
HABITAT. garrigue (ined.); mining landfills covered by pioneer plants (ined.); olive-grove (ined.).
Arbus, Ingurtosu, Naracauli, m 170, mining landfills covered by pioneer plants, 1 female 19.V-
Steatoda paykulliana (Walckenaer, 1805)
*Theridion hannoniae Denis, 1944
28
4.VI.2009; m 200, garrigue, 1 male 4-16.VI.2009,
1 male 25.VIII-8.IX.2009 (ISE) (all pitfall traps and
by pioneer plants, 1 male 18.V-3.VI.2009, 1 male
15-30.VI.2009; m 235, garrigue, 1 male 25.VIII8.IX.2009 (ISE) (all pitfall traps and leg. Sassu A.,
Verdinelli M.).
SASSARI: Sassari, Gioscari, olive-grove, chemical
knockdown, 1 male 4.IX.2009 leg. Ist. Ent. SS (ISE).
Theridion malagaense Wunderlich, 2011
Theridion melanurum Hahn, 1831
REFERENCES. (13) Garneri, 1902 (sub T. denticulatum).
PROVINCE. (13); Sassari (ined.).
Mannu, Nali farm, olive-grove, beating tray, 1 female 18.VI.1993 leg. Ist. Ent. SS (ISE).
Theridion mystaceum L. Koch, 1870
CHOROTyPE. Asiatic-European.
REFERENCES. (94) Arnò et al., 1998; (116) Helsdingen, 2005; (127) Loru et al., 2008.
PROVINCE. (127); Cagliari (ined.); Nuoro (ined.);
Ogliastra (116); Olbia-Tempio (ined.); Sassari (94).
trees (94); inside house (116); hazelnut-orchard
(127, ined.); pinewood (ined.); forest of Quercus
suber (ined.).
Manna, m 140, pinewood, emergence cages, 2
males, 16.IV.2009, 1 male 29.IV.2009, 1 female
25.V.2009 leg. Ist. Ent. SS (ISE). Uta, S’Appassiu,
Agris farm, citrus-grove, beating tray, 1 female
19.VI.2000, 1 female 10.VII.2000 leg. CRAS (ISE).
NUORO: Belvì, Su Ajone, m 585, hazelnut-orchard, beating tray, 1 female 19.V.2005 (MBG), 1
female 4.V.2005 (ISE) (all leg. I.S.E.).
OLBIA-TEMPIO: Buddusò, near Nuraghe Lo-
elle, forest of Quercus suber, beating tray, 1 male,
24.V.1999, leg. Ist. Ent. SS (ISE).
Theridion nasutum Wunderlich, 1995
PROVINCE. Nuoro/Ogliastra (87); Ogliastra (87).
HABITAT. Shrubs (87); under stones (87); under
the bark (87).
Theridion pinastri L. Koch, 1872
(ined.); Sassari (ined.).
HABITAT. Hazelnut-orchard (127, ined.); olivegrove (ined.); forest of Quercus suber (ined.).
29.VI.2005 leg. I.S.E. (ISE).
31.V.1999, 1 male and 1 female 21.VI.1999 leg. Ist.
Ent. SS (ISE).
olive-grove, 1 female 9.VI.1993 (ISE), olive-grove,
beating tray, 1 male and 1 female 9.VI.1993, 2
males and 1 female, 18.VI.1993 leg. Ist. Ent. SS
Theridion varians Hahn, 1833
REFERENCES. (12) Costa, 1886 (sub T. rusticum);
(13) Garneri, 1902; (127) Loru et al., 2008.
PROVINCE. (127); Cagliari (13, ined.); Medio
Campidano (12); Nuoro (ined.); Ogliastra (13);
Sassari (ined.).
HABITAT. Citrus-grove (ined.); wetland (12); orchard (ined.); hazelnut-orchard (127, ined.); olivegrove (ined.).
EXAMINED MATERIAL. CAGLIARI: Uta, S’Appassiu, Agris farm, citrus-grove, beating tray, 13 females 19.VI.2000, 11 females 29.VI.2000, 2 females
31.VII.2000, 9 females 10.VII.2000, 1 female
20.VII.2000, 1 female 11.IX.2000 leg. CRAS (ISE).
NUORO: Belvì, Su Ajone, m 585, hazelnut-or-
chard, beating tray, 2 males 16.VI.2005, 2 males
13.VII.2005 (MBG), hazelnut-orchard, bark traps,
1 female 5-10.VIII.2005 (ISE); Su Enazzu, m 550,
hazelnut-orchard, beating tray, 4 females 16.VI.2005,
1 female 29.VI.2005, 1 female 27.VII.2005 (MBG)
(all leg. I.S.E.). Tiana, m 522, hazelnut-orchard, beating tray, 1 male and 1 female 16.VI.2005 (ISE), 1 females 13.VII.2005 (MBG) (all leg. I.S.E.).
olive-grove, 4 males 3.VI.1993, orchard, 1 male
and 2 females 27.V.1998 (ISE) (all beating tray and
leg. Ist. Ent. SS).
Family MySMENIDAE
Mysmenella jobi (Kraus, 1967)
male 19.V.2005 leg. I.S.E. (ISE).
Family LINyPHIIDAE
Acartauchenius sardiniensis Wunderlich, 1995
Araeoncus longiusculus (O.P.-Cambridge, 1875)
REFERENCES. (87) Wunderlich, 1995b; (132)
Pantini & Sassu, 2009.
PROVINCE. Nuoro (ined.); Ogliastra (87); Sassari (132).
HABITAT. Cherry-orchard (ined.).
m 683, cherry-orchard, 1 male 16-23.V.2007 (ISE);
Mannaritzò, m 706, cherry-orchard, 1 male 1826.IV.2007 (ISE) (all pitfall traps and leg. I.S.E.).
29
Bolyphantes nigropictus Simon, 1884
REFERENCES. (9) Simon, 1884; (13) Garneri,
1902; (17) Simon, 1929.
PROVINCE. (17); Sassari (9, 13).
Centromerus bonaeviae Brignoli, 1979
REFERENCES. (64) Brignoli, 1979d; (84) Pesarini, 1994.
PROVINCE. (84); Carbonia-Iglesias (64).
Centromerus puddui Brignoli, 1979
REFERENCES. (64) Brignoli, 1979d; (84) Pesarini, 1994.
PROVINCE. (64, 84).
*Centromerus subalpinus Lessert, 1907
HABITAT. Hazelnut-orchard (ined.).
male 30.V-4.VII.2007 leg. I.S.E. (MBG).
*Centromerus succinus (Simon, 1884)
Enazzu, m 550, hazelnut-orchard, pitfall traps, 1 female, 8-24.VIII.2006, leg. I.S.E. (MBG).
REMARKS. This data represents the first record
for Italian fauna.
Cresmatoneta eleonorae (Costa, 1883)
REFERENCES. (7b) Costa, 1883 (sub Formicina
eleonorae); (8) Costa, 1884 (sub Formicina Eleonorae); (13) Garneri, 1902 (sub Formicina eleonorae); (84) Pesarini, 1994.
30
PROVINCES. (84); Sassari (7b, 8, 13).
REMARKS. Species described from a female in
1883 and never found since. The validity of the
species requires further confirmation.
Cresmatoneta mutinensis (Canestrini, 1868)
PROVINCE. (127); Nuoro (ined.); Sassari (ined.).
male, 4.X.2005 leg. I.S.E. (ISE).
chemical knockdown, 1 male and 4 females,
6.VIII.2009, 1 female 7.X.2009 leg. Ist. Ent. SS (ISE).
Diplocephalus graecus (O.P.-Cambridge, 1872)
REFERENCES. (127) Loru et al., 2008; (132) Pantini & Sassu, 2009.
PROVINCE. (127); Cagliari (ined.); Medio Campidano (ined.); Nuoro (ined.); Sassari (132, ined.).
HABITAT. Citrus-grove (ined.); cherry-orchard
(ined.); garrigue (ined.); hazelnut-orchard (127,
ined.); olive-grove (ined.); under Juniperus sp.
(ined.).
male, 2.XI.2000, 1 female, 13.XI.2000, 1 male
4.XII.2000, leg. CRAS (ISE).
plants, 1 male 21.IV-5.V.2009, 1 female 6-20.X.2009;
m 200, garrigue, 1 male 19.V-4.VI.2009 (ISE) (all pitfall traps and leg. Sassu A., Verdinelli M.). Guspini,
Montevecchio, Piccalinna, m 225, mining landfills
covered by pioneer plants, 1 male 6-20.X.2009 (ISE);
m 235, garrigue, 1 female 6-20.X.2009 (MBG) (all
pitfall traps and leg. Sassu A., Verdinelli M.).
NUORO: Belvì, Antonitzò, m 683, cherry-orchard, pitfall traps, 1 male 11-18.IV.2007 (ISE);
Mannaritzò, m 706, cherry-orchard, pitfall traps, 1
male and 1 female, 16-23.V.2007 (ISE); Su Ajone,
1.VI.2005 (MBG); Su Enazzu, m 550, hazelnut-orchard, pitfall traps, 1 male 26.IV-30.V.2007 (MBG),
1 male and 1 female, 30.V-4.VII.2007 (ISE) (all leg.
I.S.E.). Fonni, Gennargentu, mountain refuge, m
1550, under Juniperus sp., pitfall traps, 1 male
19.VI-1.VII.2007, leg. Fiori F. (ISE).
chemical knockdown, 1 male 30.IX.2009, 2 females
20.X.2009, 1 female 13.X.2009, 1 female 27.X.2009
leg. Ist. Ent. SS (ISE). Sorso, Pianu Mannu, Nali
farm, olive-grove, beating tray, 2 males and 2 females 13.V.1993 leg. Ist. Ent. SS (ISE).
*Diplostyla concolor (Wider, 1834)
Enazzu, m 550, hazelnut-orchard, pitfall traps, 1 female 13-27.VII.2006 (MBG), 1 female 24.VIII5.IX.2006 (ISE) 1 female 24.VIII-5.IX.2006
(MBG), 1 male 19.III-26.IV.2007 leg. I.S.E. (ISE).
Entelecara acuminata (Wider, 1834)
Enazzu, m 550, hazelnut-orchard, beating tray, 1 female, 19.V.2005, 1 female 1.VI.2005 (MBG), 1 female 29.VI.2005 (ISE), 1 female 27.VII.2005
(MBG) (all leg. I.S.E.).
Erigone dentipalpis (Wider, 1834)
REFERENCES. (132) Pantini & Sassu, 2009.
PROVINCE. Medio-Campidano (ined.); Nuoro
(ined.); Sassari (132).
H ABITAT. Cherry-orchard (ined.); hazelnutorchard (ined.); mining landfills covered by pioneer
plants (ined.).
Guspini, Montevecchio, Piccalinna, m 225, mining
landfills covered by pioneer plants, pitfall traps, 1
male 21.IV-5.V.2009, 1 female 18.V-3.VI.2009 leg.
NUORO: Belvì, Mannaritzò, m 706, cherry-orchard, 1 female 11-18.IV.2007 (ISE); Su Enazzu, m
550, hazelnut-orchard, 1 male 13-27.VII.2006
(MBG), 3 males 30.V-4.VII.2007, 3 males 30.V4.VII.2007 (MBG) (all pitfall traps and leg. I.S.E.).
Frontinellina frutetorum (C.L. Koch, 1834)
REFERENCES. (10) Costa, 1885 (sub Linyphia frutetorum); (13) Garneri, 1902 (sub Linyphia frutetorum); (116) Helsdingen, 2005; (139) Trotta, 2011.
(139); Nuoro (116); Ogliastra (13).
HABITAT. Shurbs and lower branches of trees
(116).
Gnathonarium dentatum (Wider, 1834)
REFERENCE. (135) Tanasevitch, 2011.
PROVINCE. Oristano (135).
Gonatium biimpressum Simon, 1884
REFERENCES. (13) Garneri, 1902; (16) Simon,
1926; (84) Pesarini, 1994.
Gonatium nemorivagum (O. P.-Cambridge, 1875)
REFERENCES. (10) Costa, 1885 (sub G. nemorivaga).
*Hybocoptus corrugis (O.P.-Cambridge, 1875)
HABITAT. Forest of Quercus suber (ined.).
EXAMINED MATERIAL. OLBIA-TEMPIO: Buddusò,
near Nuraghe Loelle, forest of Quercus suber, beating tray, 1 male 20.XI.1999 leg. Ist. Ent. SS (ISE).
for Italian fauna.
31
Incestophantes crucifer (Menge, 1866)
REFERENCES. (13) Garneri, 1902 (sub Leptyphantes crucifer).
Lepthyphantes eleonorae Wunderlich, 1995
HABITAT. Under stones along a stream (87).
Lepthyphantes minutus (Blackwall, 1833)
Lepthyphantes paoloi Wunderlich, 1995
PROVINCE. Nuoro/Ogliastra (87); Ogliastra (87);
Olbia-Tempio (87).
HABITAT. Under stones along a stream (87).
Lepthyphantes sardous Gozo, 1908
REFERENCES. (14) Gozo, 1908 (sub L. sardoa
Gozo, 1908); (30) Cerruti, 1968 (sub L. sardoa);
(84) Pesarini, 1994; (119) Lana et al., 2006.
PROVINCE. (84); Ogliastra (14, 30, 119).
Lessertia dentichelis (Simon, 1884)
REFERENCES. (41) Brignoli, 1971c; (52) Puddu
& Pirodda, 1974; (84) Pesarini, 1994.
Linyphia hortensis Sundevall, 1830
32
REFERENCES. (7a) Costa, 1882 (sub L. pratensis); (8) Costa, 1884 (sub L. pratensis).
PROVINCE. Sassari (7a, 8).
Linyphia triangularis (Clerck, 1757)
1902 (sub L. montana) ; (139) Trotta, 2011.
Campidano (139); Nuoro (10, 139); Nuoro/Ogliastra (13); Ogliastra (13, 139); Sassari (ined.).
EXAMINED MATERIAL. SASSARI: Alghero, Mamuntanas, olive-grove, chemical knockdown, 1 male
27.X.2009 leg. Ist. Ent. SS (ISE). Sassari, Gioscari,
olive-grove, 1 male 13.X.2009 leg. Ist. Ent. SS (ISE).
Maso gallicus Simon, 1894
HABITAT. Grassland along the mouth of the
river (116).
Mecopisthes millidgei Wunderlich, 1995
REFERENCES. (87) Wunderlich, 1995b; (94)
Arnò et al., 1998 (sub M. silus)
PROVINCE. Ogliastra (87); Sassari (94).
HABITAT. Under stones (87); avenue of lime
trees (94).
REMARKS. After re-examination of the material
on M. silus from Arnò et al. (1998) it was attributed
to M. millidgei.
*Meioneta mollis (O.P.-Cambridge, 1871)
Enazzu, m 550, hazelnut-orchard, pitfall traps, 1 female 27.VII-8.VIII.2006 (ISE), 1 male, 8.I21.II.2007, 4 males 30.V-4.VII.2007 (MBG), 3
males 30.V-4.VII.2007 (ISE) (all leg. I.S.E.).
Meioneta pseudorurestris Wunderlich, 1980
REFERENCES. (87) Wunderlich, 1995b (sub Agyneta pseudorurestris); (132) Pantini & Sassu, 2009;
(134) Marras et al., 2011.
PROVINCE. (134); Cagliari (ined.); Medio Campidano (ined.); Nuoro (ined.); Ogliastra (87); Sassari (132, ined.).
(ined.); orchard (134); garrigue (ined.); mediterranean maquis (ined.); hazelnut-orchard (ined.);
walnut-orchard (ined.); olive-grove (ined.); under
Juniperus sp. (ined.); under stones (87); mining
landfills covered by pioneer plants (ined.).
male 29.VI.2000, 1 male and 1 female 19.VI.2000,
1 male 10.VII.2000 leg. CRAS (ISE).
MEDIO CAMPIDANO: Arbus, Ingurtosu, Naracauli, m 120, mediterranean maquis, 2 males 1629.VI.2009; m 170, mining landfills covered by
pioneer plants, 1 female 19.V-4.VI.2009; m 200,
garrigue, 2 males and 3 females 19.V-4.VI.2009, 1
male 5-19.V.2009 (ISE), 1 male and 1 female 19.V4.VI.2009 (MBG), 1 female 6-20.X.2009 (ISE) (all
pitfall traps and leg. Sassu A., Verdinelli M.). Guspini, Montevecchio, Piccalinna, m 235, garrigue,
pitfall traps, 2 males 18.V-3.VI.2009, leg. Sassu A.,
Verdinelli M. (ISE).
NUORO: Belvì, Antonitzò, m 683, cherry-orchard, 3 females 18-26.IV.2007, 1 male 1623.V.2007, 1 female 11-18.I.2008 (ISE); Canale e'
Figu, m 576, walnut-orchard, 1 female 26.IV4.V.2007, 1 female 16-23.V.2007, 1 female 2128.VI.2007, 1 female 17-24.VII.2007 (ISE);
Mannaritzò, m 706, cherry-orchard, 1 male 1623.V.2007, 1 male 4-11.X.2007, 2 males and 1 female, 19-26.XI.2007, 1 female 11-18.I.2008 (ISE);
Su Enazzu, m 550, hazelnut-orchard, 1 male 30.V4.VII.2007 (ISE) (all pitfall traps and leg. I.S.E.).
Desulo, Gennargentu, mountain refuge, m 1500,
under Juniperus sp., pitfall traps, 1 male 17.VI1.VII.2007, leg. Fiori F. (ISE). Tonara, Su Azzime,
m 682, walnut-orchard, pitfall traps, 1 male, 1826.IV.2007, 1 male 16-23.V.2007 leg. I.S.E. (ISE).
SASSARI: Sassari, Gioscari, olive-grove, chemical knockdown, 1 female 22.IX.2008, 1 female
*Micrargus subaequalis (Westring, 1851)
HABITAT. Under Juniperus sp. (ined.).
EXAMINED MATERIAL. NUORO: Fonni, Gennargentu, mountain refuge, m 1550, under Juniperus
sp., pitfall traps, 1 male 5-19.VI.2007 leg. Fiori
F. (MBG).
Microctenonyx subitaneus (O.P.-Cambridge, 1875)
CHOROTyPE.Holarctic.
REFERENCES. (46) Thermes, 1972 (sub Aulacocyba subitanea); (52) Puddu & Pirodda, 1974 (sub
Aulacocyba subitanea); (67) Puddu,1980 (sub Aulacocyba subitanea); (84) Pesarini, 1994; (94) Arnò
et al., 1998; (104) Muccedda et al.,2002 (sub Aulacocyba subitanea); (132) Pantini & Sassu, 2009;
(135) Tanasevitch, 2011.
PROVINCE. (84); Cagliari (135); Cagliari/Nuoro
(67); Carbonia-Iglesias (46, 52); Nuoro (ined.); Sassari (94, 104, 132, ined.).
habitat (46, 52, 67, 104); cherry-orchard (ined.);
hazelnut-orchard (ined.); walnut-orchard (ined.);
olive-grove (ined.).
EXAMINED MATERIAL. NUORO: Belvì, Antonitzò, m 683, cherry-orchard, 1 male 1826.IV.2007, 8 male 19-26.XI.2007, 2 males
22-29.II.2008 (ISE); Canale e' Figu, m 576, walnut-orchard, 2 males 11-18.I.2008 (ISE); Mannaritzò, m 706, cherry-orchard, 1 male 11-18.IV.2007,
1 male 18-26.IV.2007, 1 male 16-23.V.2007, 1 male
19-26.XI.2007 (ISE); Su Enazzu, m 550, hazelnut-orchard, 1 male 24.VIII-5.IX.2006, 2 females
30.V-4.VII.2007 (MBG), 1 male, 1 female 30.V4.VII.2007 (ISE) (all pitfall traps and leg. I.S.E.).
traps, 1 male 19-26.XI.2007 leg. I.S.E. (ISE).
chemical knockdown, 1 male 6.VIII.2009 leg. Ist.
Ent. SS (ISE). Sorso, Pianu Mannu, Nali farm,
olive-grove, beating tray, 1 female 29.III.1993 leg.
Ist. Ent. SS (ISE).
Microlinyphia pusilla (Sundevall, 1830)
REFERENCES. (8) Costa, 1884 (sub Linyphia pusilla); (13) Garneri, 1902 (sub Linyphia pusilla).
33
(13); Sassari (13).
*Nematogmus sanguinolentus (Walckenaer, 1841)
EXAMINED MATERIAL. SASSARI: Sassari, Lu
male 13.X.2008 leg. Ist. Ent. SS (MBG).
Neriene clathrata (Sundevall, 1830)
Neriene furtiva (O.P.-Cambridge,1871)
HABITAT. In shurbs along a little stram (116).
Ouedia rufithorax (Simon, 1881)
PROVINCE. Sassari (132, ined.).
EXAMINED MATERIAL. SASSARI: Alghero, Mamuntanas, olive-grove, chemical knockdown, 1 female 20.X.2009, 2 males 27.X.2009, 1 male
Palliduphantes angustiformis (Simon, 1884)
REFERENCES. (9) Simon, 1884; (13) Garneri,
1902; (17) Simon, 1929; (41) Brignoli, 1971c; (52)
Puddu & Pirodda, 1974; (64) Brignoli, 1979d; (84)
Pesarini, 1994 (all sub Leptyphantes angustiformis);
(132) Pantini & Sassu, 2009; (139) Trotta, 2011.
PROVINCE. (17); (84); Carbonia-Iglesias (41, 52,
64, 139); Medio Campidano (ined.); Nuoro (ined.);
Sassari (9, 13, 64, 132, ined.).
HABITAT. Hypogean habitat (41, 52, 64); cherryorchard (ined.); garrigue (ined.); mediterranean
34
maquis (ined.); hazelnut-orchard (ined.); walnutorchard (ined.); alder-grove (ined.); mining landfills covered by pioneer plants (ined.).
maquis, 1 male and 2 females 19.V-4.VI.2009
(MBG), 2 females 29.VI-14.VII.2009, 1 female 416.VI.2009, 1 female 22.IX-6.X.2009, 1 male 620.X.2009; m 170, mining landfills covered by
pioneer plants, 1 female 22.IX-6.X.2009 (ISE) (all
pitfall traps and leg. Sassu A., Verdinelli M.). Guspini,
covered by pioneer plants, 1 female 5-18.V.2009, 1
male 8-22.IX.2009, 1 male 6-20.X.2009, m 235; garrigue, 2 femals 22.IX-6.X.2009; m 260, mediterranean maquis, 1 female 21.IV-5.V.2009, 3 females
18.V-3.VI.2009, 2 females 5-18.V.2009, 1 female 1530.VI.2009, 1 male, 1 female 13-27.VII.2009, 1 female 22.IX-6.X.2009 (ISE) (all pitfall traps and leg.
NUORO: Belvì, Antonitzò, m 683, cherry-orchard, 1 male and 2 females 16-23.V.2007 (ISE);
Canale e' Figu, m 576, walnut-orchard, pitfall traps,
4 females 16-23.V.2007, 1 female 19-26.XI.2007
(ISE); Mannaritzò, m 706, cherry-orchard, 3 females
16-23.V.2007, 2 males and 1 female 21-28.VI.2007,
1 female 4-11.X.2007, 1 male 19-26.XI.2007 (ISE);
Su Enazzu, m 550, hazelnut-orchard, 1 female
27.VII-8.VIII.2006, 1 female 8-24.VIII.2006, 1
male 21.IX-5.X.2006, 1 female 8.I-21.II.2007, 1
male and 3 females 30.V-4.VII.2007 (MBG), 1 male
and 2 females 30.V-4.VII.2007 (ISE) (all pitfall
traps and leg. I.S.E.). Fonni, Gennargentu, mountain
refuge, m 1550, alder-grove, pitfall traps, 1 female
5-19.VI.2007 leg. Fiori F. (ISE). Tonara, Su Azzime,
m 682, walnut-orchard, pitfall traps, 2 males 1118.IV.2007, 1 male, 3 females 21-28.VI.2007, 2
males and 1 female 24-30.VIII.2007, 1 male 1926.XI.2007, 1 female 11.I-18.I.2008, 1 female 2229.II.2008 leg. I.S.E. (ISE).
OLBIA-TEMPIO: Olbia, Isola Molarotto, 1 female 23.III.1966 leg. Gruppo Isole C.N.R. (MBG).
Pelecopsis bucephala (O.P.-Cambridge, 1875)
Pelecopsis elongata (Wider, 1834)
HABITAT. Cherry-orchard (ined.); hazelnut-orchard (127, ined.); walnut-orchard (ined.).
EXAMINED MATERIAL. NUORO: Belvì, Antonitzò, m 683, cherry-orchard, bark traps, 1 female
4.X-26.XI.2007 (ISE); Canale e' Figu, m 576, walnut-orchard, bark traps, 1 female 19-26.XI.2007, 1
female 11-18.I.2008 (ISE); Su Ajone, m 585, hazelnut-orchard, beating tray, 1 female 4.V.2005 (ISE);
Su Enazzu, m 550, hazelnut-orchard, pitfall traps,
1 male 21.IX.2006-8.I.2007 (ISE) (all leg. I.S.E.).
Tonara, Su Azzime, m 682, walnut-orchard, bark
traps, 1 female 18-26.IV.2007, 1 female 1118.IV.2007 leg. I.S.E. (ISE).
Pelecopsis inedita (O.P.-Cambridge, 1875)
REFERENCES. (127) Loru et al., 2008; (132) Pantini & Sassu, 2009.
(ined.); hazelnut-orchard (127, ined.); olive-grove
(ined.); pinewood (ined.).
EXAMINED MATERIAL. CAGLIARI: Uta, S’Appassiu, Agris farm, citrus-grove, beating tray, 1 female 23.XI.2000, 2 males 4.XII.2000 leg. CRAS
(ISE).
NUORO: Belvì, Antonitzò, m 683, cherry-orchard, 1 male 19-26.XI.2007 (ISE); Mannaritzò, m
706, cherry-orchard, 2 males and 3 females, 1826.IV.2007, 4 males, 4 females 16-23.V.2007, 1
male and 2 females 16-23.V.2007, 2 females 2128.VI.2007 (ISE); Su Enazzu, m 550, hazelnut-orchard, 1 male 21.II-19.III.2007 (ISE), 1 male
21.II-19.III.2007 (MBG) (all pitfall traps and leg.
I.S.E.). Tiana, m 522, hazelnut-orchard, beating
tray, 1 female 4.V.2005 leg. I.S.E. (ISE).
SASSARI: Alghero, Porto Conte, m 35, pinewood, emergence cages, 1 female 13.IV.2009, 2 females 15.V.2009, leg. Ist. Ent. SS (ISE);
Mamuntanas, olive-grove, chemical knockdown, 1
female 21.VIII.2009, 2 males and 1 female
6.IX.2009, 1 female 7.IX.2009, 1 male and 3 females 30.IX.2009 leg. Ist. Ent. SS (ISE).
Pelecopsis mengei (Simon, 1884)
HABITAT. Orchard (ined.).
Mannu, Nali farm, orchard, pitfall traps, 1 male and
1 female 11-18.III.1998 leg. Ist. Ent. SS (ISE).
Pityohyphantes phrygianus (C.L. Koch, 1836)
REFERENCES. (10) Costa, 1885 (sub Linyphia
phrygiana).
PROVINCE. Carbonia-Iglesias (10); Sassari (10).
*Pocadicnemis juncea Locket et Millidge, 1953
male 30.V-4.VII.2007 leg. I.S.E. (ISE).
Porrhomma convexum (Westring, 1851)
Prinerigone vagans (Audouin, 1826)
REFERENCES. (13) Garneri, 1902 (sub Erigone
vagans); (84) Pesarini, 1994 (sub Erigone vagans).
PROVINCE. (84); Cagliari (ined.); Medio-Campidano (ined.); Nuoro (ined.); Ogliastra (13).
HABITAT. Citrus-grove (ined.); hazelnut-orchard
(ined.); alder-grove (ined.); mining landfills covered by pioneer plants (ined.).
male 14.XII.2000 leg. CRAS (ISE).
35
MEDIO CAMPIDANO: Guspini, Montevecchio, Piccalinna, m 225, mining landfills covered
by pioneer plants, pitfall traps, 1 male 18.V3.VI.2009 leg. Sassu A., Verdinelli M. (ISE).
NUORO: Belvì, Su Enazzu, m 550, hazelnutorchard, pitfall traps, 3 males 30.V-4.VII.2007
(MBG), 2 males 30.V-4.VII.2007 (ISE) (all leg.
1550, alder-grove, pitfall traps, 1 male 19.VI1.VII.2007 leg. Fiori F. (ISE).
Sintula retroversus (O.P.-Cambridge, 1875)
CHOROTyPE. Turano-European.
PROVINCE. Nuoro (ined.); Ogliastra (64); OlbiaTempio (ined.).
HABITAT. Hypogean habitat (64); cherry-orchard (ined.); hazelnut-orchard (ined.); walnut-orchard (ined.).
m 683, cherry-orchard, 2 males 19-26.XI.2007, 1 female 22-29.II.2008 (ISE); Canale e' Figu, m 576,
walnut-orchard, 1 male 26.XI-3.XII.2007, 1 male
19-26.XI.2007, 1 male 11.I-18.I.2008, 1 male 2229.II.2008 (ISE); Mannaritzò, m 706, cherry-orchard, 1 female 11-18.IV.2007 (ISE); Su Enazzu, m
550, hazelnut-orchard, 1 female 21.II-19.III.2007
(ISE) (all pitfall traps and leg. I.S.E.). Tonara, Su
Azzime, m 682, walnut-orchard, pitfall traps, 3
males 19-26.XI.2007 leg. I.S.E. (ISE).
OLBIA-TEMPIO: Monte Limbara, Punta Balisteri, m 1310, 1 female 10.XII.2004 leg. Lodovici
O., Pantini P. (MBG).
*Syedra nigrotibialis Simon, 1884
HABITAT. Walnut-orchard (ined.); under Juniperus sp. (ined.).
EXAMINED MATERIAL. NUORO: Belvì, Canale e'
Figu, m 576, walnut-orchard, 1 female 2128.VI.2007, 1 female 17-24.VII.2007 leg. I.S.E.
(MBG). Desulo, Gennargentu, mountain refuge, m
1500, alder-grove, 1 female 17-23.VIII.2006 (ISE);
under Juniperus sp., 2 females 2-17.VIII.2006
(ISE), 1 male 9-15.I.2007 (MBG) (all pitfall traps
and leg. Fiori F.).
36
Tenuiphantes flavipes (Blackwall, 1854)
HABITAT. Cherry-orchard (ined.); hazelnut-orchard (127, ined.); under Juniperus sp. (ined.).
EXAMINED MATERIAL. NUORO: Belvì, Antonitzo, m 683, cherry-orchard, pitfall traps, 1 male
16-23.V.2007 leg. I.S.E. (MBG); Su Enazzu, m 550,
hazelnut-orchard, beating tray, 1 male 4.V.2005 leg.
I.S.E. (ISE). Desulo, Gennargentu, mountain refuge, m 1500, under Juniperus sp., pitfall traps, 1
female 10-24.V.2007 leg. Fiori F. (ISE).
Tenuiphantes herbicola (Simon, 1884)
HABITAT. Mediterranean maquis (ined.); hazelnut-orchard (ined.); walnut-orchard (ined.); aldergrove (ined.).
maquis, pitfall traps, 1 male and 1 female 721.IV.2009 (MBG), 2 females 21.IV-5.V.2009, 2
females 7-21.IV.2009, 1 female 5-19.V.2009, 2
males, 3 females and 1 juv. 19.V-4.VI.2009, 1 female 4-16.VI.2009, 2 females 16-29.VI.2009, 1
female 29.VI-14.VII.2009, 1 female 28.VII10.VIII.2009, 1 male 10-25.VIII.2009, 1 female
22.IX-6.X.2009, 1 male and 1 female 8-22.IX.2009
leg. Sassu A., Verdinelli M. (ISE). Guspini, Montevecchio, Piccalinna, m 260, mediterranean maquis,
pitfall traps, 2 females 7-21.IV.2009, 2 females
18.V-3.VI.2009, 1 male and 1 female 1530.VI.2009, 3 females 30.VI-13.VII.2009, 2 males
8-22.IX.2009, 2 females 22.IX-6.X.2009 leg. Sassu
A., Verdinelli M. (ISE).
NUORO: Belvì, Su Enazzu, m 550, hazelnut-orchard, pitfall traps, 1 female 5-16.X.2006 (ISE), 1
female 8.I-21.II.2007 (MBG) (all leg. I.S.E.).
alder-grove, pitfall traps, 1 female 19.VII2.VIII.2006, 1 female 17-30.VIII.2006, 1 female
19.VI-1.VII.2007 leg. Fiori F. (ISE). Tonara, Su Azzime, m 682, walnut-orchard, pitfall traps, 1 female
11-18.I.2008 leg. I.S.E. (ISE).
Tenuiphantes mengei (Kulczyński, 1887)
REFERENCES. (114) Grill et al., 2005 (sub Lepthyphantes mengei).
PROVINCE. (114).
HABITAT. Low shurbs (114).
Tenuiphantes tenebricola (Wider, 1834)
REFERENCES. (13) Garneri, 1902 (sub Leptyphantes tenebricola).
Tenuiphantes tenuis (Blackwall, 1852)
REFERENCES. (94) Arnò et al., 1998 (sub Lepthyphantes tenuis); (127) Loru et al., 2008; (132) Pantini & Sassu, 2009.
PROVINCE. (127); Cagliari (ined.); Medio-Campidano (ined.); Nuoro (ined.); Olbia-Tempio (ined.);
Sassari (94, 132, ined.).
trees (94); cherry-orchard (ined.); orchard (ined.);
mediterranean maquis (ined.); hazelnut-orchard
(127, ined.); walnut-orchard (ined.); olive-grove
(ined.); alder-grove (ined.); under Juniperus (ined.);
forest of Quercus suber (ined.); vineyard (ined.).
EXAMINED MATERIAL. CAGLIARI: Uta, S’Appassiu, Agris farm, citrus-grove, beating tray, 2 females
19.VI.2000, 1 male 10.VII.2000 leg. CRAS (ISE).
MEDIO CAMPIDANO: Guspini, Montevecchio,
Piccalinna, m 260, mediterranean maquis, pitfall
traps, 1 female 30.VI-13.VII.2009, 1 male 1530.VI.2009, 2 females 3-15.VI.2009 leg. Sassu A.,
NUORO: Belvì, Antonitzò, m 683, cherry-orchard, pitfall traps, 1 female 18-26.IV.2007, 1 male
and 1 female 11-18.IV.2007, 1 female 16-23.V.2007
(ISE); Canale e' Figu, m 576, walnut-orchard, pitfall
traps, 1 female 27.IX-4.X.2007 (ISE); Mannaritzò, m
706, cherry-orchard, pitfall traps, 2 males and 1 female 16-23.V.2007 (ISE); Su Ajone, m 585, hazelnutorchard, beating tray, 1 female 19.V.2005, 2 females
29.VI.2005, 1 female 16.VI.2005 (MBG); Su Enazzu,
m 550, hazelnut-orchard, beating tray, 1 female and
2 juv. 16.VI.2005, 3 females and 1 juv. 29.VI.2005
(MBG); hazelnut-orchard, pitfall traps, 1 female
27.VII-8.VIII.2006, 1 female 8-24.VIII.2006, 1 female 24.VIII-5.IX.2006, 1 female 21.IX-5.X.2006, 2
females 8.I-21.II.2007, 1 male and 2 females 21.II19.III.2007, 1 male and 1 female 19.III-26.IV.2007,
4 females 26.IV-30.V.2007, 3 males, 13 females and
1 juv. 30.V-4.VII.2007, 4 females and 1 juv. 30.V4.VII.2007 (ISE) (all leg. I.S.E.). Desulo, Gennargentu, mountain refuge, m 1500, under Juniperus sp.,
3 males and 1 female 10-24.V.2007, alder-grove, 1
male and 2 females 10-24.V.2007 (ISE) (all pitfall
traps and leg. Fiori F.). Fonni, Gennargentu, mountain
refuge, m 1550, alder-grove, 1 female, 30.VIII12.IX.2006, 11 males and 1 female 5-19.VI.2007, 2
males and 1 female 19.VI-1.VII.2007 (ISE), under
Juniperus sp., 1 female, 2-17.VIII.2006, 3 males and
3 females 5-19.VI.2007, 4 males and 2 females
19.VI-1.VII.2007 (ISE) (all pitfall traps and leg. Fiori
F.). Tiana, m 522, hazelnut-orchard, beating tray, 1
male 16.VI.2005 leg. I.S.E. (ISE). Tonara, Su Azzime, m 682, walnut-orchard, pitfall traps, 1 female
11-18.IV.2007, 1 male and 2 females 21-28.VI.2007,
1 female 24-30.VIII.2007, 1 male 17.VIII.200724.VIII.2007, 1 female 19-26.XI.2007, 1 female 2229.II.2008 leg. I.S.E. (ISE).
31.V.1999, 1 female 28.VI.1999 leg. Ist. Ent. SS
(ISE). Olbia, Isola Tavolara, macchia alta Nord, 1
female 4.XI.1965 leg. Gruppo Isole C.N.R. (MBG).
SASSARI: Sassari, Gioscari, olive-grove, chemical knockdown, 2 females 22.IX.2008 leg. Ist.
olive-grove, beating tray, 1 female 13.V.1993 (ISE);
orchard, beating tray, 1 female 18.III.1998 (ISE);
orchard, pitfall traps, 2 males 11-18.III.1998 (ISE)
(all leg. Ist.Ent.SS). Torralba, vineyard, 1 mal,
1.VII.1995 leg. Ist. Ent. SS (ISE).
Tenuiphantes zimmermanni (Bertaku, 1884)
Tiso vagans (Blackwall, 1834)
37
EXAMINED MATERIAL. NUORO: Belvì, Su Enazzu,
4.V.2005 (MBG); m 550, hazelnut-orchard, pitfall
traps, 4 females 13-27.VII.2006 (ISE), 1 female
27.VII-8.VIII.2006, 5 females 13-27.VII.2006, 1
male and 1 female, 24.VIII-5.IX.2006, 1 female 824.VIII.2006, 2 males and 2 females 16-31.X.2006
(MBG), 2 males 16-31.X.2006 (ISE), 1 female 516.X.2006, 1 male and 2 females 21.XI.20068.I.2007, 9 males and 1 female, 8.I-21.II.2007, 9
males 21.II-19.III.2007, 3 females 19.III-26.IV.2007,
6 males and 2 females, 26.IV-30.V.2007, 3 males and
51 females 30.V-4.VII.2007 (MBG) (all leg. I.S.E.).
Typhochrestus sardus Bosmans, 2008
REFERENCES. (125) Bosmans, 2008.
HABITAT. In litter (125).
*Walckenaeria antica (Wider, 1834)
Enazzu, m 550, hazelnut-orchard, pitfall traps, 1 female 27.VII-8.VIII.2006 (MBG), 1 female 30.V4.VII.2007 (ISE) (all leg. I.S.E.).
*Walckenaeria stylifrons (O.P.-Cambridge, 1875)
HABITAT. Cherry-orchard (ined.).
EXAMINED MATERIAL. NUORO: Belvì, Antonitzò, m 683, cherry-orchard, pitfall traps, 1 male
22-29.II.2008 leg. I.S.E. (ISE).
Family TETRAGNATHIDAE
Meta bourneti Simon, 1922
REFERENCES. (34) Dresco & Hubert, 1969; (41)
38
Brignoli, 1971c; (45) Sanna et al., 1972; (46) Thermes, 1972; (64) Brignoli, 1979d; (69) Cannas,
1972; (84) Pesarini, 1994; (105) Grafitti, 2003;
(119) Lana et al., 2006; (121) Grafitti, 2007a; (122)
Grafitti, 2007b; (126) Grafitti & Merella, 2008.
PROVINCE. (84); Cagliari (46, 69); CarboniaIglesias (41, 45, 46, 64, 105, 119, 126); Nuoro (34,
41); Ogliastra (34, 46); Oristano (122); Sassari (41,
121, ined.).
69, 105, 119, 121, 126).
EXAMINED MATERIAL. SASSARI: Sassari, La
Corte, 1 female 23.XI.2003 (ISE).
Meta menardi (Latreille, 1804)
REFERENCES. (38) Puddu, 1970.
Metellina mengei (Blackwall, 1870)
REFERENCES. (64) Brignoli, 1979d (sub Meta
mengei); (139) Trotta, 2011.
PROVINCE. Carbonia-Iglesias (64, 139).
Metellina merianae (Scopoli, 1763)
REFERENCES. (8) Costa, 1884 (sub Meta merianae); (13) Garneri, 1902 (sub Meta Merianae); (14)
Gozo, 1908 (sub Meta merianae); (19) Roewer,
1953 (sub Meta merianae); (22) Roewer, 1956 (sub
Meta merianae); (30) Cerruti, 1968 (sub Meta merianae); (34) Dresco & Hubert, 1969 (sub Meta
merianae); (38) Puddu, 1970 (sub Meta merianae);
(41) Brignoli, 1971c (sub Meta merianae); (44)
Casale, 1972 (sub Meta merianae); (46) Thermes,
1972 (sub Meta merianae); (47) Bartolo et al.,
1973 (sub Meta merianae); (64) Brignoli, 1979d
(sub Meta merianae); (65) Grafitti, 1979 (sub Meta
merianae); (69) Cannas, 1972 (sub Meta bourneti
merianae); (75) Grafitti, 1984 (sub Meta merianae); (82) Grafitti, 1993 (sub Meta merianae); (83)
Pons, 1993 (sub Meta merianae); (84) Pesarini,
1994 (sub Meta merianae); (121) Grafitti, 2007a
(sub Meta merianae); (126) Grafitti & Merella,
2008; (139) Trotta, 2011.
PROVINCE. (13, 84); Cagliari (13, 46, 69); Carbonia-Iglesias (14, 41, 46, 47, 64, 83, 126, 139); Nuoro
(22, 34, 41, 44, 64, 83); Ogliastra (13, 38, 41, 46);
Sassari (8, 13, 14, 19, 30, 41, 65, 75, 82, 121, 139).
38, 41, 44, 46, 47, 64, 65, 69, 75, 82, 83, 121, 126).
EXAMINED MATERIAL. NUORO: Dorgali, Cave
Su Anzu, 1 male VII.1983 leg. Giachino P.M.
(MBG).
Metellina segmentata (Clerck, 1757)
REFERENCES. (8) Costa, 1884 (sub Meta segmentata); (13) Garneri, 1902 (sub Meta segmentata);
(19) Roewer, 1953 (sub Meta segmentata); (30) Cerruti, 1968 (sub Meta segmentata); (46) Thermes,
1972 (sub Meta segmentata); (84) Pesarini, 1994
(sub Meta segmentata); (127) Loru et al., 2008.
PROVINCE. (84, 127); Cagliari (8, 13); Nuoro
(ined.); Ogliastra (46); Sassari (19, 30, ined.).
HABITAT. Hypogean habitat (19, 30, 46); hazelnut-orchard (127, ined.); olive-grove (ined.).
m 585, hazelnut-orchard, beating tray, 1 female,
21.IX.2005 (MBG), 1 female, 21.IX.2005 (ISE) (all
leg. I.S.E.). Tiana, m 522, hazelnut-orchard, beating
tray, 1 male 4.X.2005 leg. I.S.E. (ISE).
olive-grove, beating tray, 1 male 18.III.1994 leg. Ist.
Ent. SS (ISE).
Pachygnatha clercki Sundevall, 1823
REFERENCES. (10) Costa, 1885; (13) Garneri, 1902.
PROVINCE. Cagliari (13); Oristano (10); Sassari (10).
Pachygnatha degeeri Sundevall, 1830
REFERENCES. (10) Costa, 1885.
PROVINCE. Nuoro (ined.); Sassari (10, ined.).
Enazzu, m 550, hazelnut-orchard, pitfall traps, 3 females 13-27.VII.2006, 1 male and 3 females,
24.VIII-5.IX.2006, 2 males 8-24.VIII.2006, 1 female 5-16.X.2006, 1 female 8.I-21.II.2007 (MBG),
1 male and 3 females, 21.II-19.III.2007 (ISE), 1
male and 2 females, 21.II-19.III.2007, 1 male and 2
juv. 19.III-26.IV.2007, 3 females 26.IV-30.V.2007,
2 females 30.V-4.VI.2007 (MBG) (all leg. I.S.E.).
OLBIA-TEMPIO: Olbia, Isola Molara, 1 male
13.VI.1967 leg. Gruppo Isole C.N.R. (MBG).
Tetragnatha extensa (Linnaeus, 1758)
REFERENCES. (6) Magretti, 1880; (7a) Costa,
1882; (8) Costa, 1884; (13) Garneri, 1902; (84) Pesarini, 1994; (133) Trotta, 2009.
PROVINCE. (84); Cagliari (13); Ogliastra (13);
Olbia-Tempio (133); Oristano (6, 13); Oristano/Nuoro (7a, 8).
Tetragnatha intermedia Kulczyński, 1891
PROVINCE. (136); Cagliari (ined.); Sassari (ined.).
HABITAT. Between rocks (136); citrus-grove
(ined.); orchard (ined.); olive-grove (ined.).
EXAMINED MATERIAL. CAGLIARI: Uta, S’Appassiu, Agris farm, citrus-grove, 1 male 21.IX.2005
leg. CRAS (MBG).
olive-grove, 1 male 21.V.1993 (ISE); olive-grove,
beating tray, 1 male 18.VI.1993, 4 males, 1 female
and 1 juv. 3.VI.1993, 3 males and 2 juv.
25.VI.1993, 1 male and 1 female, 9.VII.1993, 1
male 26.VIII.1993, 1 male and 6 juv. 6.VIII.1993,
1 male 19.VIII.1993 (ISE); orchard, beating tray, 1
male 31.VII.1997 (ISE) (all leg. Ist. Ent. SS).
Tetragnatha montana Simon, 1874
REFERENCES. (13) Garneri, 1902; (127) Loru et
al., 2008.
Ogliastra (13); Olbia-Tempio (ined.); Oristano (13);
Sassari (ined.).
39
EXAMINED MATERIAL. NUORO: Belvì, Scala Cerasia, m 660, 1 male, 1 female 21.V.2008 leg. Pantaleoni R.A. (ISE); Su Ajone, m 585,
hazelnut-orchard, beating tray, 1 male 29.VI.2005
(ISE), 1 male and 8 juv. 29.VI.2005 (MBG), 1 female 1.VI.2005 leg. I.S.E. (ISE) (all leg I.S.E.).
OLBIA-TEMPIO: Telti, Rio Almiddina, 1 male
20.V.2008 leg. Sassu A. (ISE).
chemical knockdown, 2 males and 1 juv. 6.VIII.2008,
2 males 21.VIII.2009 leg. Ist. Ent. SS (ISE).
Tetragnatha nitens (Audouin, 1826)
REFERENCES. (13) Garneri, 1902; (84) Pesarini, 1994.
(13); Ogliastra (13).
REMARKS ON FAMILy TETRAGNATHIDAE. Tetragnatha chrysochlora (Audouin, 1826) cited by de
Carlini (1885) and Garneri (1902) is an unknown
species in our fauna, the citations should be linked
to one of the cited species.
FAMILy ARANEIDAE
Aculepeira armida (Audouin, 1826)
REFERENCES. (10) Costa, 1885 (sub Epeira armida); (11) de Carlini, 1885 (sub Epeira armida);
(13) Garneri, 1902 (sub Epeira armida); (63) Brignoli, 1979c; (84) Pesarini, 1994.
PROVINCE. (63, 84); Carbonia-Iglesias (11, 13);
Oristano (10); Sassari (ined.).
EXAMINED MATERIAL. SASSARI: Sassari, Monte
Bianchinu, 1 female 24.VI.2008 leg. Cossu S. (ISE).
Aculepeira ceropegia (Walckenaer, 1802)
REFERENCES. (10) Costa, 1885 (sub Epeira ceropegia); (63) Brignoli, 1979c; (84) Pesarini, 1994.
40
PROVINCE. (10, 63, 84); Nuoro (ined.).
HABITAT. Grassland (ined.).
EXAMINED MATERIAL. NUORO: Bolotana, Badde
'e Salighes, m 850, grassland, 1 male and 3 females
Agalenatea redii (Scopoli, 1763)
REFERENCES. (10) Costa, 1885 (sub Epeira
Redii); (11) de Carlini, 1885 (sub Epeira redii); (13)
Garneri, 1902 (sub Epeira Redii); (63) Brignoli,
1979c; (84) Pesarini, 1994; (132) Pantini & Sassu,
2009; (139) Trotta, 2011.
PROVINCE. (84); Cagliari (10, 13, 63); CarboniaIglesias (11, 13, 139); Medio Campidano (139);
Ogliastra (13, 63, 139); Olbia-Tempio (ined.); Oristano (10, 13); Sassari (132).
Isola Tavolara, high plateau, 1 male 27.IV.1966 leg.
Gruppo Isole CNR (MBG).
Araneus diadematus Clerck, 1757
REFERENCES. (8) Costa, 1884 (sub Epeira diademata); (13) Garneri, 1902 (sub Epeira diademata); (63) Brignoli, 1979c; (84) Pesarini, 1994;
(139) Trotta, 2011.
PROVINCE. (8, 84); Cagliari (13); Carbonia-Iglesias (139); Ogliastra (63); Sassari (ined.).
EXAMINED MATERIAL. SASSARI: Sorso, Lu Tuvaraggiu, m 25, 1 female 24.XI.2009 leg. Loru L.
(ISE). Torralba, vineyard, 1 male 15.X.1994 leg. Ist.
Ent. SS (ISE).
*Araneus marmoreus Clerck, 1757
Mannu, Nali farm, orchard, beating tray, 1 male
*Araneus quadratus Clerck, 1757
EXAMINED MATERIAL. NUORO: Belvì, Scala
Cerasia, m 660, 1 male 28.VIII.2000 leg. Molinu
A. (ISE).
Araniella alpica (L. Koch, 1869)
REFERENCES. (13) Garneri, 1902 (sub Epeira alpica); (84) Pesarini, 1994.
Araniella cucurbitina (Clerck, 1757)
REFERENCES. (8) Costa, 1884 (sub Epeira curcubitina); (13) Garneri, 1902 (sub Epeira cucurbitina); (84) Pesarini, 1994; (114) Grill et al., 2005;
(127) Loru et al., 2008; (139) Trotta, 2011.
PROVINCE. (84, 114, 127); Carbonia-Iglesias (13,
139); Nuoro (8, 13, ined.); Ogliastra (13); OlbiaTempio (8, 13); Oristano (8, 13); Sassari (ined.).
HABITAT. Low shurbs, grass and shurbland,
shurblands and trees (114); hazelnut-orchard (127,
ined.); grassland (ined.); vineyard (ined.).
Enazzu, m 550, hazelnut-orchard, beating tray, 1 female 1.VI.2005, 1 male 16.VI.2005 (ISE), 1 male
16.VI.2005 (MBG) (all leg. I.S.E.). Bolotana,
Badde e'Salighes, m 850, grassland, 1 male
27.V.2007 leg. Sassu A. (ISE). Tiana, m 522, hazelnut-orchard, beating tray, 1 male 16.VI.2005, 1
male 1.VI.2005, 1 female 29.VI.2005, 1 female
27.VII.2005 leg. I.S.E. (MBG).
8.VII.1995 leg. Ist. Ent. SS (ISE).
Argiope bruennichi (Scopoli, 1772)
REFERENCES. (1) Canestrini & Pavesi, 1868
(sub Nephila fasciata); (2a) Canestrini & Pavesi,
1870 (sub Nephila fasciata); (3) Pavesi, 1876;
(13) Garneri, 1902; (84) Pesarini, 1994; (133)
Trotta, 2009.
PROVINCE. (1, 2a, 84); Cagliari (3, 13); OlbiaTempio (133); Oristano (13); Sassari (ined.).
EXAMINED MATERIAL. SASSARI: Sassari, 1 female 30.IX.2008 leg. Cabula P. (ISE).
Argiope lobata (Pallas, 1772)
REFERENCES. (3) Pavesi, 1876; (5) Pavesi, 1878;
(8) Costa, 1884; (13) Garneri, 1902; (84) Pesarini,
1994; (114) Grill et al., 2005.
PROVINCE. (8, 84, 114); Cagliari (13); CarboniaIglesias (3, 13); Nuoro (ined.); Olbia-Tempio (5, 13).
HABITAT. Agricultural land (114).
EXAMINED MATERIAL. NUORO: Siniscola, Monte
Albo, cant. di S. Anna, m 600, 1 female
27.VII.1999 leg. Fontana P. (MBG).
Argiope trifasciata (Forskål, 1775)
CHOROTyPE. Subcosmopolitan.
REFERENCES. (138) Di Pompeo et al., 2011.
HABITAT. Costal areas (138).
Cercidia prominens (Westring, 1851)
REFERENCES. (13) Garneri, 1902; (84) Pesarini, 1994.
Cyclosa conica (Pallas, 1772)
1902; (84) Pesarini, 1994; (127) Loru et al., 2008.
PROVINCE. (84, 127); Cagliari (13); Nuoro
(ined.); Oristano (8, 13).
Enazzu, m 550, hazelnut-orchard, beating tray, 1 female 4.V.2005 leg. I.S.E. (ISE).
Cyclosa insulana (Costa, 1834)
CHOROTyPE. Afrotropico-Indo-Mediterranean.
REFERENCES. (3) Pavesi, 1876; (5) Pavesi, 1878
(sub Cyrtophora or Cyclosa insulana); (7a) Costa,
1882 (sub Epeira insulana); (8) Costa, 1884 sub Ciclosa insulana (tritubercolata Luc.); (13) Garneri,
1902; (84) Pesarini, 1994; (114) Grill et al., 2005.
41
PROVINCE. (84, 114); Cagliari (3, 13); CarboniaIglesias (7a, 8, 13); Olbia-Tempio (5); Oristano (8,
13); Sassari (ined.).
HABITAT. Agricultural land (114); olive-grove
(ined.); on marsh plants (8).
EXAMINED MATERIAL. SASSARI: Sassari, Gioscari, olive-grove, chemical knockdown, 1 female
4.IX.2008, 1 male 6.VIII.2009 leg. Ist. Ent. SS (ISE).
Cyclosa oculata (Walckenaer, 1802)
REFERENCES. (134) Marras et al., 2011.
PROVINCE. (134); Cagliari (ined.).
HABITAT. Citrus-grove (ined.); orchard (134).
EXAMINED MATERIAL. CAGLIARI: Uta, S’Appassiu, Agris farm, citrus-grove, beating tray, 1 female, 19.VI.2000, 2 males and 1 female,
29.VI.2000, 5 males and 4 females 20.VII.2000, 6
males and 1 female 10.VII.2000, 1 male and 1 female, 31.VII.2000, 1 female 31.VIII.2000, 2 females 10.VIII.2000, 3 males and 1 female
21.VIII.2000, 1 male 21.IX.2000, 1 female
11.IX.2000, 2 females 4.X.2000, 1 female
12.X.2000, leg. CRAS (ISE).
Cyclosa sierrae Simon, 1870
1994; (139) Trotta, 2011.
(139).
Cyrtarachne ixoides (Simon, 1870)
REFERENCES. (129) Colombo & Manunza, 2009.
PROVINCE. Cagliari (ined.); Sassari (129, ined.).
HABITAT. Citrus-grove (ined.); bushes and hedges inside a private garden (129); orchard (ined.);
olive-grove (ined.).
EXAMINED MATERIAL. CAGLIARI: Uta, S’Appassiu, Agris farm, citrus-grove, beating tray, 1 juv
13.XI.2000 leg. CRAS (ISE).
olive-grove, 1 male 1.VII.1993, 1 male 22.VII.1993
42
(ISE); orchard, 1 female 27.V.1998 (ISE) (all beating tray and leg. Ist. Ent. SS).
Cyrtophora citricola (Forskål, 1775)
REFERENCES. (3) Pavesi, 1876; (7a) Costa, 1882
(sub Epeira cajetana); (8) Costa, 1884 (sub C.
opuntiae); (11) de Carlini, 1885; (13) Garneri, 1902
(sub Cyrtaphora citricola); (84) Pesarini, 1994.
PROVINCE. (13, 84); Cagliari (8, 13, ined.); Carbonia-Iglesias (3, 11, 13); Oristano (7a, 8, 13).
HABITAT. On agaves (8).
EXAMINED MATERIAL. CAGLIARI: Villasimius,
around the village, 1 female 25.VI.2010 leg. zucchelli D. (MBG).
Gibbaranea bituberculata (Walckenaer, 1802)
REFERENCES. (8) Costa, 1884 (sub Epeira dromedaria); (13) Garneri, 1902 (sub Epeira dromedaria); (63) Brignoli, 1979c; (84) Pesarini, 1994;
(139) Trotta, 2011.
PROVINCE. (8, 63, 84); Cagliari (13); CarboniaIglesias (139); Ogliastra (13).
Gibbaranea gibbosa (Walckenaer, 1802)
REFERENCES. (10) Costa, 1885 (sub Epeira gibbosa); (84) Pesarini, 1994; (132) Pantini & Sassu,
2009; (139) Trotta, 2011.
PROVINCE. (84); Carbonia-Iglesias (139); Nuoro
(10); Sassari (132).
Gibbaranea omodea (Thorell, 1870)
Hypsosinga albovittata (Westring, 1851)
REFERENCES. (10) Costa, 1885 (sub Singa albovittata); (84) Pesarini, 1994.
PROVINCE. (84); Oristano (10).
Hypsosinga pygmaea (Sundevall, 1831)
REFERENCES. (10) Costa, 1885 (sub Singa pygmaea); (84) Pesarini, 1994.
PROVINCE. (84); Cagliari (10); Oristano (10).
Hypsosinga sanguinea (C.L. Koch, 1844)
REFERENCES. (10) Costa, 1885 (sub Singa sanguinea).
Larinioides cornutus (Clerck, 1757)
REFERENCES. (8) Costa, 1884 (sub Epeira cornuta); (13) Garneri, 1902 (sub Epeira cornuta); (84)
Pesarini, 1994; (139) Trotta, 2011.
PROVINCE. (84); Cagliari (8, 13); Oristano (139).
Larinioides patagiatus (Clerck, 1757)
REFERENCES. (10) Costa, 1885 (sub Epeira patagiata (dumetorum hbn.).); (13) Garneri, 1902 (sub
Epeira patagiata); (84) Pesarini, 1994.
PROVINCE. (84); Cagliari (13); Oristano (10, 13).
Larinioides suspicax (O.P.-Cambridge, 1876)
REFERENCES. (70) Grasshoff, 1983 (sub L. folium); (116) Helsdingen, 2005.
PROVINCE. Nuoro (116); Olbia-Tempio (70).
HABITAT. In the centre of the city (116).
Leviellus kochi (Thorell, 1870)
REFERENCES.(41) Brignoli, 1971c (sub Zygiella
kochi); (46) Thermes, 1972 (sub Zygiella kochi);
(52) Puddu & Pirodda, 1974 (sub Zygiella kochi);
(63) Brignoli, 1979c (sub Zygiella kochi); (83)
Pons, 1993 (sub Zygiella kochi); (84) Pesarini, 1994
(sub Zygiella kochi).
PROVINCE. (63, 84); Carbonia-Iglesias (41, 46,
52, 83).
Mangora acalypha (Walckenaer, 1802)
REFERENCES. (10) Costa, 1885 (sub Epeira acalypha); (13) Garneri, 1902 (sub Epeira acalypha);
(63) Brignoli, 1979c; (84) Pesarini, 1994; (114) Grill
et al., 2005; (127) Loru et al., 2008; (132) Pantini &
Sassu, 2009; (133) Trotta, 2009; (139) Trotta, 2011.
PROVINCE. (84, 114, 127); Cagliari (13, 63); Carbonia-Iglesias (10, 13, 139); Medio Campidano
(139); Nuoro (10, 63, ined.); Ogliastra (13, 63);
Olbia-Tempio (10, 133); Oristano (10, 63); Sassari
(63, 132, ined.).
HABITAT. Grass and shurbland, shurblands and
trees, agricultural land (114); hazelnut-orchard
(127, ined.).
hazelnut-orchard, beating tray, 1 female 16.VI.2005,
1 male 1.VI.2005 leg. I.S.E. (ISE).
OLBIA-TEMPIO: Olbia, Isola Tavolara, macchia
alta, 2 males, 2 females and 11 juv. 24.IV.1966 leg.
Gruppo Isole CNR (MBG).
male and 1 female 22.V.2008 leg. Sassu A. (ISE).
Neoscona adianta (Walckenaer, 1802)
REFERENCES. (5) Pavesi, 1878 (sub Epeira
adianta); (8) Costa, 1884 (sub Epeira adianta); (13)
Garneri, 1902 (sub Epeira adianta); (63) Brignoli,
1979c; (84) Pesarini, 1994; (114) Grill et al., 2005;
PROVINCE. (84, 114); Cagliari (13, 63); Carbonia-Iglesias (13, 139); Medio Campidano (139);
Nuoro (63, 139); Ogliastra (63); Olbia-Tempio (5,
13, ined.); Oristano (13, 63); Oristano/Nuoro (8);
HABITAT. Low shurbs, grass and shurbland
(114); vineyard (ined.).
EXAMINED MATERIAL. OLBIA-TEMPIO: Telti, Rio
Almiddina, 1 male 20.V.2008 leg. Sassu A. (ISE).
SASSARI: Torralba, vineyard, 2 juv.
21.V.1994, 1 female and 1 juv. 4.VI.1994, 1 fe-
43
male 17.VI.1995, 1 male 24.VI.1995 leg. Ist. Ent.
SS (ISE).
Neoscona subfusca (C.L. Koch, 1837)
REFERENCES. (10) Costa, 1885 (sub Epeira dalmatica); (63) Brignoli, 1979c (sub "Araneus" subfuscus); (127) Loru et al., 2008; (139) Trotta, 2011.
PROVINCE. (63, 127); Cagliari (10); CarboniaIglesias (139); Nuoro (139, ined.); Olbia-Tempio
(ined.); Sassari (ined.).
HABITAT. Hazelnut-orchard (127, ined.); olivegrove (ined.); forest of Quercus suber (ined.); vineyard (ined.).
13.VII.2005, leg. I.S.E. (ISE).
forest of Quercus suber, beating tray, 1 female
16.VII.1999, leg. Ist. Ent. SS (ISE).
chemical knockdown, 1 male, 13.X.2009, leg. Ist.
olive-grove, beating tray, 1 male 25.VI.1993, 1 male
1.VII.1993 leg. Ist. Ent. SS (ISE). Torralba, vineyard, 1 female 16.VII.1994, 1 male 2.VII.1994, 1
male 15.VII.1995, leg. Ist. Ent. SS (ISE); vineyard,
beating tray, 1 male 6.VIII.1994 leg. Pirisi (ISE).
Parazygiella montana (C.L. Koch, 1834)
REFERENCES. (13) Garneri, 1902 (sub Zilla montana); (63) Brignoli, 1979c (sub Zygiella montana);
(84) Pesarini, 1994 (sub Zygiella montana).
PROVINCE. (63, 84); Nuoro/Ogliastra (13).
Singa hamata (Clerck, 1757)
1979c; (84) Pesarini, 1994.
PROVINCE. (63, 84); Cagliari (13); CarboniaIglesias (13).
Zilla diodia (Walckenaer, 1802)
44
REFERENCES. (10) Costa, 1885 (sub Epeira dioidia); (13) Garneri, 1902 (sub Epeira diodia); (63)
Brignoli, 1979c; (84) Pesarini, 1994; (114) Grill et
al., 2005; (127) Loru et al., 2008.
PROVINCE. (84, 114, 127); Cagliari (13, 63); Carbonia-Iglesias (10, 13); Nuoro (ined.); Ogliastra
(13); Olbia-Tempio (10); Sassari (ined.).
HABITAT. Orchard (ined.); low shrubs (14); hazelnut-orchard (127, ined.).
male 19.V.2005 leg. I.S.E. (ISE).
SASSARI: Sassari, Platamona, at sea level, 1 female 22.V.2008 leg. Sassu A. (ISE). Sorso, Pianu
Mannu, Nali farm, orchard, beating tray, 1 male
22.IV.1998 leg. Ist. Ent. SS (ISE).
Zygiella atrica (C.L. Koch, 1845)
REFERENCES. (10) Costa, 1885 (sub Zilla atrica);
(11) de Carlini, 1885 (sub Zilla atrica); (13) Garneri, 1902 (sub Zilla atrica); (63) Brignoli, 1979c;
PROVINCE. (63, 84); Cagliari (10); CarboniaIglesias (11, 13); Ogliastra (13).
Zygiella x-notata (Clerck, 1757)
REFERENCES. (13) Garneri, 1902 (sub Zilla x-notata); (41) Brignoli, 1971c; (46) Thermes, 1972;
(63) Brignoli, 1979c; (72) Puddu,1983; (83) Pons,
1993; (84) Pesarini, 1994; (94) Arnò et al., 1998;
(127) Loru et al., 2008; (132) Pantini & Sassu,
2009; (139) Trotta, 2011.
PROVINCE. (63, 84, 127); Cagliari (ined.); Carbonia-Iglesias (41, 139); Medio Campidano (139);
Nuoro (ined.); Ogliastra (13, 46, 72); Sassari (83,
94, 132, ined.).
trees (94); hypogean habitat (41, 46, 72, 83);
cherry-orchard (ined.); orchard (ined.); inside house
(ined.); in orto (ined.); hazelnut-orchard (127,
ined.); olive-grove (ined.); vineyard (ined.).
EXAMINED MATERIAL. CAGLIARI: Uta, S’Appassiu, Agris farm, citrus-grove, beating tray, 2 males
31.VIII.2000, 1 male and 1 female 21.IX.2000, 4 fe-
males 4.X.2000, 1 female 12.X.2000, 6 females
23.X.2000, 1 female 2.XI.2000, 2 females
23.XI.2000, 2 females 4.XII.2000 leg. CRAS (ISE).
NUORO: Belvì, Antonitzò, m 683, cherry-orchard, 1 female 4.X-26.XI.2007, 1 female 2229.II.2008 (ISE); Su Enazzu, m 550,
hazelnut-orchard, 1 female 8-15.XI.2004 (MBG)
(all in bark traps and leg. I.S.E.). Tiana, m 522,
hazelnut-orchard, beating tray, 1 female
21.IX.2005, 1 male 4.X.2005, leg. I.S.E. (ISE).
SASSARI: Alghero, inside house, 1 male
21.XII.1999 leg. Pantaleoni R.A. (ISE); Mamuntanas, olive-grove, chemical knockdown, 1 male
6.IX.2009, 1 male 7.X.2009 leg. Ist. Ent. SS (ISE).
Osilo, in a vegetable garden, 1 male 20.X.1999
(ISE). Sassari, Li Punti, Baldinca, m 110, inside
house, 1 female 23.V.2008 leg. Sassu A. (ISE); Lu
male 29.IX.2009 leg. Ist. Ent. SS (ISE). Sorso,
Pianu Mannu, Nali farm, orchard, bark traps, 1 female 2-9.XII.1997 leg. Ist. Ent. SS (ISE). Torralba,
vineyard, 1 male 9.IX.1995, 1 female 7.X.1995 leg.
Ist. Ent. SS (ISE); vineyard, beating tray, 2 males
30.IX.1995 leg. Pirisi (ISE).
REMARKS ON FAMILy ARANEIDAE. Singa aussereri Thorell, 1873 cited by Costa (1884, sub S. rufula); Garneri (1902, sub S. rufula); Brignoli
(1979c, sub Hypsosinga aussereri) e Pesarini (1994,
sub Hypsosinga aussereri) was considered a species
inquirenda by Thaler & Knoflach (2003) and therefore excluded from the catalogue. Singa simoniana Costa, 1885 described by Costa in the
Campidano Cagliari area is a species inquirenda
(Brignoli, 1979: 27) and therefore excluded from
the catalogue.
Family LyCOSIDAE
Allocosa oculata (Simon, 1876)
REFERENCES. (13) Garneri, 1902 (sub Lycosa
oculata).
Alopecosa accentuata (Latreille, 1817)
REFERENCES. (13) Garneri, 1902 (sub Lycosa accentuata); (84) Pesarini, 1994.
PROVINCE. (84); Nuoro/Ogliastra (13).
Alopecosa albofasciata (Brullé, 1832)
REFERENCES. (10) Costa, 1885 [sub Lycosa albofasciata (sagittata K.)]; (13) Garneri, 1902 (sub
Lycosa albofasciata); (20) Krauss, 1955 (sub Xerolycosa albofasciata); (36) Lugetti & Tongiorgi,
1969; (84) Pesarini, 1994; (132) Pantini & Sassu,
2009; (139) Trotta, 2011.
PROVINCE. - (20, 84); Cagliari (10, 13, 36); Carbonia-Iglesias (10, 13, 139); Medio-Campidano
(ined.); Nuoro (36, ined.); Nuoro/Ogliastra (36);
Ogliastra (13); Olbia-Tempio (36); Sassari (36, 132).
HABITAT. Cherry-orchard (ined.); garrigue
(ined.); mediterranean maquis (ined.); hazelnut-orchard (ined.); walnut-orchard (ined.).
maquis, 1 male 19.V-4.VI.2009. (ISE); m 200, garrigue, 1 male 5-19.V.2009 (ISE) (all pitfall traps
and leg. Sassu A., Verdinelli M.). Guspini, Montevecchio, Piccalinna, m 235, garrigue, pitfall traps,
2 males 21.IV-5.V.2009, 3 males 5-18.V.2009, 1
male and 1 female 18.V-3.VI.2009 leg. Sassu A.,
NUORO: Belvì, Antonitzò, m 683, cherry-orchard, 1 male 16-23.V.2007 (ISE); Canale e' Figu,
m 576, walnut-orchard, 1 male 26.IV-4.V.2007, 4
males and 1 female 16-23.V.2007, 2 females 2128.VI.2007 (ISE); Mannaritzò, m 706, cherry-orchard, 3 males and 3 females 18-26.IV.2007, 4
males and 6 females 11-18.IV.2007, 11 males 1623.V.2007 (ISE); Su Enazzu, m 550, hazelnut-orchard, 6 males and 2 females 19.III-26.IV.2007
(MBG), 2 males and 1 female, 19.III-26.IV.2007
(ISE), 17 males and 1 female, 30.V-4.VII.2007
(MBG) (all pitfall traps and leg. I.S.E.). Tonara, Su
Azzime, m 682, walnut-orchard, pitfall traps, 3
males 16-23.V.2007 leg. I.S.E. (ISE).
Alopecosa fabrilis (Clerck, 1757)
REFERENCES. (13) Garneri, 1902 (sub Lycosa fabrilis); (84) Pesarini, 1994.
45
Ogliastra (13); Oristano (13).
EXAMINED MATERIAL. NUORO: Fonni, Gennargentu, mountain refuge, m 1550, under Juniperus
sp., pitfall traps, 1 male 26.IX-10.X.2006 leg.
Fiori F. (ISE).
Alopecosa laciniosa (Simon, 1876)
REFERENCES. (13) Garneri, 1902 (sub Lycosa laciniosa); (84) Pesarini, 1994.
REMARKS. The presence of this species in Italy
requires further confirmation as stated by Lugetti &
Tongiorgi (1969: 76).
Alopecosa pulverulenta (Clerck, 1757)
REFERENCES. (6) Magretti, 1880 (sub Tarentula
pulverulenta); (10) Costa, 1885 [sub Lycosa pulverulenta (cuineata, K.)]; (13) Garneri, 1902 (sub Lycosa pulverulenta); (84) Pesarini, 1994.
(13); Ogliastra (13); Oristano (6, 13).
Arctosa cinerea (Fabricius, 1777)
REFERENCES. (8) Costa, 1884 (sub Lycosa cinerea); (13) Garneri, 1902 (sub Lycosa cinerea); (20)
Krauss, 1955; (84) Pesarini, 1994; (132) Pantini &
Sassu, 2009.
PROVINCE. (8, 20, 84); Cagliari (13); CarboniaIglesias (13); Ogliastra (13); Oristano (13); Sassari (132).
HABITAT. Common on the banks of rivers (8).
Arctosa fulvolineata (Lucas, 1846)
CHOROTyPE. European-Mediterranean.
REFERENCES. (84) Pesarini, 1994.
PROVINCE. (84).
Arctosa lacustris (Simon, 1876)
46
REFERENCES. (24) Lugetti & Tongiorgi, 1965;
(84) Pesarini, 1994; (139) Trotta, 2011.
PROVINCE. (84); Cagliari (24); Medio Campidano (139).
Arctosa leopardus (Sundevall, 1833)
REFERENCES. (10) Costa, 1885 (sub Lycosa leopardus); (13) Garneri, 1902 (sub Lycosa leopardus); (20) Krauss, 1955 (sub Megarctosa
leopardus); (24) Lugetti & Tongiorgi, 1965; (84)
Pesarini, 1994.
PROVINCE. (20, 24, 84); Cagliari (13);
Cagliari/Oristano (13); Carbonia-Iglesias (10);
Ogliastra (13).
Arctosa perita (Latreille, 1799)
REFERENCES. (13) Garneri, 1902 (sub Lycosa perita); (84) Pesarini, 1994; (117) Buchar et al., 2006.
(13); Ogliastra (13); Olbia-Tempio (117).
Arctosa personata (L. Koch, 1872)
REFERENCES. (10) Costa, 1885 (sub Lycosa personata).
PROVINCE. Cagliari (10); Sassari (10).
Arctosa similis Schenkel, 1938
REFERENCES. (117) Buchar et al., 2006; (132)
Pantini & Sassu, 2009; (139) Trotta, 2011.
PROVINCE. Cagliari (117); Carbonia-Iglesias
(139); Sassari (132).
HABITAT. Riverbank with gravel (117).
Arctosa variana C.L. Koch, 1847
REFERENCES. (24) Lugetti & Tongiorgi, 1965;
(24) Lugetti & Tongiorgi, 1965 (sub A. perita
latitorax); (84) Pesarini, 1994; (117) Buchar et
al., 2006.
PROVINCE. (24, 84); Cagliari (24); Medio Campidano (117).
HABITAT. On muddy shore (117).
Aulonia albimana (Walckenaer, 1805)
Hogna radiata (Latreille, 1817)
REFERENCES. (3) Pavesi, 1876 (sub Tarentula radiata var. liguriensis); (8) Costa, 1884 (sub Lycosa
radiata); (11) de Carlini, 1885 (sub Tarentula radiata); (13) Garneri, 1902 (sub Lycosa balearica);
(20) Krauss, 1955; (41) Brignoli, 1971c (sub Lycosa
radiata); (84) Pesarini, 1994; (132) Pantini &
Sassu, 2009; (133) Trotta, 2009.
PROVINCE. (20, 84); Cagliari (13); CarboniaIglesias (3, 11, 13); Medio-Campidano (ined.);
Nuoro (8, 13, ined.); Olbia-Tempio (13, 41); Oristano (13); Sassari (132, 133, ined.).
HABITAT. Hypogean habitat (41); cherry-orchard
(ined.); garrigue (ined.); mediterranean maquis
(ined.); hazelnut-orchard (ined.); walnut-orchard
(ined.); under Juniperus sp. (ined.).
maquis, 1 male and 1 female 28.VII-10.VIII.2009,
1 male 10-25.VIII.2009 (ISE); m 200, garrigue, 1
female 10-25.VIII.2009 (ISE) (all pitfall traps and
leg. Sassu A., Verdinelli M.). Guspini, Montevecchio, Piccalinna, m 235, garrigue, pitfall traps, 1
male 25.VIII-8.IX.2009, 3 males 10-25.VIII.2009
leg. Sassu A., Verdinelli M. (ISE).
NUORO: Belvì, Canale e' Figu, m 576, walnutorchard, 1 male 24-30.VIII.2007 (ISE); Mannaritzò,
m 706, cherry-orchard, 1 male 24-30.VIII.2007
(ISE); Su Enazzu, m 550, hazelnut-orchard, 2 females 27.VII-8.VIII.2006 (ISE), 1 male, 1327.VII.2006, 1 male 27.VII-8.VIII.2006, 2 males
8-24.VIII.2006 (MBG) (all pitfall traps and leg.
1550, under Juniperus sp., pitfall traps, 1 male
19.VII-2.VIII.2006 leg. Fiori F. (ISE). Tonara, Su
Azzime, m 682, walnut-orchard, pitfall traps, 1 male
and 1 female, 17-24.VII.2007, leg. I.S.E. (ISE).
OLBIA-TEMPIO: Olbia, Isola Tavolara, 1 female 24.II.1966 leg. Gruppo Isole C.N.R. (MBG).
Lycosa narbonensis Walckenaer, 1806
REFERENCES. (13) Garneri, 1902; (46) Thermes,
1972.
Pardosa cribrata Simon, 1876
REFERENCES. (26) Tongiorgi, 1966; (84) Pesarini, 1994.
PROVINCE. (26, 84).
Pardosa gefsana Roewer, 1959
REFERENCES. (26) Tongiorgi, 1966 (sub P. naevia); (80) Tongiorgi & Sola, 1991 (sub P. nigristernis); (84) Pesarini, 1994.
PROVINCE. (84); Cagliari (80); Medio Campidano (26); Oristano (80).
HABITAT. Edge of brackish pond (80); salt
marshes (80).
REMARKS. The citation of P. naevia around the
Guspini area (Tongiorgi, 1966) is later corrected in
P. nigristernis (Tongiorgi & Sola, 1991: 241). Pesarini reports P. naevia in his Checklist maybe because he was not aware of the publication or maybe
deceived by a printing error. On page 241 Tongiorgi
& Sola’s article states in fact: “A revision of the Italian Wolf Spiders of the Genus Pardosa by Tongiorgi (1966) had assigned, with reserve, three
female specimens of a species collected in Sardinia
to Pardosa naevia … thus, our specimens are now
more correctly assigned to the closely related Pardosa naevia Denis, 1966 (figure 1).” It seems evident
(even from the subtitle of the figure) that it is actually
assigned to P. nigristernis later synonimised with P.
gefsana (Alderweireldt & Jocqué, 1992)
Pardosa hortensis (Thorell, 1872)
47
REFERENCES. (6) Magretti, 1880 (sub Lycosa
hortensis); (13) Garneri, 1902.
PROVINCE. Cagliari (13); Medio Campidano (6);
Nuoro (ined.); Oristano (6, 13).
HABITAT. Alder-grove (ined.); under Juniperus
sp. (ined.).
EXAMINED MATERIAL. NUORO: Desulo, Gennargentu, mountain refuge, m 1500, alder-grove, pitfall
traps, 2 males 11-25.VII.2006 leg. Fiori F. (ISE).
under Juniperus sp., pitfall traps, 1 male 19.VI1.VII.2007 leg. Fiori F. (ISE).
Pardosa nigra (C.L. Koch, 1834)
Pardosa occidentalis Simon, 1881
REFERENCES. (53) Tongiorgi, 1974.
PROVINCE. Medio Campidano (53).
Pardosa prativaga (L. Koch, 1870)
REFERENCES. (13) Garneri, 1902 (sub P. fervida).
PROVINCE. Cagliari (13); Nuoro (ined.).
male 19.III-26.IV.2007 leg. I.S.E. (ISE).
Pardosa proxima (C.L. Koch, 1847)
REFERENCES. (5) Pavesi, 1878 (sub Lycosa proxima); (8) Costa, 1884; (13) Garneri, 1902; (20)
Sassu, 2009.
PROVINCE. (20, 84); Cagliari (8, 13); CarboniaIglesias (13); Medio-Campidano (ined.); Nuoro
(ined.); Olbia-Tempio (5, 13); Sassari (132).
(ined.); alder-grove (ined.).
48
Guspini, Montevecchio, Piccalinna, m 235, garrigue, pitfall traps, 1 male and 1 female 18.V3.VI.2009 leg. Sassu A., Verdinelli M. (ISE).
NUORO: Belvì, Antonitzò, m 683, cherry-orchard, 2 males and 2 females, 11-18.IV.2007 (ISE);
Canale e' Figu, m 576, walnut-orchard, 1 female
17-24.VII.2007 (ISE); Mannaritzò, m 706, cherryorchard, 1 female 11-18.IV.2007, 6 males and 4 females 21-28.VI.2007, 3 males and 1 female,
17-24.VII.2007, 1 male 24-30.VIII.2007, 6 males
and 3 females 22-29.II.2008 (ISE); Su Enazzu, m
550, hazelnut-orchard, 3 females 27.VII8.VIII.2006, 1 male and 4 females, 1327.VII.2006, 5 males 21.II-19.III.2007, 1 male and
1 female, 19.III-26.IV.2007, 1 male and 3 females,
26.IV-30.V.2007, 1 female 30.V-4.VII.2007
(MBG), 2 males and 3 females, 30.V-4.VII.2007
(ISE) (all pitfall traps and leg. I.S.E.). Fonni, Gennargentu, mountain refuge, m 1550, alder-grove,
pitfall traps, 1 female 19.VII-2.VIII.2006, 1 female
17-30.VIII.2006 leg. Fiori F. (ISE).
REMARKS. P. proxima is easily confused with
the similar species P. vlijmi known in France and
recently found in Northern Italy. The separation of
the two species is very clear when considering
courtship displays while from a morphological
point of view, only the males are identifiable (Isaia
in litteris). While awaiting the results of current research, all specimens examined have been assigned
to P. proxima.
Pardosa pullata (Clerck, 1757)
Pardosa tatarica (Thorell, 1875)
REFERENCES. (13) Garneri, 1902 (sub P. strigillata); (26) Tongiorgi, 1966 (sub P. strigillata); (84)
Pesarini, 1994 (sub Pardosa strigillata).
PROVINCE. (84); Cagliari (13, 26); CarboniaIglesias (13); Ogliastra (13); Sassari (26).
REMARKS. Buchar & Thaler (2002: 446) consider this species and P. strigillata as synonyms of P.
atomaria.
Pirata knorri (Scopoli, 1763)
Pirata latitans (Blackwall, 1841)
REFERENCES. (13) Garneri, 1902; (139) Trotta,
2011.
PROVINCE. Carbonia-Iglesias (139); Nuoro
(ined.); Ogliastra (13).
male 30.V-4.VII.2007 leg. I.S.E. (ISE).
Pirata piraticus (Clerck, 1757)
1902; (20) Krauss, 1955 (sub P. piratica); (139)
Trotta, 2011.
PROVINCE. Cagliari (ined.); Carbonia-Iglesias
(13, 139); Oristano (10); Sassari (20).
EXAMINED MATERIAL. CAGLIARI: Pula, m 715, 1
female 18.VIII.2006 leg. Ruzzante Wilkin J. (MBG).
Pirata piscatorius (Clerck, 1757)
1902.
Oristano (13).
*Pirata tenuitarsis Simon, 1876
HABITAT. Alder-grove (ined.).
EXAMINED MATERIAL. NUORO: Desulo, Gennargentu, mountain refuge, m 1500, alder-grove, pitfall
traps, 2 females 11-25.VII.2006, 1 female 25.VII9.VIII.2006 leg. Fiori F. (ISE). Fonni, Gennargentu,
mountain refuge, m 1550, alder-grove, pitfall traps,
1 female 17-30.VIII.2006, 1 female 2-17.VIII.2006,
3 males 19.VI-1.VII.2007 leg. Fiori F. (ISE).
Trochosa hispanica Simon, 1870
REFERENCES. (13) Garneri, 1902 (sub Lycosa caliginosa); (20) Krauss, 1955; (84) Pesarini, 1994.
PROVINCE. (20, 84); Carbonia-Iglesias (13);
Nuoro (ined.).
HABITAT. Cherry-orchard (ined.); hazelnut-orchard (ined.).
m 683, cherry-orchard, 1 male 11-18.IV.2007, 1 male
18-26.IV.2007 (ISE); Su Enazzu, m 550, hazelnut-orchard, 3 males, 1 female and 1 juv. 24.VIII-5.IX.2006
(MBG), 2 males 8-24.VIII.2006 (ISE), 3 males, 1 female and 1 juv. 5-21.IX.2006, 4 males and 1 juv.
21.IX-5.X.2006, 1 male and 1 female 5-16.X.2006, 1
male 16-31.X.2006, 1 male 21.II-19.III.2007, 71
males and 2 females 19.III-26.IV.2007, 26 males
26.IV-30.V.2007 (MBG), 13 males and 1 female
30.V-4.VII.2007 (ISE), 86 males 30.V-4.VII.2007
(MBG) (all pitfall traps and leg. I.S.E.).
Trochosa ruricola (De Geer, 1778)
REFERENCES. (13) Garneri, 1902 (sub Lycosa ruricola).
PROVINCE. Cagliari (13); Nuoro/Ogliastra (13);
Oristano (13).
Trochosa terricola Thorell, 1856
REFERENCES. (13) Garneri, 1902 (sub Lycosa
terricola).
PROVINCE. Cagliari (13); Cagliari/Oristano (13);
Ogliastra (13).
Xerolycosa nemoralis (Westring, 1861)
REFERENCES. (13) Garneri, 1902 (sub Lycosa nemoralis).
PROVINCE. Cagliari (13); Cagliari/Oristano (13);
Ogliastra (13).
REMARKS ON FAMILy LyCOSIDAE. Pardosa agricola (Thorell, 1856) is cited in Sardinia by Garneri
49
(1902). The presence of this species in Italy is very
doubtful, Tongiorgi (1966: 284) has never seen Italian specimens of this species and there have not
been recent reports. It is possible that old reports are
due to mistaken identification of other species of
Pardosa of the monticola group. It was decided
therefore not to include this species in the catalogue.
Family PISAURIDAE
Dolomedes plantarius (Clerck, 1757)
REFERENCES. (20) Krauss, 1955 (sub D. clercki);
Pisaura mirabilis (Clerck, 1757)
REFERENCES. (8) Costa, 1884 (sub Ocyale mirabilis); (13) Garneri, 1902 (sub Ocyale mirabilis);
(20) Krauss, 1955; (59) Brignoli, 1977; (74) Brignoli, 1984 (sub "Form A"); (84) Pesarini, 1994;
(127) Loru et al., 2008; (139) Trotta, 2011.
PROVINCE. (13, 20, 84, 127); Cagliari (59, 74);
Carbonia-Iglesias (59, 139); Medio Campidano (59,
139); Nuoro (59, ined.); Ogliastra (59); Olbia-Tempio (59); Oristano/Nuoro (8); Sassari (59, ined.).
HABITAT. Orchard (ined.); hazelnut-orchard
(127, ined.).
Enazzu, m 550, hazelnut-orchard, beating tray, 1 female 16.VI.2005 leg. I.S.E. (ISE).
OLBIA-TEMPIO: Loiri-Porto San Paolo, Porto
S. Paolo, 1 juv. 27.XI.1966 leg. Gruppo Isole
C.N.R. (MBG). Olbia, Isola Tavolara, 1 juv.
24.IV.1966 leg. Gruppo Isole C.N.R. (MBG).
SASSARI: Sorso, Pianu Mannu, Nali farm, orchard, pitfall traps, 1 male 15-22.IV.1998 leg. Ist.
Ent. SS (ISE).
Pisaura quadrilineata (Lucas, 1838)
REFERENCES. (74) Brignoli, 1984 (sub "Form
D"); (84) Pesarini, 1994 (sub P. maderiana); (132)
PROVINCE. (84); Ogliastra (74); Sassari (132).
50
Family OXyOPIDAE
Oxyopes heterophthalmus (Latreille, 1804)
REFERENCES. (8) Costa, 1884; (11) de Carlini,
1885; (13) Garneri, 1902; (59) Brignoli, 1977; (84)
Pesarini, 1994; (114) Grill et al., 2005; (132) Pantini
& Sassu, 2009; (139) Trotta, 2011.
PROVINCE. (8, 84, 114); Cagliari (13); CarboniaIglesias (11, 13, 139); Medio Campidano (139);
Ogliastra (13); Olbia-Tempio (59); Oristano (13);
HABITAT. Low shurbs, shurblands and trees, agricultural land (114).
EXAMINED MATERIAL. OLBIA-TEMPIO: LoiriPorto San Paolo, Porto S. Paolo, 2 males 1.V.1964
leg. Gruppo Isole C.N.R. (MBG).
Oxyopes lineatus Latreille, 1806
1902; (84) Pesarini, 1994.
PROVINCE. (8, 84); Cagliari (13); Nuoro (ined.).
Enazzu, m 550, hazelnut-orchard, pitfall traps, 1 female 27.VII-8.VIII.2006 leg. I.S.E. (ISE).
Oxyopes nigripalpis Kulczyński, 1891
REFERENCES. (127) Loru et al., 2008; (139)
Trotta, 2011.
PROVINCE. (127); Carbonia-Iglesias (139);
Nuoro (ined.); Sassari (ined.).
Enazzu, m 550, hazelnut-orchard, beating tray, 2 females 27.VII.2005 leg. I.S.E. (ISE).
male 22.V.2008 leg. Sassu A. (ISE).
Family zOROPSIDAE
Zoropsis media Simon, 1878
Zoropsis spinimana (Dufour, 1820)
REFERENCES. (8) Costa, 1884 (sub Z. ocreata);
(13) Garneri, 1902 (sub Z. ocreata); (84) Pesarini,
1994; (94) Arnò et al., 1998; (95) Thaler & Knoflach, 1998; (127) Loru et al., 2008; (133) Trotta,
2009; (139) Trotta, 2011.
PROVINCE. (84, 127); Cagliari (13, 139); Carbonia-Iglesias (13, 139); Medio-Campidano (ined.);
Nuoro (95, ined.); Olbia-Tempio (133); Oristano (8,
13); Sassari (94, ined.).
HABITAT. Avenue of lime trees (94); orchard
(ined.); inside house (ined.); mediterranean maquis
(ined.); hazelnut-orchard (127, ined.).
Guspini, Montevecchio, Piccalinna, m 260, mediterranean maquis, pitfall traps, 1 male 21.IV-5.V.2009,
1 male and 2 females 18.V-3.VI.2009, 1 female 1530.VI.2009 leg. Sassu A., Verdinelli M. (ISE).
NUORO: Belvì, Su Ajone, m 585, hazelnut-orchard, bark traps, 1 female 16-23.VI.2005, 1 juv.
14.VII.2005, 1 female 21-29.IX.2005, 1 female
29.IX-6.X.2005 (MBG), 1 male 29.IX-6.X.2005
(ISE), 2 males 6-12.X.2005. (MBG) (all leg. I.S.E.).
Tiana, m 522, hazelnut-orchard, bark traps, 1 female 17-26.VIII.2005 leg. I.S.E. (ISE).
SASSARI: Sassari, inside house, 1 male,
6.II.2008, 1 female I.2004 leg. Sassu A. (ISE).
Sorso, Pianu Mannu, Nali farm, orchard, bark
traps, 1 male and 1 female 24.XI-2.XII.1997 leg.
Ist. Ent. SS (ISE).
Family zORIDAE
*Zora manicata Simon, 1878
CHOROTyPE. Europeo-Mediterranean.
HABITAT. Walnut-orchard (ined.).
EXAMINED MATERIAL. NUORO: Belvì, Canale e’
Figu, m 576, walnut-orchard, pitfall traps, 1 male
26.IV-4.V.2007 leg. I.S.E. (ISE).
Zora spinimana (Sundevall, 1833)
REFERENCES. (20) Krauss, 1955.
PROVINCE. (20).
Family AGELENIDAE
Agelena labyrinthica (Clerck, 1757)
1902; (20) Krauss, 1955; (84) Pesarini, 1994; (139)
Trotta, 2011.
PROVINCE. (20, 84); Cagliari (13); CarboniaIglesias (13, 139); Oristano (8, 13).
Allagelena gracilens (C.L. Koch, 1841)
REFERENCES. (13) Garneri, 1902 (sub Agelena
similis); (84) Pesarini, 1994.
Aterigena soriculata (Simon, 1873)
REFERENCES. (13) Garneri, 1902 (sub Tegenaria
soriculata); (84) Pesarini, 1994 (sub Tegenaria soriculata).
PROVINCE. (13, 84); Nuoro/Ogliastra (13); Sassari (20).
REMARKS. The presence in Sardinia of this species
is doubtful, the citation by Krauss (1955) concerns
T. parietina (Bolzern et al., 20: 776).
Lycosoides coarctata (Dufour, 1831)
REFERENCES. (3) Pavesi, 1876 (sub Textrix coarctata); (6) Magretti, 1880 (sub Textrix coarctata);
(10) Costa, 1885 (sub Textrix coarctata); (11) de
Carlini, 1885 (sub Textrix coactata); (13) Garneri,
1902 (sub Textrix Coarctata); (20) Krauss, 1955
(sub Textrix Coarctata); (37) Argiolas et al., 1970
(sub Textrix coarctata); (59) Brignoli, 1977 (sub
Textrix coarctata); (84) Pesarini, 1994; (132) Pantini & Sassu, 2009.
PROVINCE. (84); Cagliari (6, 13, 37, 59, ined.);
Carbonia-Iglesias (3, 6, 11, 13, 59); Medio Campi-
51
dano (ined.); Olbia-Tempio (59); Oristano (10, 13,
ined.); Sassari (6, 13, 20, 59, 132, ined.).
HABITAT. Hypogean habitat (20, 37); orchard
(ined.); garrigue (ined.); garden (ined.); inside
house (ined.); mining landfills covered by pioneer
plants (ined.).
Maria, Punta su Pisu, m 14, 2 males and 2 females
XII.2006 leg. Ruzzante (MBG).
plants, 2 males 6-20.X.2009 (ISE); m 200, garrigue,
1 male 6-20.X.2009 (MBG) (all pitfall traps and
ORISTANO: Bonarcado, residential area, m 283,
garden, 1 male 25.X.2008 leg. Sassu A. (ISE).
OLBIA-TEMPIO: Olbia, Isola Molara, 8 female
25.II.1966, 1 female, 20.IV.1966, 11 females,
25.XI.1966 (MBG); Isola Tavolara, 2 females,
26.XI.1966 (MBG) (all leg. Gruppo Isole C.N.R.).
SASSARI: Alghero, 1 male, leg. Pantaleoni R.A.
(ISE). Ozieri, 1 female 21.III.1978 leg. Bianchi R.,
Moretti G. (MBG). Sassari, inside house, 1 male
9.X.2007 leg. Cesaroni C. (ISE). Sorso, Pianu
Mannu, Nali farm, orchard, pitfall traps, 1 male 2027.X.1997 (ISE); orchard, bark traps, 1 female 29.XII.1997 (ISE) (all leg. Ist. Ent. SS).
Malthonica campestris (C.L. Koch, 1834)
REFERENCES. (13) Garneri, 1902 (sub Tegenaria
campestris); (84) Pesarini, 1994 (sub Tegenaria
campestris).
PROVINCE. (84); Nuoro/Ogliastra (13).
Malthonica dalmatica (Kulczyński, 1906)
CHOROTyPE. Turano-European-Mediterranean.
REFERENCES. (20) Krauss, 1955 (sub Tegenaria
pagana); (39) Brignoli, 1971a (sub Tegenaria drescoi); (84) Pesarini, 1994 (sub Tegenaria drescoi);
(87) Wunderlich, 1995b (sub Tegenaria henroti);
(124) Bolzern et al., 2008.
PROVINCE. (84); Nuoro (ined.); Ogliastra (87,
124); Sassari (20, 39, 124).
HABITAT. Hypogean habitat (20, 39, 124); hazelnut-orchard (ined.).
52
male 10-31.X.2006 leg. I.S.E. (ISE).
Malthonica eleonorae Brignoli, 1974
REFERENCES. (39) Brignoli, 1971a (sub Tegenaria henroti); (51) Brignoli, 1974 (sub Tegenaria
eleonorae); (59) Brignoli, 1977 (sub Tegenaria
eleonorae); (83) Pons, 1993 (sub Tegenaria eleanorae); (84) Pesarini, 1994 (sub Tegenaria eleonorae); (124) Bolzern et al., 2008.
PROVINCE. (84); Carbonia-Iglesias (39, 51, 59,
83, 124); Ogliastra (59, 124).
Malthonica pagana (C.L. Koch, 1840)
REFERENCES. (6) Magretti, 1880 (sub Tegenaria
pagana); (13) Garneri, 1902 (sub Tegenaria pagana); (30) Cerruti, 1968 (sub Tegenaria pagana);
(39) Brignoli, 1971a (sub Tegenaria pagana); (59)
Brignoli, 1977 (sub Tegenaria pagana); (84) Pesarini, 1994 (sub Tegenaria pagana).
PROVINCE. (84); Cagliari (6, 13, 39, 59); Sassari
(30, 39).
Malthonica sardoa Brignoli, 1977
REFERENCES. (59) Brignoli, 1977; (83) Pons,
1993; (84) Pesarini, 1994; (124) Bolzern et al.,
2008.
PROVINCE. (124); (84); Carbonia-Iglesias (59,
124); Medio Campidano (ined.); Nuoro (59, 83, 124,
ined.); Olbia-Tempio (59, 124); Oristano (59, 124).
HABITAT. Hypogean habitat (83, 124); garrigue
(ined.); mediterranean maquis (ined.); walnut-orchard (ined.); open Quercus forest with mosses
(124); Quercus pubescens, humid (124); under Juniperus sp. (ined.); mining landfills covered by pioneer plants (ined.).
maquis, 1 female 19.V-4.VI.2009, 1 female 416.VI.2009, 1 female 14-28.VII.2009, 2 females
28.VII-10.VIII.2009 (ISE); m 170, mining landfills
covered by pioneer plants, 1 female 19.V-4.VI.2009
(ISE); m 200, garrigue, 1 female 5-19.V.2009 (ISE),
1 female 19.V-4.VI.2009 (MBG) (all pitfall traps
and leg. Sassu A., Verdinelli M.). Guspini, Montevecchio, Piccalinna, m 225, mining landfills covered by pioneer plants, 1 female 15-30.VI.2009
(ISE); m 235, garrigue, 1 female 18.V-3.VI.2009, 2
females 30.VI-13.VII.2009 (ISE); m 260, mediterranean maquis, 1 female 30.VI-13.VII.2009 (ISE),
1 female 13-27.VII.2009 (MBG) (all pitfall traps
and leg. Sassu A., Verdinelli M.).
NUORO: Fonni, Gennargentu, mountain refuge,
m 1550, under Juniperus sp., pitfall traps, 1 female
19.VI-1.VII.2007 leg. Fiori F. (ISE). Tonara, Su Azzime, m 682, walnut-orchard, pitfall traps 2 males
22-29.II.2008 leg. I.S.E. (ISE).
Malthonica sicana Brignoli, 1976
REFERENCES. (124) Bolzern et al., 2008.
HABITAT. Open Quercus forest, rich leaf litter
(124); Quercus forest open, not wet (124).
Tegenaria agrestis (Walckenaer, 1802)
REFERENCES. (14) Gozo, 1908; (59) Brignoli,
1977; (84) Pesarini, 1994.
PROVINCE. (84); Carbonia-Iglesias (14); Sassari
(59).
Tegenaria armigera Simon, 1873
REFERENCES. (19) Roewer, 1953; (30) Cerruti,
1968; (39) Brignoli, 1971a; (84) Pesarini, 1994.
PROVINCE. (84); Ogliastra (39); Sassari (19, 30).
Tegenaria domestica (Clerck, 1757)
1902; (30) Cerruti, 1968; (84) Pesarini, 1994.
PROVINCE. (84); Cagliari (13); Nuoro (30); Sassari (8, 13).
Tegenaria henroti Dresco, 1956
REFERENCES. (21) Dresco, 1956; (22) Roewer,
1956 sub T. domestica (Clerck, 1757); (30) Cerruti,
1968; (51) Brignoli, 1974; (52) Puddu & Pirodda,
1974; (83) Pons, 1993; (84) Pesarini, 1994; (124)
Bolzern et al., 2008.
PROVINCE. (124); (84); Carbonia-Iglesias (52);
Nuoro (21, 22, 30, 51, 83, 124); Ogliastra (124).
83, 124).
Tegenaria parietina (Fourcroy, 1785)
REFERENCES. (13) Garneri, 1902; (20) Krauss,
1955; (37) Argiolas et al., 1970; (38) Puddu, 1970;
(39) Brignoli, 1971a; (42) Puddu,1971; (43) Bartolo
et al., 1972; (46) Thermes, 1972; (84) Pesarini,
1994; (139) Trotta, 2011.
PROVINCE. (84); Cagliari (13, 37, 42, 46); Carbonia-Iglesias (39, 43); Medio Campidano (139);
Ogliastra (38); Olbia-Tempio (20); Sassari (39).
HABITAT. Hypogean habitat (37, 38, 39, 42, 43, 46).
Textrix caudata L. Koch, 1872
REFERENCES. (5) Pavesi, 1878 (sub T. violantis); (13) Garneri, 1902 (sub T. violantis); (84) Pesarini, 1994.
PROVINCE. (84); Cagliari (ined.); Medio Campidano (ined.); Olbia-Tempio (5, 13).
HABITAT. Mediterranean maquis (ined.).
EXAMINED MATERIAL. CAGLIARI: Pula, Is Cannoneris, m 715, 1 female 18.VIII.2006 leg. Ruzzante-Wilkins (MBG).
traps, 2 males 18.V-3.VI.2009 (MBG), 2 males
18.V-3.VI.2009, 6 males and 1 female 315.VI.2009 (ISE), 3 males 3-15.VI.2009, 3 males
30.VI-13.VII.2009 (MBG), 5 males and 1 female
53
15-30.VI.2009 (ISE), 2 males and 1 female 1530.VI.2009 (MBG), 6 males and 2 females 30.VI13.VII.2009, 1 female 27.VII-10.VIII.2009 (ISE),
1 male 13-27.VII.2009 (MBG), 1 male 1327.VII.2009 (ISE) (all leg. Sassu A., Verdinelli M.).
Textrix denticulata (Olivier, 1789)
1994.
PROVINCE. (84); Nuoro/Ogliastra (13); Ogliastra
(13).
*Textrix pinicola Simon, 1875
Guspini, Montevecchio, Piccalinna, m 260, mediterranean maquis, pitfall traps, 1 male 18.V-3.VI.2009
(ISE), 1 male 18.V-3.VI.2009 (MBG), 4 males 1530.VI.2009, 15 males 3-15.VI.2009 (ISE), 7 males
15-30.VI.2009, 1 male 15-30.VI.2009 (MBG), 7
males 30.VI-13.VII.2009 (ISE), 3 males 315.VI.2009, 1 male 13-27.VII.2009 (MBG) (all leg.
Family HAHNIIDAE
Hahnia candida Simon, 1875
REFERENCES. (48) Brignoli, 1973.
PROVINCE. Nuoro (48), Medio Campidano
(ined.).
HABITAT. Mining landfills covered by pioneer
plants (ined.).
EXAMINED MATERIAL. MEDIO CAMPIDANO: Guspini, Montevecchio, Piccalinna, m 260, mining landfills covered by pioneer plants, pitfall traps, 1 female
21.IV-5.V.2009 leg. Sassu A., Verdinelli M. (MBG).
Family DICTyNIDAE
Argenna patula (Simon, 1874)
54
Dictyna arundinacea (Linnaeus, 1758)
PROVINCE. Ogliastra (13); Oristano (13).
Dictyna civica (Lucas, 1850)
REFERENCES. (94) Arnò et al., 1998 (sub Brigittea civica).
PROVINCE. Olbia-Tempio (ined.); Sassari (94).
HABITAT. Avenue of lime trees (94); forest of
EXAMINED MATERIAL. OLBIA-TEMPIO: Buddusò, near Nuraghe Loelle, forest of Quercus
suber, beating tray, 2 females 7.VI.1999 leg. Ist.
Ent. SS (ISE).
Dictyna latens (Fabricius, 1775)
PROVINCE. Cagliari (13); Sassari (ined.).
Mannu, Nali farm, olive-grove, beating tray, 1 female 29.IV.1993 leg. Ist. Ent. SS (ISE).
Lathys humilis (Blackwall, 1855)
REFERENCES. (94) Arnò et al., 1998; (127) Loru
et al., 2008.
HABITAT. Avenue of lime trees (94); cherry-orchard (ined.); hazelnut-orchard (127, ined.); forest
of Quercus suber (ined.).
m 683, cherry-orchard, 1 female, 11-18.IV.2007
(ISE); Su Ajone, m 585, hazelnut-orchard, 1 female
25.V-1.VI.2005 (ISE); Su Enazzu, m 550, hazelnutorchard, 1 female 13-20.X.2004 (ISE) (all bark traps
and leg. I.S.E.).
Marilynia bicolor (Simon, 1870)
HABITAT. In debris on the sandydeposits at the
mouth of the Rio Girasole (116).
Nigma puella (Simon, 1870)
REFERENCES. (10) Costa, 1885 (sub Dictyna
puella); (13) Garneri, 1902 (sub Dictyna puella);
(116) Helsdingen, 2005; (127) Loru et al., 2008.
PROVINCE. (127); Cagliari (10, 13); Nuoro (116,
ined.); Ogliastra (116); Olbia-Tempio (ined.); Oristano (ined.); Sassari (ined.).
HABITAT. In shurbs along a little stream (116);
hazelnut-orchard (127, ined.); olive-grove (ined.);
forest of Quercus suber (ined.); vineyard (ined.).
m 585, hazelnut-orchard, 1 male 16.VI.2005 (MBG),
1 female 13.VII.2005 (ISE); Su Enazzu, m 550, hazelnut-orchard, beating tray, 1 male 19.V.2005 (ISE),
3 males and 1 female, 19.V.2005 leg. I.S.E. (MBG)
(all beating tray and leg. I.S.E.). Tiana, m 522, hazelnut-orchard, beating tray, 1 male 19.V.2005, 4
males and 2 females, 1.VI.2005, 1 female,
13.VII.2005, 1 female and 1 juv., 26.VIII.2005 leg.
I.S.E. (MBG).
OLBIA-TEMPIO: Buddusò, near Nuraghe Loelle, forest of Quercus suber, beating tray, 1 female,
31.V.1999, 1 male 24.V.1999 leg. Ist. Ent. SS (ISE).
ORISTANO: Fondorgianus, Tirso river, m 35, 1
male 22.IV.2003 leg. Pantini P., Magnati F. (MBG).
SASSARI: Bonnannaro, Cannisones, 2 males
17.V.2002, 2 males 28.V.2002 leg. Nuvoli T.
(ISE). Sorso, Pianu Mannu, Nali farm, olivegrove, 1 female 23.III.1993 leg. Ist. Ent. SS (ISE).
Torralba, vineyard, 1 female 27.V.1995 leg. Ist.
Ent. SS (ISE).
Nigma walckenaeri (Roewer, 1951)
REFERENCES. (10) Costa, 1885 (sub Dictyna viridissima); (13) Garneri, 1902 (sub Dictyna vividissima); (94) Arnò et al., 1998; (116) Helsdingen, 2005.
PROVINCE. Cagliari (13, ined.); Nuoro (10);
Ogliastra (116); Sassari (94, ined.).
trees (94); on trees lining a street (ined.); in the valley of the rio (116).
male 2.XI.2000, 5 males 13.XI.2000, 1 male and 1
female 4.XII.2000 leg. CRAS (ISE).
SASSARI: Sassari, on trees lining a street, 1 female 29.IX.1994 leg. Ist. Ent. SS (ISE).
Family AMAUROBIIDAE
Amaurobius erberi (Keyserling, 1863)
REFERENCES. (41) Brignoli, 1971c; (52) Puddu
& Pirodda, 1974; (139) Trotta, 2011.
PROVINCE. Carbonia-Iglesias (41, 52, 139); Sassari (ined.).
HABITAT. Hypogean habitat (41, 52); orchard
(ined.).
Mannu, Nali farm, orchard, bark traps, 1 female
20-27.X.1997 leg. Ist. Ent. SS (ISE).
Amaurobius fenestralis (Ström, 1768)
PROVINCE. Cagliari (13); Oristano (13).
Amaurobius ferox (Walckenaer, 1830)
Family TITANOECIDAE
Nurscia albomaculata (Lucas, 1846)
55
REFERENCES. (10) Costa, 1885 (sub Titanoeca
albomaculata); (13) Garneri, 1902 (sub Titanoeca
albo-maculata); (84) Pesarini, 1994 (sub Titanoeca
albomaculata); (94) Arnò et al., 1998 (sub Titanoeca albomaculata); (133) Trotta, 2009.
Ogliastra (13); Olbia-Tempio (10, 133); Oristano
(13); Sassari (94, ined.).
HABITAT. Avenue of lime trees (94); cherry-orchard (ined.); orchard (ined.).
m 683, cherry-orchard, pitfall traps, 1 male, 1 female,
17-24.VII.2007, leg. I.S.E. (ISE).
SASSARI: Sorso, Pianu Mannu, Nali farm, orchard, pitfall traps, 1 male 26.VI-2.VII.1997 leg. Ist.
Ent. SS (ISE).
Family MITURGIDAE
Cheiracanthium angulitarse Simon, 1878
PROVINCE. Olbia-Tempio (ined.); Sassari (132).
Murta Maria, light trap, 1 male 17.V.1994 leg. Comotti G., Pantini P. (MBG).
Cheiracanthium mildei L. Koch, 1864
1902; (94) Arnò et al., 1998; (116) Helsdingen, 2005;
PROVINCE. (ined.); Cagliari (13, ined.); Carbonia-Iglesias (139); Medio-Campidano (139, ined.);
Nuoro (10); Ogliastra (116); Oristano (10, ined.);
trees (94); orchard (ined.); inside house (ined.);
mediterranean maquis (ined.); olive-grove (ined.);
vineyard (ined.); wasteland plot (116).
males 19.VI.2000, 1 male and 1 female 29.VI.2000,
2 males 20.VII.2000, 1 female 31.VII.2000, 1 female 10.VIII.2000 leg. CRAS (ISE).
MEDIO CAMPIDANO: Arbus, Ingurtosu, Nara-
56
cauli, m 120, mediterranean maquis, pitfall traps, 1
female 14-28.VII.2009, 1 female 10-25.VIII.2009
leg. Sassu A., Verdinelli M. (ISE). Guspini, Montevecchio, Piccalinna, m 260, mediterranean maquis,
pitfall traps, 1 male 15-30.VI.2009 leg. Sassu A.,
ORISTANO: Bonarcado, inside house, 1 female
6.VI.2007 leg. Sassu M. (ISE).
olive-grove, 1 male 21.V.1993 (ISE); olive-grove,
beating tray, 2 males and 2 females 9.VII.1993, 1
male 1.VII.1993, 1 female 22.VII.1993, 1 female
26.VIII.1993 (ISE); orchard, bark traps, 1 female
10-17.VII.1997 (ISE); orchard, beating tray, 1 male
20.X.1997 (ISE) (all leg. Ist.Ent.SS). Torralba, vineyard, 1 female 13.V.1995, 1 male 24.V.1995, 1 male
Cheiracanthium pelasgicum (C.L. Koch, 1837)
PROVINCE. Ogliastra (13); Oristano (10).
Cheiracanthium pennyi O. P.-Cambridge, 1873
Cheiracanthium punctorium (Villers, 1789)
CHOROTyPE.Turano-European.
REFERENCES. (13) Garneri, 1902 (sub Chiracanthiun nutrix).
PROVINCE. Cagliari (13); Nuoro/Ogliastra (13).
Cheiracanthium seidlitzi L. Koch, 1864
1902; (18) Simon, 1932; (84) Pesarini, 1994.
PROVINCE. (18, 84); Cagliari (13); Oristano (10).
Cheiracanthium striolatum Simon, 1878
REFERENCES. (10) Costa, 1885; (114) Grill et
al., 2005.
HABITAT. Shurblands and trees (114).
Family ANyPHAENIDAE
Anyphaena accentuata (Walckenaer, 1802)
PROVINCE. (84, 127); Nuoro (8, 13, ined.).
HABITAT. Cherry-orchard (ined.); hazelnut-orchard (127, ined.).
EXAMINED MATERIAL. NUORO: Belvì, Mannaritzò, m 706, cherry-orchard, bark traps, 1 male 1826.IV.2007 (ISE); Su Ajone, m 585,
hazelnut-orchard, beating tray, 9 males, 2 females
and 1 juv. 4.V.2005 (MBG), 1 male 1 female,
4.V.2005, 1 male and 2 females 19.V.2005 (ISE), 5
females, 19.V.2005, 1 male 1.VI.2005 (MBG); Su
Enazzu, m 550, hazelnut-orchard, bark traps, 1 female 15-22.III.2005, 1 female 27.IV-4.V.2005, 1 female 4-11.V.2005 (MBG); m 550, hazelnut-orchard,
beating tray, 2 males and 2 females 4.V.2005, 2
males, 5 females and 1 juv. 19.V.2005, 1 female and
2 juv. 16.VI.2005, 3 females 1.VI.2005, 1 female
29.VI.2005, 2 females and 40 juv. 10.VIII.2005
(MBG) (all leg. Ist. Ent. SS).
Anyphaena sabina L. Koch, 1866
1994; (94) Arnò et al., 1998; (127) Loru et al., 2008;
PROVINCE. (84, 127); Cagliari (13, 139); Carbonia-Iglesias (139); Medio-Campidano (139, ined.);
Nuoro (ined.); Oristano (ined.); Sassari (94, 132,
ined.).
(ined.); mediterranean maquis (ined.); hazelnut-orchard (127, ined.); olive-grove (ined.); artichoke
field (ined.).
maquis, pitfall traps, 1 male 19.V-4.VI.2009 leg.
Sassu A., Verdinelli M. (ISE). Guspini, Montevecchio, Piccalinna, m 260, mediterranean maquis, pit-
fall traps, 1 female 15-30.VI.2009 leg. Sassu A.,
NUORO: Belvì, Su Enazzu, m 550, hazelnut-orchard, beating tray, 1 female 4.V.2005 leg. I.S.E.
(MBG). Tiana, m 522, hazelnut-orchard, beating
tray, 1 female 4.V.2005 (MBG), 1 female 1.VI.2005
(ISE); hazelnut-orchard, bark traps, 1 female 411.V.2005 (MBG) (all leg. Ist. Ent. SS).
ORISTANO: Bosa, artichoke field, 1 female
4.III.2009 leg. Padiglia P. (ISE).
OLBIA-TEMPIO: Olbia, Isola Tavolara, penisolotto, 1 male 23.IV.1966 leg. Gruppo Isole C.N.R.
(MBG).
olive-grove, beating tray, 1 male, 23.III.1993 (ISE),
olive-grove, 1 female 6.V.1993 (ISE); orchard, beating tray, 1 male and 1 female 22.IV.1998 (ISE) (all
leg. Ist. Ent. SS).
Family LIOCRANIDAE
*Agraecina lineata (Simon, 1878)
HABITAT. Garrigue (ined.).
Guspini, Montevecchio, Piccalinna, m 235, garrigue, pitfall traps, 1 male 18.V-3.VI.2009, 1 male 315.VI.2009 (ISE), 1 male 30.VI-13.VII.2009
(MBG), 1 male 15-30.VI.2009 leg. Sassu A., Verdinelli M. (ISE).
Agroeca annulipes Simon, 1878
REFERENCES. (18) Simon, 1932.
PROVINCE. (18).
Agroeca inopina O.P.-Cambridge, 1886
CHOROTyPE.European.
PROVINCE. Carbonia-Iglesias (139); Nuoro (ined.).
HABITAT. Alder-grove (ined.); cherry-orchard
(ined.); hazelnut-orchard (ined.); under Juniperus
sp. (ined.); walnut-orchard (ined.).
EXAMINED MATERIAL. NUORO: Belvì, Antonitzò, m 683, cherry-orchard, 1 female 16-
57
23.V.2007, 1 female 21-28.VI.2007, 1 male 1118.I.2008 (ISE); Canale e' Figu, m 576, walnut-orchard, 1 female 21-28.VI.2007 (ISE); Mannaritzò,
m 706, cherry-orchard, 1 male 19-26.XI.2007
(ISE); Su Enazzu, m 550, hazelnut-orchard, 1 female 13-27.VII.2006, 1 female 27.VII-8.VIII.2006,
4 juv. 8-24.VIII.2006, 6 males 24.VIII-5.IX.2006, 1
male and 1 female 5-21.IX.2006, 7 males and 1 female 21.IX-5.X.2006, 2 males 5-16.X.2006, 3 males
31.X-21.XI.2006 (MBG), 1 male and 1 female 516.X.2006 (ISE), 3 males 16-31.X.2006, 4 males and
1 female, 21.XI.2006-8.I.2007 (MBG), 1 male and
1 female 21.XI.2006-8.I.2007 (ISE), 1 female 8.I21.II.2007, 1 female 21.II-19.III.2007, 2 females
19.III-26.IV.2007, 3 females 26.IV-30.V.2007, 3 females 30.V-4.VII.2007 (MBG) (all pitfall traps and
leg. I.S.E.). Desulo, Gennargentu, mountain refuge,
m 1500, under Juniperus sp., 2 males 19.IX3.X.2006 (ISE); alder-grove, 1 male, 5-19.IX.2006,
1 male, 3-17.X.2006 (ISE) (all pitfall traps and leg.
Fiori F.). Fonni, Gennargentu, mountain refuge, m
1550, alder-grove, 1 female, 19.VII-2.VIII.2006, 1
male, 17-30.VIII.2006, 6 males and 2 females 217.VIII.2006, 14 males and 1 female 30.VIII12.IX.2006, 11 males 12-26.IX.2006, 1 female
29.XI.2006-9.I.2007 (ISE); under Juniperus sp. 1
male 17-30.VIII.2006, 3 males 30.VIII-12.IX.2006,
2 females 15-29.XI.2006, 1 female 5-19.VI.2007
(ISE) (all pitfall traps and leg. Fiori F.). Tonara, Su
Azzime, m 682, walnut-orchard, pitfall traps, 1 female 11-18.IV.2007, 1 male 4-11.X.2007, 1 male 1926.XI.2007, 1 male 11-18.I.2008 leg. I.S.E. (ISE).
Cybaeodes sardus Platnick et Di Franco, 1992
REFERENCES. (81) Platnick & Di Franco, 1992.
Liocranum giersbergi Kraus, 1955
REFERENCES. (20) Krauss, 1955; (46) Thermes,
1972; (52) Puddu & Pirodda, 1974; (60) Brignoli &
Gaddini, 1979; (64) Brignoli, 1979d; (84) Pesarini,
1994; (87) Wunderlich, 1995b; (127) Loru et al.,
2008; (139) Trotta, 2011.
PROVINCE. (84, 127); Cagliari (46, 52, 139,
ined.); Carbonia-Iglesias (139); Medio Campidano
(139, ined.); Nuoro (60, 64, ined.); Nuoro/Ogliastra
(87); Ogliastra (87); Sassari (20).
58
HABITAT. Hypogean habitat (46, 52, 64);
cherry-orchard (ined.); mediterranean maquis
(ined.); hazelnut-orchard (127, ined.); walnut-orchard (ined.); under the bark (87).
EXAMINED MATERIAL. CAGLIARI: Pula, Is Cannoneris, m 715, 1 female 18.VIII.2006, 1 female
1.XII.2006 leg. Ruzzante-Wilkins (MBG).
MEDIO CAMPIDANO: Arbus, Ingurtosu, Naracauli, m 120, mediterranean maquis, pitfall traps, 1
male 19.V-4.VI.2009, 1 male 4-16.VI.2009 (ISE), 1
male 16-29.VI.2009, 2 females 29.VI-14.VII.2009
(MBG), 1 female 16-29.VI.2009, 1 female 29.VI14.VII.2009, 1 female 14-28.VII.2009, 1 male 822.IX.2009, 1 male 6-20.X.2009 (ISE) (all leg.
Sassu A., Verdinelli M.). Guspini, Montevecchio,
traps, 1 female 7-21.IV.2009, 3 females 518.V.2009, 2 females 30.VI-13.VII.2009, 2 females
15-30.VI.2009, 1 female 13-27.VII.2009 (ISE), 1
male 22.IX-6.X.2009 (MBG), 1 male 8-22.IX.2009,
1 male, 1 female 22.IX-6.X.2009, 1 male 620.X.2009 (ISE) (all leg. Sassu A., Verdinelli M.).
NUORO: Belvì, Mannaritzò, m 706, cherry-orchard, pitfall traps, 1 female 17-24.VII.2007 leg.
I.S.E. (ISE). Tiana, m 522, hazelnut-orchard, bark
traps, 1 female 11-18.V.2005 leg. I.S.E. (ISE). Tonara, light trap, 1 male 8.IX.2006 leg. Fiori F. (ISE);
Su Azzime, m 682, walnut-orchard, pitfall traps, 1
female 17-24.VIII.2007 leg. I.S.E. (ISE).
Liocranum rupicola (Walckenaer, 1830)
REFERENCES. (13) Garneri, 1902; (77) Grimm,
1986; (139) Trotta, 2011.
PROVINCE. (13); Nuoro (77); Ogliastra (139).
HABITAT. Under bark (77).
Mesiotelus tenuissimus (L. Koch, 1866)
1994.
Scotina celans (Blackwall, 1841)
PROVINCE. Carbonia-Iglesias (139); Nuoro
(ined.).
HABITAT. Hazelnut-orchard (ined.); walnut-orchard (ined.).
male 30.X-21.XI.2006 leg. I.S.E. (MBG). Tonara,
Su Azzime, m 682, walnut-orchard, 1 male 2229.II.2008 leg. I.S.E. (MBG).
Family CLUBIONIDAE
Clubiona comta C.L. Koch, 1839
Trotta, 2011.
PROVINCE. (127); Carbonia-Iglesias (139); Medio
Campidano (ined.); Nuoro (ined.); Sassari (ined.).
HABITAT. Orchard (ined.); garrigue (ined.); hazelnut-orchard (127, ined.); walnut-orchard (ined.).
Guspini, Montevecchio, Piccalinna, m 235, garrigue, pitfall traps, 1 female 30.VI-13.VII.2009 leg.
NUORO: Belvì, Canale e' Figu, m 576, walnutorchard, 2 females 28.V-4.VI.2007 (ISE); Su
Enazzu, m 550, hazelnut-orchard, 1 female, 27.IV4.V.2005 1 female 4-11.V.2005, 1 female 714.VII.2005, 1 female 14-21.IX.2005 (ISE) (all
bark traps and leg. I.S.E.).
SASSARI: Sorso, Pianu Mannu, Nali farm, orchard, pitfall traps, 1 female 24-31.VII.1997 leg.
Ist. Ent. SS (ISE).
Clubiona genevensis L. Koch, 1866
Clubiona leucaspis Simon, 1932
Trotta, 2011.
PROVINCE. (127); Cagliari (ined.); Carbonia-
Iglesias (139); Nuoro (ined.); Olbia-Tempio (ined.);
Sassari (ined.).
(ined.); orchard (ined.); hazelnut-orchard (127,
ined.); walnut-orchard (ined.); olive-grove (ined.);
pinewood (ined.); forest of Quercus suber (ined.);
vineyard (ined.).
EXAMINED MATERIAL. CAGLIARI: Uta, S’Appassiu, Agris farm, citrus-grove, beating tray, 1 female 29.VI.2000, 1 female 10.VII.2000, 1 female
20.VII.2000 leg. CRAS (ISE).
NUORO: Belvì, Antonitzò, m 683, cherry-orchard, bark traps, 5 males, 11-18.IV.2007, 4 females, 16-24.V.2007, 1 female, 21-28.VI.2007
(ISE); Canale e' Figu, m 576, walnut-orchard,
bark traps, 1 female, 26.IV-4.V.2007, 2 females 1724.VII.2007, 1 male 19-26.XI.2007 (ISE); Mannaritzò, m 706, cherry-orchard, bark traps, 6 males and
1 female 18-26.IV.2007, 3 females 16-23.V.2007, 4
males 11.I-18.I.2008, 1 male 22-29.II.2008 (ISE);
Su Ajone, m 585, hazelnut-orchard, bark traps, 1
male 27.IV-4.V.2005 (ISE), 1 female 4-11.V.2005,
3 females 25.V-1.VI.2005 (MBG), 1 female 25.V1.VI.2005 (ISE) 2 females 14-19.VII.2005 (MBG);
Su Ajone, m 585, hazelnut-orchard, beating tray, 1
female 1.VI.2005, 1 female, 19.V.2005 (MBG); Su
Enazzu, m 550, hazelnut-orchard, bark traps, 1 female 13-20.X.2004, 3 females 8-15.XI.2004, 1 male
27.IV-4.V.2005, 1 female 25.V-1.VI.2005, 2 females
18-25.V.2005, 1 female, 29.VI-6.VII.2005, 1 female
16-23.VI.2005, 2 females 1-8.VI.2005 (MBG); hazelnut-orchard, beating tray, 1 male, 3 females and
1 juv. 19.V.2005, 1 female 1.VI.2005 (MBG) (all
leg. I.S.E.). Tiana, m 522, hazelnut-orchard, bark
traps, 1 male 27.IV-4.V.2005, 2 females 1825.V.2005, 4 females 11-18.V.2005, 1 male and 3 females, 4-11.V.2005, 1 female 6-12.X.2005 leg.
I.S.E. (MBG). Tonara, Su Azzime, m 682, walnutorchard, bark traps, 1 female 11-18.IV.2007, 1 female 16-23.V.2007 leg. I.S.E. (ISE).
SASSARI: Alghero, Porto Conte, m 35, pinewood, emergence cages, 1 female,13.IV.2009 leg. Ist.
Ent. SS (ISE). Sorso, Pianu Mannu, Nali farm, olivegrove, beating tray, 1 female 29.III.1993, 1 male and
1 female 21.IV.1993 (ISE); olive-grove, 1 female
18.VI.1993 (ISE); orchard, bark traps, 1 male 29.XII.1997 (ISE) (all leg. I.S.E.). Torralba, vineyard,
1 female 23.IV.1994, 1 female 18.VI.1994, 1 male
59
and 3 juv. 19.XI.1994, 3 males 12.XI.1994, 1 female
and 9 juv. 26.VIII.1995, 1 male 21.X.1995 leg. Ist.
Ent. SS (ISE).
Clubiona neglecta O.P.-Cambridge, 1862
Clubiona pallidula (Clerck, 1757)
Clubiona phragmitis C.L. Koch, 1843
Clubiona subsultans Thorell, 1875
REFERENCES. (13) Garneri, 1902 (sub C. erratica).
Clubiona terrestris Westring, 1851
HABITAT.Hazelnut-orchard (127, ined.).
m 585, hazelnut-orchard, bark traps, 1 female,
27.IV-4.V.2005, 1 male 1-8.VI.2005, 1 female 1726.VIII.2005, 1 female 5-10.VIII.2005 (ISE), 1 female 20-27.X.2004, 1 female 6-13.X.2004 leg.
I.S.E. (MBG).
Clubiona vegeta Simon, 1918
REFERENCES. (116) Helsdingen, 2005; (132)
60
Family CORINNIDAE
Cetonana laticeps (Canestrini, 1868)
REFERENCES. (77) Grimm, 1986 (sub Ceto laticeps); (84) Pesarini, 1994 (sub Ceto laticeps).
PROVINCE. (84); Nuoro (77, ined.).
EXAMINED MATERIAL. NUORO: Tonara, Su Azzime, m 682, walnut-orchard, bark traps, 1 female
11-18.IV.2007 leg. I.S.E. (ISE).
Liophrurillus flavitarsis (Lucas, 1846)
REFERENCES. (60) Brignoli & Gaddini, 1979
(sub Phrurolitus flavitarsis); (84) Pesarini, 1994
(sub Phrurolitus flavitarsis); (139) Trotta, 2011.
PROVINCE. (84); Cagliari (60, ined.); CarboniaIglesias (139); Medio Campidano (60, ined.);
Olbia-Tempio (139); Nuoro (60, ined.); Oristano
(60); Sassari (60, 139).
(ined.); mediterranean maquis (ined.); hazelnut-orchard (ined.); walnut-orchard (ined.); mining landfills covered by pioneer plants (ined.).
EXAMINED MATERIAL. CARBONIA-IGLESIAS:
Gonnesa, 1 male 24.IV.2003 leg. Ferrario E. (MBG).
MEDIO CAMPIDANO: Arbus, Ingurtosu, Naracauli, m 120, mediterranean maquis, 1 male 21.IV5.V.2009, 1 female, 19.V-4.VI.2009, 2 males
5-19.V.2009, 1 female, 29.VI-14.VII.2009 (ISE); m
170, mining landfills covered by pioneer plants, 1 female 5-19.V.2009 (ISE); m 200, garrigue, 2 males
21.IV-5.V.2009 (ISE) (all pitfall traps and leg. Sassu
A., Verdinelli M.). Guspini, Montevecchio, Piccalinna, m 235, garrigue, 2 males 7-21.IV.2009, 2
males, 21.IV-5.V.2009, 3 males and 1 female 518.V.2009, 1 male and 1 female, 18.V-3.VI.2009
(ISE); m 260, mediterranean maquis, 5 males and 1
female 21.IV-5.V.2009, 18 males and 3 females 518.V.2009, 6 males and 14 females 18.V-3.VI.2009,
6 females 3-15.VI.2009, 1 female 15-30.VI.2009, 1
female 30.VI-13.VII.2009 (ISE) (all pitfall traps and
NUORO: Belvì, Antonitzò, m 683, cherry-orchard, 17 males 18-26.IV.2007, 1 male 1118.IV.2007, 2 males and 4 females 16-23.V.2007, 1
male, 1 female 21-28.VI.2007 (ISE); Canale e'
Figu, m 576, walnut-orchard, 2 females 1623.V.2007, 4 females, 21-28.VI.2007, 1 female 1724.VII.2007 (ISE); Mannaritzò, m 706,
cherry-orchard, 6 males and 1 female 18-26.IV.2007,
12 males and 1 female 11-18.IV.2007, 1 male and 13
females 16-23.V.2007, 5 females 21-28.VI.2007, 2
females 17-24.VII.2007 (ISE); Su Enazzu, m 550,
hazelnut-orchard, 1 male and 16 females 27.VII8.VIII.2006 (MBG), 13 female 13-27.VII.2006
(ISE), 15 female 13-27.VII.2006, 2 females and 1
juv. 8-24.VIII.2006, 1 juv. 31.X-21.XI.2006, 3 juv.
16-31.X.2006, 1 juv. 8.I-21.II.2007, 1 female 19.III26.IV.2007, 5 males and 13 females 30.V-4.VII.2007
(MBG), 1 male 30.V-4.VII.2007 (ISE) (all pitfall
traps and leg. I.S.E.). Tonara, Su Azzime, m 682,
walnut-orchard, pitfall traps, 1 male and 1 female 1826.IV.2007, 2 males and 2 females 16-23.V.2007, 2
females 21-28.VI.2007 leg. I.S.E. (ISE).
Phrurolithus corsicus (Simon, 1878)
PROVINCE. Nuoro (116, ined.).
HABITAT. At the margin of a field with stones
and shurbs (116); under Juniperus sp. (ined.).
EXAMINED MATERIAL. NUORO: Desulo, Gennargentu, mountain refuge, m 1500, under Juniperus
sp., pitfall traps, 1 female 10-24.V.2007 leg. Fiori
F. (MBG). Fonni, Gennargentu, mountain refuge,
m 1550, under Juniperus sp., pitfall traps, 2 males
5-19.VI.2007 leg. Fiori F. (MBG).
REMARKS. The male palp of P. corsicus is illustrated in figures 4-5.
Trachelas minor O.P.-Cambridge, 1872
CHOROTyPE. W-Palearctic.
HABITAT. In grassland along the mouth of the
river (116).
Family zODARIIDAE
Selamia reticulata (Simon, 1870)
61
Figures 4, 5. Phrurolithus corsicus (Simon, 1878) - Fonni (NU) Gennargentu m 1550, 5-19.VI.2007 leg. Fiori F. (scale
0.2 mm): (4) retrolateral view of left male pedipalp; (5) ventral view of left male pedipalp.
PROVINCE. Carbonia-Iglesias (133); Oristano
(ined.).
EXAMINED MATERIAL. ORISTANO: San Vero
Milis, Mandriola, 1 female 28.V.2005 leg. Mulas
A. (UCA).
Zodarion elegans (Simon, 1873)
REFERENCES. (13) Garneri, 1902 (sub Enyo elegans); (84) Pesarini, 1994; (93) Bosmans, 1997;
(112) Thaler & Knoflach, 2004; (132) Pantini &
Sassu, 2009; (139) Trotta, 2011.
(13, 93, 139); Medio Campidano (ined.); Nuoro
(93, ined.); Ogliastra (112); Oristano (ined.); Sassari
(132, ined.).
(ined.); garrigue (ined.); mediterranean maquis
(ined.); mining landfills covered by pioneer
plants (ined.).
EXAMINED
MATERIAL.
MEDIO CAMPIDANO:
maquis, 1 female 14-28.VII.2009, 1 female 19.V4.VI.2009, 1 male 5-19.V.2009 (ISE); m 170, mining landfills covered by pioneer plants, 1 female
29.VI-14.VII.2009, 1 male and 1 female 1629.VI.2009, 1 female 4-16.VI.2009, 2 males and 5
females 28.VII-10.VIII.2009, 2 males and 2 females 14-28.VII.2009, 1 female 10-25.VIII.2009, 1
female 25.VIII-8.IX.2009 (ISE), m 200, garrigue,
3 males 19.V-4.VI.2009, 2 females 16-29.VI.2009,
3 males and 1 female 14-28.VII.2009, 5 males and
5 females 28.VII-10.VIII.2009, 1 male 1025.VIII.2009 (ISE) (all pitfall traps and leg. Sassu
A., Verdinelli M.). Guspini, Montevecchio, Piccalinna, m 225, mining landfills covered by pioneer
plants, 2 males, 13-27.VII.2009 (ISE); m 235, garrigue, 1 female and 5 juv. 21.IV-5.V.2009, 17 males
and 6 juv. 5-18.V.2009, 30 males, 5 females and 1
juv. 18.V-3.VI.2009, 2 females 15-30.VI.2009, 1
male and 7 females 30.VI-13.VII.2009, 5 males and
1 female 3-15.VI.2009, 32 males and 39 females
13-27.VII.2009, 33 males and 27 females 27.VII10.VIII.2009, 5 males and 16 females 1025.VIII.2009, 5 females 25.VIII-8.IX.2009 (ISE),
62
16 males and 23 females 10-25.VIII.2009 (MBG),
4 females 8-22.IX.2009 (ISE) (all pitfall traps and
NUORO: Belvì, Antonitzò, m 683, cherry-orchard, 1 female 16-23.V.2007, 6 males 2128.VI.2007, 1 male 17-24.VII.2007, 1 female
27.IX-4.X.2007 (ISE); Canale e' Figu, m 576, walnut-orchard, 1 female, 16-23.V.2007, 3 males and
2 females 21-28.VI.2007, 1 female 24-30.VIII.2007
(ISE), Su Enazzu, m 550, hazelnut-orchard, 1 female 27.VII-8.VIII.2006, 1 male 8-24.VIII.2006
(ISE) (all pitfall traps and leg. I.S.E.).
SASSARI: Sorso, Pianu Mannu, Nali farm, orchard, pitfall traps, 2 males, 1 female and 1 juv. 2431.VII.1997 leg. Ist. Ent. SS (ISE).
ORISTANO: Cabras, Isola Mal di Ventre, 1
male, 1 female, IV.2006, leg. Mulas A. (UCA).
*Zodarion gallicum (Simon, 1873)
Guspini, Montevecchio, Piccalinna, m 260, mediterranean maquis, pitfall traps, 1 female 30.VI13.VII.2009 leg. Sassu A., Verdinelli M. (MBG).
Zodarion italicum (Canestrini, 1868)
REFERENCES. (13) Garneri, 1902 (sub Enyo italica).
Zodarion nigriceps (Simon, 1873)
REFERENCES. (3) Pavesi, 1876 (sub Enyo nigriceps); (13) Garneri, 1902 (sub Enyo nigriceps);
(15) Simon, 1914; (84) Pesarini, 1994; (93) Bosmans, 1997.
PROVINCE. (15, 84); Cagliari (13); CarboniaIglesias (3, 13); Nuoro (ined.); Ogliastra (13, 93);
Oristano (93).
HABITAT. Cherry-orchard (ined.); walnut-orchard (ined.); under Juniperus sp. (ined.).
EXAMINED MATERIAL. NUORO: Belvì, Antonitzò, m 683, cherry-orchard, 6 male, 1 female 1623.V.2007, 1 male and 1 female 21-28.VI.2007, 1
male 27.IX-4.X.2007 (ISE); Canale e' Figu, m 576,
walnut-orchard, 1 male 21-28.VI.2007 (ISE); Mannaritzò, m 706, cherry-orchard, 1 male 1826.IV.2007, 1 female 16-23.V.2007, 2 males
21-28.VI.2007 (ISE) (all pitfall traps and leg.
I.S.E.). Desulo, Gennargentu, mountain refuge, m
1500, under Juniperus sp., pitfall traps, 1 male 923.VIII.2006 leg. Fiori F. (ISE). Tonara, Su Azzime,
m 682, walnut-orchard, pitfall traps, 1 male 1826.IV.2007, 1 male 16-23.V.2007, 2 males and 3 females 21-28.VI.2007, 2 males 24-30.VIII.2007 leg.
I.S.E. (ISE).
Zodarion ruffoi Caporiacco, 1951
CHOROTyPE. Italian endemic.
REFERENCES. (93) Bosmans, 1997.
(ined.); under Juniperus sp. (ined.); mining landfills
covered by pioneer plants (ined.).
maquis, 1 male 4-16.VI.2009 (MBG); m 200, garrigue, 1 female 14-28.VII.2009 (MBG) (all pitfall
traps and leg. Sassu A., Verdinelli M.). Guspini,
covered by pioneer plants, 1 female 10-25.VIII.2009
(MBG); m 235, garrigue, 16 males 5-18.V.2009, 1
female 6-20.X.2009 (MBG) (all pitfall traps and leg.
NUORO: Fonni, Gennargentu, mountain refuge,
m 1550, under Juniperus sp., pitfall traps, 1 male
5-19.VI.2007 leg. Fiori F. (ISE).
Zodarion sardum Bosmans, 1997
REFERENCES. (49) Wunderlich, 1973 (sub Zodarion (?) vicinum); (93) Bosmans, 1997.
PROVINCE. (49, 93).
Family PRODIDOMIDAE
*Zimirina brevipes Pérez et Blasco, 1986
63
plants (ined.).
Arbus, Ingurtosu, Naracauli, m 170, mining landfills
covered by pioneer plants, pitfall traps, 1 female
19.V-4.VI.2009 leg. Sassu A., Verdinelli M. (MBG).
REMARKS. a new record for Italy of this family
and this species known only from Southern Spain.
The epigynum is illustrated in figure 6.
Family GNAPHOSIDAE
Aphantaulax cincta (L. Koch, 1866)
CHOROTyPE. Europeo-Mediterranean.
REFERENCES. (76) Grimm, 1985; (84) Pesarini,
1994; (127) Loru et al., 2008; (139) Trotta, 2011.
HABITAT. Hazelnut-orchard (127, ined.); sandstrand (76).
m 585, hazelnut-orchard, bark traps, 1 female 411.V.2005 leg. I.S.E. (ISE). Tiana, m 522, hazelnutorchard, beating tray, 1 female 19.V.2005, 1 male
16.VI.2005 leg. I.S.E. (ISE).
Aphantaulax trifasciata (O.P.-Cambridge, 1872)
REFERENCES. (8) Costa, 1884 (sub A. seminiger);
(13) Garneri, 1902 (sub A. semi-niger); (76) Grimm,
1985 (sub A. seminigra); (84) Pesarini, 1994 (sub A.
seminigra); (132) Pantini & Sassu, 2009.
PROVINCE. (84); Cagliari (8, 13); Medio-Campidano (ined.); Nuoro (8, 13, 76, ined.); Olbia-Tempio
(8, 13); Sassari (8, 13, 132, ined.).
HABITAT. Garrigue (ined.); inside house (ined.);
grassland (ined.).
Arbus, Ingurtosu, Naracauli, m 200, garrigue, pitfall traps, 1 male 5-19.V.2009 leg. Sassu A., Verdinelli M. (ISE).
NUORO: Bolotana, Badde e'Salighes, m 850,
grassland, 1 male 27.V.2007 leg. Sassu A. (ISE).
SASSARI: Sassari, Li Punti, Baldinca, m 110,
inside house, 1 male 2.VII.2008 leg. Cossu S. (ISE).
Figure 6. Zimirina brevipes Pérez & Blasco, 1986 - epigyne,
ventral view (scale 0.1 mm), Arbus (VS), Ingurtosu, Naracauli m 120, 4.VI.2009 leg. Sassu A., Verdinelli M.
Drassodes lapidosus (Walckenaer, 1802)
REFERENCES. (5) Pavesi, 1878 (sub Drassus lapidicola); (8) Costa, 1884 (sub Drassus lapidosus);
(13) Garneri, 1902 (sub Drassus lapidosus); (20)
Krauss, 1955; (84) Pesarini, 1994; (127) Loru et al.,
2008; (132) Pantini & Sassu, 2009; (133) Trotta,
2009; (139) Trotta, 2011.
PROVINCE. (20, 84, 127); Cagliari (13); Carbonia-Iglesias (139); Medio Campidano (ined.); Nuoro
(8, 13, ined.); Nuoro/Ogliastra (13); Olbia-Tempio
(5, 8, 13, 133); Oristano (ined.); Sassari (132, 133).
HABITAT. Cherry-orchard (ined.); hazelnut-orchard (127, ined.); alder-grove (ined.); under Juniperus sp. (ined.); mining landfills covered by
pioneer plants (ined.).
covered by pioneer plants, pitfall traps, 1 male 519.V.2009 (MBG), 1 male 5-19.V.2009 (ISE) (all
leg. Sassu A., Verdinelli M.). Guspini, Montevecchio, Piccalinna, m 225, mining landfills covered by
pioneer plants, pitfall traps, 1 male 21.IV-5.V.2009
(ISE), 2 males 18.V-3.VI.2009 (MBG), 1 male 315.VI.2009 (ISE) (all leg. Sassu A., Verdinelli M.).
NUORO: Belvì, Mannaritzò, m 706, cherry-orchard, pitfall traps, 1 male 16-23.V.2007, 1 female
64
17-24.VII.2007 (ISE); Su Enazzu, m 550, hazelnutorchard, bark traps, 1 male 18-25.V.2005, 1 female
14-19.VII.2005 (ISE) (all leg. I.S.E.). Desulo, Gennargentu, mountain refuge, m 1500, alder-grove,
bark traps, 1 female 25.VI-2.VII.2007 leg. Fiori F.
(ISE). Fonni, Gennargentu, mountain refuge, m
1550, alder-grove, 1 female 19.VII-2.VIII.2006, 1
female 19.VI-1.VII.2007 (ISE); under Juniperus sp.,
2 females 5-19.VI.2007 (ISE) (all pitfall traps and
leg. Fiori F.). Oliena, Cedrino lake, m 80, light trap,
1 male 20.V.1994 leg. Pantini P., Valle M. (MBG).
ORISTANO: Cabras, Isola Mal di Ventre, 3
males 1 female 14-18.IV.2006 leg. Mulas A. (UCA).
*Drassodes luteomicans (Simon, 1878)
HABITAT. Garrigue (ined.); mediterranean maquis (ined.); mining landfills covered by pioneer
plants (ined.).
maquis, 1 male 16-29.VI.2009 l, 1 female 416.VI.2009 (MBG); m 170, mining landfills covered
by pioneer plants, 1 female 21.IV-5.V.2009, 1 female
28.VII-10.VIII.2009 (MBG); m 200, garrigue, 1 female 19.V-4.VI.2009, 1 female 29.VI-14.VII.2009,
1 female 6-20.X.2009 (ISE) (all pitfall traps and leg.
traps, 1 female 18.V-3.VI.2009 (ISE), 1 female 315.VI.2009 (MBG) (all leg. Sassu A., Verdinelli M.).
REMARKS. The male palp, the epigynum and the
vulva of this species are illustrated in figures 7-11.
Drassodes lutescens (C.L. Koch, 1839)
CHOROTyPE. Turano-European-Mediterranean.
REFERENCES. (13) Garneri, 1902 (sub Drassus
lutescens); (20) Krauss, 1955; (84) Pesarini, 1994;
(139) Trotta, 2011.
PROVINCE. (20, 84); Carbonia-Iglesias (139); Medio
Campidano (ined.); Nuoro (ined.); Nuoro/Ogliastra
(13); Ogliastra (13); Oristano (ined.).
HABITAT. Garrigue (ined.); mediterranean maquis (ined.); alder-grove (ined.); under Juniperus
sp. (ined.); mining landfills covered by pioneer
plants (ined.).
maquis, 1 male 5-19.V.2009, 1 male 29.VI14.VII.2009, 1 male, 14-28.VII.2009 (ISE); m 170,
mining landfills covered by pioneer plants, 1 male
and 1 female 21.IV-5.V.2009, 1 male 7-21.IV.2009
(ISE); m 200, garrigue, 1 male 21.IV-5.V.2009, 2
males 7-21.IV.2009, 3 males and 2 females 19.V4.VI.2009 (ISE), 3 males and 3 females 5-19.V.2009
(MBG), 1 male 5-19.V.2009 (ISE) (all pitfall traps
and leg. Sassu A., Verdinelli M.). Guspini, Montevecchio, Piccalinna, m 225, mining landfills covered
by pioneer plants, pitfall traps, 1 female 21.IV5.V.2009, 1 male 18.V-3.VI.2009, 1 female 1530.VI.2009, 1 male 18.V-3.VI.2009 leg. Sassu A.,
NUORO: Desulo, Gennargentu, mountain refuge,
m 1500, alder-grove, bark traps, 1 female 29.VIII5.IX.2006, 1 female and 1 juv. 26.IX-3.X.2006, 3
females 13-19.IX.2006, 1 male and 1 female 24.IV1.V.2007 leg. Fiori F. (ISE). Fonni, Gennargentu,
mountain refuge, m 1550, alder-grove, pitfall traps,
1 female 17-30.VIII.2006 (ISE); alder-grove, bark
traps, 1 female 24.IV-1.V.2007 (ISE); under Juniperus sp., pitfall traps, 1 female 19.VI-1.VII.2007, 2
males 5-19.VI.2007 (ISE) (all leg. Fiori F.).
ORISTANO: Cabras, Isola Mal di Ventre, 1 male,
2 females 12-21.II.2005, 5 females 17.IV.2006 leg.
Mulas A. (UCA).
Drassodes rubidus (Simon, 1878)
rubidus); (84) Pesarini, 1994.
PROVINCE. (84); Carbonia-Iglesias (13); Nuoro/
Ogliastra (13); Ogliastra (13).
Drassodex hispanus (L. Koch, 1866)
hispanus); (84) Pesarini, 1994 (sub Drassodes hispanus).
Drassodex hypocrita (Simon, 1878)
hypocrita).
65
Figures 7-9. Drassodes luteomicans (Simon, 1878), Ingurtosu, Naracauli, m 120, 16-29.VI.2009 leg. Sassu A., Verdinelli
M. (scale 0.5 mm): (7) retrolateral view of left male pedipalp; (8) ventral view of left male pedipalp; (9) chelicerae, posterior
view. Figures 10-11. D. luteomicans (Simon, 1878), Ingurtosu, Naracauli, m 170, 28.VII-10.VIII.2009 leg. Sassu A., Verdinelli M. (scale 0.2 mm): (10) epigyne, ventral view; (11) vulva, ventral view.
Gnaphosa alacris Simon, 1878
(ined.); Ogliastra (13); Sassari (ined.).
HABITAT. Orchard (ined.); garrigue (ined.); walnut-orchard (ined.); mining landfills covered by
Arbus, Ingurtosu, Naracauli, m 170, mining
landfills covered by pioneer plants, 1 male 416.VI.2009, 1 male and 3 females 29.VI14.VII.2009 (ISE), 1 female 16-29.VI.2009
(MBG), 1 female 28.VII-10.VIII.2009 (ISE); m
200, garrigue, 1 male 29.VI-14.VII.2009 (ISE)
landfills covered by pioneer plants, 1 male 1327.VII.2009, 1 female 27.VII-10.VIII.2009, 1 male
8-22.IX.2009 (ISE); m 235, garrigue, 1 male 1327.VII.2009 (MBG) (all pitfall traps and leg. Sassu
A., Verdinelli M.).
NUORO: Belvì, Canale e' Figu, m 576, walnutorchard, pitfall traps, 1 male 17-24.VII.2007 leg.
I.S.E. (ISE).
SASSARI: Sorso, Pianu Mannu, Nali farm, orchard, pitfall traps, 1 female 26.VI-2.VII.1997 leg.
Ist. Ent. SS (ISE).
Gnaphosa bicolor (Hahn, 1833)
PROVINCE. Cagliari (13); Nuoro/Ogliastra (13).
66
Gnaphosa lucifuga (Walckenaer, 1802)
1994.
PROVINCE. (84); Cagliari (13); Carbonia-Iglesias (13).
Gnaphosa lugubris (C.L. Koch, 1839)
Haplodrassus dalmatensis (L. Koch, 1866)
minusculus); (20) Krauss, 1955; (116) Helsdingen,
2005; (132) Pantini & Sassu, 2009.
PROVINCE. (20); Cagliari (13); Medio Campidano (ined.); Ogliastra (116); Sassari (132).
HABITAT. Garrigue (ined.); in coastal drift on the
shore (116); mining landfills covered by pioneer
plants (ined.).
Arbus, Ingurtosu, Naracauli, m 170, mining landfills covered by pioneer plants, pitfall traps, 1 male
7-21.IV.2009, 2 males and 1 female 21.IV-5.V.2009
(ISE), 2 males and 1 female 5-19.V.2009 (MBG), 2
females 5-19.V.2009, 1 female 16-29.VI.2009, 2
males 7-21.IV.2009 (ISE) (all leg. Sassu A., Verdinelli M.). Guspini, Montevecchio, Piccalinna, m
225, mining landfills covered by pioneer plants, 1
male 7-21.IV.2009, 2 males and 3 females 21.IV5.V.2009, 3 females 18.V-3.VI.2009, 1 male and 1
female 5-18.V.2009, 2 females 15-30.VI.2009
(ISE); m 235, garrigue, 2 males 18.V-3.VI.2009 (all
Haplodrassus macellinus (Thorell, 1871)
REFERENCES. (11) de Carlini, 1885 (sub Drassus
macellinus); (13) Garneri, 1902 (sub Drassus macellinus); (20) Krauss, 1955; (84) Pesarini, 1994.
PROVINCE. (84); Carbonia-Iglesias (11, 13); Sassari (20).
Haplodrassus severus (C.L. Koch, 1839)
REFERENCES. (10) Costa, 1885 (sub Drassus severus); (20) Krauss, 1955; (30) Cerruti, 1968; (84)
Pesarini, 1994; (132) Pantini & Sassu, 2009.
PROVINCE. (20, 30, 84); Cagliari (10); Nuoro
EXAMINED MATERIAL. NUORO: Gavoi, S'eremu
e sa mela, m 1000, 1 male 13.VI.2006 leg. Mulas
A. (UCA).
Haplodrassus signifer (C.L. Koch, 1839)
REFERENCES. (6) Magretti, 1880 (sub Drassus
troglodytes); (8) Costa, 1884 (sub Drassus troglodytes); (13) Garneri, 1902 (sub Drassus troglodytes); (20) Krauss, 1955; (84) Pesarini, 1994; (116)
Helsdingen, 2005; (133) Trotta, 2009.
PROVINCE. (20, 84); Cagliari (13); CarboniaIglesias (6); Nuoro (116, ined.); Nuoro/Ogliastra (8,
13); Olbia-Tempio (133).
HABITAT. At the margin of a field with stones and
shurbs (116); cherry-orchard (ined.).
EXAMINED MATERIAL. NUORO: Belvì, Mannaritzò, m 706, cherry-orchard, pitfall traps, 1 female
18-26.IV.2007 leg. I.S.E. (ISE).
*Leptodrassus femineus (Simon, 1873)
landfills covered by pioneer plants, 1 female
27.VII-10.VIII.2009 (ISE); m 235, garrigue, 1 female 25.VIII-8.IX.2009 (ISE) (all pitfall traps and
*Micaria albovittata (Lucas, 1846)
HABITAT. Under Juniperus sp. (ined.); walnutorchard (ined.).
EXAMINED MATERIAL. NUORO: Fonni, Gennargentu, mountain refuge, m 1550, under Juniperus sp.,
pitfall traps, 1 male 19.VI-1.VII.2007 leg. Fiori F.
(ISE). Tonara, Su Azzime, m 682, walnut-orchard,
pitfall traps, 1 female 21-28.VI.2007 leg. I.S.E. (ISE).
Micaria coarctata (Lucas, 1846)
REFERENCES. (8) Costa, 1884 (sub M. smaragdula); (13) Garneri, 1902 (sub M. smaragdula).
(ined.); Sassari (8, 13).
HABITAT. Cherry-orchard (ined.); garrigue (ined.);
covered by pioneer plants, 1 female 29.VI14.VII.2009, 1 male 4-16.VI.2009 (ISE); m 200,
garrigue, 1 male 16-29.VI.2009 (ISE) (all pitfall
Montevecchio, Piccalinna, m 235, garrigue, pitfall
traps, 1 male 15-30.VI.2009 (ISE), 1 male 1530.VI.2009 (MBG) (all leg. Sassu A., Verdinelli M.).
NUORO: Belvì, Antonitzò, m 683, cherry-orchard, pitfall traps, 1 female 17-24.VII.2007 leg.
I.S.E. (ISE).
*Micaria cyrnea Brignoli, 1983
HABITAT. Forest of Quercus suber (ined.); walnut-orchard (ined.).
Figu, m 576, walnut-orchard, bark traps, 1 male
26.IV-4.V.2007 leg. I.S.E. (ISE).
OLBIA-TEMPIO: Buddusò, near Nuraghe Loelle, forest of Quercus suber, beating tray, 1 female,
Micaria pulicaria (Sundevall, 1831)
Nomisia aussereri (L. Koch, 1872)
67
REFERENCES. (20) Krauss, 1955; (76) Grimm,
1985; (84) Pesarini, 1994.
PROVINCE. (76, 84); Nuoro (76); Sassari (20).
Nomisia exornata (C.L. Koch, 1839)
REFERENCES. (13) Garneri, 1902 (sub Pythonissa
exornata); (76) Grimm, 1985; (84) Pesarini, 1994.
PROVINCE. (13, 84); Cagliari (13); CarboniaIglesias (76); Medio Campidano (ined.); Nuoro
(ined.); Nuoro/Ogliastra (13).
HABITAT. Road verge (6); garrigue (ined.); mediterranean maquis (ined.); walnut-orchard (ined.); mining landfills covered by pioneer plants (ined.).
maquis, 1 male 4-16.VI.2009, 1 female 1025.VIII.2009 (ISE); m 170, mining landfills covered by pioneer plants, 8 males, 3 females and 2 juv.
19.V-4.VI.2009 (ISE), 1 male and 1 female 416.VI.2009 (MBG), 7 males and 2 females 416.VI.2009, 4 females 16-29.VI.2009, 3 females
29.VI-14.VII.2009 (ISE); m 200, garrigue, 6 males
and 2 females 19.V-4.VI.2009, 1 female 29.VI14.VII.2009, 2 males 4-16.VI.2009 (ISE) (all pitfall traps and leg. Sassu A., Verdinelli M.). Guspini,
covered by pioneer plants, 1 female 30.VI13.VII.2009, 1 male 3-15.VI.2009 (ISE); m 235,
garrigue, pitfall traps, 3 males 18.V-3.VI.2009 (ISE)
NUORO: Belvì, Canale e' Figu, m 576, walnutorchard, pitfall traps, 2 males 21-28.VI.2007 leg.
I.S.E. (ISE).
*Nomisia recepta (Pavesi, 1880)
(ined.).
maquis, 1 female 14-28.VII.2009 (ISE), m 170; mining landfills covered by pioneer plants, 1 male 416.VI.2009, 1 male 16-29.VI.2009, 4 females
29.VI-14.VII.2009, 1 female 14-28.VII.2009, 2 females 10-25.VIII.2009 (ISE); m 200, garrigue, 2
68
males and 1 female, 19.V-4.VI.2009, 2 males and 1
female 16-29.VI.2009, 2 females 14-28.VII.2009,
1 female 10-25.VIII.2009 (ISE) (all pitfall traps and
by pioneer plants, 1 male 3-15.VI.2009 (ISE); m
260, mediterranean maquis, 1 male 18.V-3.VI.2009,
1 female 15-30.VI.2009, 1 male, 30.VI13.VII.2009, 1 male 3-15.VI.2009 (ISE), 1 male
and 1 female 15-30.VI.2009 (MBG), 1 female 1327.VII.2009, 1 female 27.VII-10.VIII.2009, 1 female 10-25.VIII.2009 (ISE) (all pitfall traps and
traps, 1 male, 18.V-3.VI.2009, leg. Sassu A., Verdinelli M. (ISE).
NUORO: Belvì, Su Enazzu, m 550, hazelnut-orchard, bark traps, 1 female 25.V-1.VI.2005 leg.
I.S.E. (ISE).
SASSARI: Sorso, Pianu Mannu, Nali farm, orchard, bark traps, 1 male and 1 female 2431.VII.1997 leg. Ist. Ent. SS (ISE).
plants (ined.).
landfills covered by pioneer plants, pitfall traps, 1 female 3-15.VI.2009 leg. Sassu A., Verdinelli M. (ISE).
PROVINCE. Cagliari (ined.); Nuoro (ined.);
Olbia-Tempio (133); Oristano (ined.); Sassari
(ined.).
HABITAT. Citrus-grove (ined.); on trees lining a
street (ined.); orchard (ined.); hazelnut-orchard
(ined.).
EXAMINED MATERIAL. CAGLIARI: Uta, S’Appassiu, Agris farm, citrus-grove, beating tray, 1 female,
2.XI.2000, 1 female 13.XI.2000 leg. CRAS (ISE).
NUORO: Belvì, Su Enazzu, m 550, hazelnut-orchard, bark traps, 1 female 20-27.X.2004 leg. I.S.E.
(ISE). Gavoi, 1 male 14.IX.2009 leg. Mulas A.
(UCA).
ORISTANO: San Vero Milis, Mandriola, 1 female 29.V.2005, 1 female 21.VIII.2008 leg. Mulas
A. (UCA).
SASSARI: Alghero, inside house, at sea level, 1
male 31.IX.1998 leg. Pantaleoni R.A. (ISE). Sassari, m 522, on trees lining a street, bark traps, 1 female 10.XI.1993 leg. Ist. Ent. SS (ISE). Sorso,
Pianu Mannu, Nali farm, orchard, bark traps, 1
male, 18-25.IX.1997, 1 female 24.XI-2.XII.1997
*Phaeocedus braccatus (L. Koch, 1866)
Poecilochroa albomaculata (Lucas, 1846)
REFERENCES. (13) Garneri, 1902 (sub Poecilochroa picta); (84) Pesarini, 1994.
PROVINCE. (84); Cagliari (13); Ogliastra (13);
Oristano (13).
Scotophaeus blackwalli (Thorell, 1871)
et al., 2008.
PROVINCE. (127); Cagliari (ined.); Medio-Campidano (ined.); Nuoro (ined.); Olbia-Tempio (ined.);
trees (94); orchard (ined.); mediterranean maquis
(ined.); hazelnut-orchard (127, ined.); pinewood
(ined.); forest of Quercus suber (ined.).
Manna, m 140, pinewood, emergence cages, 1 male,
30.VI.2009 leg. Ist. Ent. SS (ISE). Uta, S’Appassiu,
Agris farm, citrus-grove, beating tray, 1 female
23.X.2000, 1 female, 2.XI.2000 leg. CRAS (ISE).
Scotophaeus scutulatus (L. Koch, 1866)
Scotophaeus validus (Lucas, 1846)
1994; (127) Loru et al., 2008.
PROVINCE. (20, 84, 127); Nuoro (ined.).
Enazzu, m 550, hazelnut-orchard, bark traps, 1 female 29.IX-6.X.2005 leg. I.S.E. (ISE).
Setaphis carmeli (O.P.-Cambridge, 1872)
REFERENCES. (2b) Canestrini, 1873 (sub Melanophora latipes); (11) de Carlini, 1885 (sub Prosthesima latipes); (13) Garneri, 1902 (sub
Prosthesima latipes and sub Prosthesima brachialis, n. sp.); (20) Krauss, 1955 (sub Zelotes carmeli);
(84) Pesarini, 1994 (sub Zelotes carmeli).
PROVINCE. (20, 2b, 84); Carbonia-Iglesias (11,
13); Medio Campidano (ined.); Nuoro (ined.);
Ogliastra (13).
Arbus, Ingurtosu, Naracauli, m 170, mining landfills covered by pioneer plants, 1 male 21.IV5.V.2009 (ISE), 1 male 5-19.V.2009 (MBG), 1
male, 5-19.V.2009 (ISE); m 200, garrigue, 1 male
and 1 female, 19.V-4.VI.2009 (ISE) (all pitfall traps
and leg. Sassu A., Verdinelli M.). Guspini, Montevecchio, Piccalinna, m 235, garrigue, pitfall traps,
4 males, 5-18.V.2009 (ISE), 1 female, 18.V3.VI.2009. (MBG), 1 female, 30.VI-13.VII.2009
(ISE) (all leg. Sassu A., Verdinelli M.).
NUORO: Belvì, Antonitzò, m 683, cherry-orchard, 1 male 21-28.VI.2007 (ISE); Canale e' Figu,
m 576, walnut-orchard, 1 male and 1 female 1623.V.2007, 2 females 21-28.VI.2007 (ISE); Mannaritzò, m 706, cherry-orchard, 1 male 18-26.IV.2007,
4 males 16-23.V.2007, 2 males and 1 female 2128.VI.2007 (ISE); Su Enazzu, m 550, hazelnut-orchard, 1 female 27.VII-8.VIII.2006, 2 females
13-27.VII.2006 (MBG), 3 females 13-27.VII.2006
(ISE), 1 female 8-24.VIII.2006 (MBG) (all pitfall
traps and leg. I.S.E.).
*Setaphis parvula (Lucas, 1846)
Arbus, Ingurtosu, Naracauli, m 170, mining land-
69
fills covered by pioneer plants, 1 male 19.V4.VI.2009 (MBG), 1 male 5-19.V.2009 (ISE); m
200, garrigue, 1 male 19.V-4.VI.2009 (ISE) (all pitfall traps and leg. Sassu A., Verdinelli M.).
for Italian fauna.
Trachyzelotes barbatus (L. Koch, 1866)
REFERENCES. (10) Costa, 1885 (sub Prosthesima
barbata).
PROVINCE. Carbonia-Iglesias (10); Medio Campidano (ined.); Nuoro (ined.).
Arbus, Ingurtosu, Naracauli, m 170, mining landfills covered by pioneer plants, 1 male 19.V4.VI.2009 (MBG), m 200; garrigue, 1 male
21.IV-5.V.2009, 5 males 5-19.V.2009, 1 male and
1 female 19.V-4.VI.2009, 1 female 4-16.VI.2009
(ISE) (all pitfall traps and leg. Sassu A., Verdinelli
M.). Guspini, Montevecchio, Piccalinna, m 235,
garrigue, pitfall traps, 1 male 21.IV-5.V.2009, 1
male 18.V-3.VI.2009 (ISE), 1 male and 3 females,
30.VI-13.VII.2009 (MBG), 2 females 30.VI13.VII.2009, 2 males 3-15.VI.2009, 2 males and
1 female 15-30.VI.2009, 3 females 27.VII10.VIII.2009, 1 female 13-27.VII.2009, 1 male 620.X.2009 (ISE) (all leg. Sassu A., Verdinelli M.).
NUORO: Belvì, Antonitzò, m 683, cherry-orchard, 1 male, 17-24.VII.2007 (ISE); Mannaritzò,
m 706, cherry-orchard, 1 male 21-28.VI.2007
(ISE) (all pitfall traps and leg. I.S.E.).
Trachyzelotes fuscipes (L. Koch, 1866)
REFERENCES. (13) Garneri, 1902 (sub Prosthesima fuscipes).
*Trachyzelotes pedestris (C.L. Koch, 1837)
Mannu, Nali farm, orchard, pitfall traps, 1 male and
70
1 female, 26.VI-2.VII.1997, 1 male 10-17.VII.1997,
1 male 11-19.VI.1998 leg. Ist. Ent. SS (ISE).
Urozelotes rusticus (L. Koch, 1872)
REFERENCES. (10) Costa, 1885 (sub Prosthesima
rustica).
*Zelotes dentatidens Simon, 1914
plants (ined.).
Arbus, Ingurtosu, Naracauli, m 170, mining
landfills covered by pioneer plants, pitfall traps,
2 females 5-19.V.2009, 1 female 19.V-4.VI.2009
(ISE), 1 male and 1 female 19.V-4.VI.2009 (MBG),
1 male and 1 female 4-16.VI.2009, 2 females 1629.VI.2009 (ISE) (all leg. Sassu A., Verdinelli M.).
male 21.IV-5.V.2009, 1 male 5-18.V.2009. (MBG),
1 male and 2 females, 18.V-3.VI.2009, 3 females
30.VI-13.VII.2009, 1 male and 3 females 1530.VI.2009 (ISE), 2 females 30.VI-13.VII.2009
(MBG), 1 male and 1 female 3-15.VI.2009 (ISE),
1 male and 2 females 3-15.VI.2009 (MBG), 2 females 27.VII-10.VIII.2009 (ISE), 1 female 1327.VII.2009, 1 female 25.VIII-8.IX.2009 (MBG),
1 female 10-25.VIII.2009 (ISE) (all leg. Sassu A.,
Verdinelli M.).
REMARKS. The epigynum and the male palp of
this species are illustrated in figures 12, 13.
*Zelotes egregius Simon, 1914
EXAMINED MATERIAL. NUORO: Desulo, Gennargentu, mountain refuge, m 1500, under Juniperus
sp., pitfall traps, 1 male 23.VIII-5.IX.2006 leg. Fiori
F. (ISE). Fonni, Gennargentu, mountain refuge, m
1550, under Juniperus sp., pitfall traps, 1 male 217.VIII.2006 leg. Fiori F. (ISE).
Figures 12, 13. Zelotes dentatidens Simon, 1914 (scale 0.2 mm): (12) epigyne, ventral view, Guspini (VS), Montevecchio,
Piccalinna m 225, 3-15.VI.2009 leg. Sassu A., Verdinelli M.; (13) ventral view of left male pedipalp, Arbus (VS), Ingurtosu,
Naracauli m 120, 19.V-4.VI.2009 leg. Sassu A., Verdinelli M.
Zelotes femellus (L. Koch, 1866)
REFERENCES. (20) Krauss, 1955; (111) Senglet,
2004.
Ogliastra (111); Olbia-Tempio (111); Sassari (111).
HABITAT. Hazelnut-orchard (ined.); walnut-orchard (ined.).
Figu, m 576, walnut-orchard, 3 males, 27.IX4.X.2007 (ISE), Su Enazzu, m 550, hazelnut-orchard, 1 male, 21.IX-5.X.2006 (ISE) (all pitfall
traps and leg. I.S.E.). Gavoi, S'eremu e sa mela, m
1000, 1 female 10.V.2005, 1 female 16.II.2006, 1
female 13.III.2006, 1 female 18.IX.2006 leg.
Mulas A. (UCA).
Zelotes fuscorufus (Simon, 1878)
REFERENCES. (111) Senglet, 2004; (139) Trotta,
2011.
PROVINCE. Medio Campidano (ined.); Oristano
(111); Sassari (111, 139).
HABITAT. Dry litter on sand (111); dry litter,
barks (111); garrigue (ined.); mediterranean maquis
(ined.).
maquis, 2 males, 16-29.VI.2009, 3 males and 1 female 29.VI-14.VII.2009, 2 females 28.VII10.VIII.2009, 1 male and 2 females, 14-28.VII.2009,
1 female 10-25.VIII.2009, 1 female 25.VIII8.IX.2009 (ISE); m 170, mining landfills covered by
pioneer plants, 1 female 16-29.VI.2009, 1 female
29.VI-14.VII.2009, 1 male and 2 females 1428.VII.2009 (ISE); m 200, garrigue, 1 female 29.VI14.VII.2009 (ISE), 1 female 28.VII-10.VIII.2009
(MBG), 1 female 14-28.VII.2009 (ISE) (all pitfall
covered by pioneer plants, 2 males and 2 females
30.VI-13.VII.2009, 1 male 15-30.VI.2009, 2 females 27.VII-10.VIII.2009, 1 male and 1 female 1327.VII.2009 (ISE); m 235, garrigue, 1 female
13-27.VII.2009 (ISE); m 260, mediterranean maquis, 3 males 15-30.VI.2009 (ISE), 1 female 30.VI-
71
13.VII.2009 (MBG), 1 male 3-15.VI.2009 (ISE), 2
males 3-15.VI.2009 (MBG), 1 female 30.VI13.VII.2009, 4 females 13-27.VII.2009, 1 female
27.VII-10.VIII.2009 (ISE) (all pitfall traps and leg.
Zelotes nilicola (O.P.-Cambridge, 1874)
(ined.); Sassari (132, ined.).
HABITAT. Cherry-orchard (ined.); orchard (ined.);
walnut-orchard (ined.); mining landfills covered by
covered by pioneer plants, pitfall traps, 2 females
29.VI-14.VII.2009 leg. Sassu A., Verdinelli M. (ISE).
NUORO: Belvì, Canale e' Figu, m 576, walnutorchard, 1 male and 3 females 21-28.VI.2007, 2 females 17-24.VII.2007 (ISE); Mannaritzò, m 706,
cherry-orchard, 1 male 21-28.VI.2007 (ISE) (all pitfall traps and leg. I.S.E.).
SASSARI: Sorso, Pianu Mannu, Nali farm, orchard, pitfall traps, 1 male 26.VI-2.VII.1997 leg.
Ist. Ent. SS (ISE).
Zelotes ruscinensis Simon, 1914
REFERENCES. (111) Senglet, 2004; (139) Trotta,
2011.
Campidano (ined.); Oristano (ined.); Sassari (111).
covered by pioneer plants, 1 male 16-29.VI.2009,
12 males 29.VI-14.VII.2009, 7 females 28.VII10.VIII.2009, 7 males and 1 female, 1428.VII.2009, 2 females 10-25.VIII.2009 (ISE), 2
females 10-25.VIII.2009 (MBG), 2 females 25.VIII8.IX.2009, 1 female 8-22.IX.2009 (ISE); m 200,
garrigue, 23 males 29.VI-14.VII.2009, 1 male 1629.VI.2009 (ISE), 5 males and 1 female, 29.VI14.VII.2009 (MBG), 1 male and 20 females
28.VII-10.VIII.2009, 22 males and 11 females 14-
72
28.VII.2009, 7 females 25.VIII-8.IX.2009, 13 females 10-25.VIII.2009, 1 female 22.IX-6.X.2009
(ISE) (all pitfall traps and leg. Sassu A., Verdinelli
M.). Guspini, Montevecchio, Piccalinna, m 225, mining landfills covered by pioneer plants, 1 male
30.VI-13.VII.2009 (ISE); m 235, garrigue, 2 males
15-30.VI.2009, 9 males 30.VI-13.VII.2009, 4 males
and 6 females, 27.VII-10.VIII.2009, 14 males and 5
females 13-27.VII.2009 (ISE), 3 females 1025.VIII.2009 (MBG), 8 females 10-25.VIII.2009, 2
females 25.VIII-8.IX.2009, 2 females 22.IX6.X.2009, 4 females 8-22.IX.2009 (ISE) (all pitfall
ORISTANO: San Vero Milis, Mandriola, 1 male
7.VIII.2004; Spinarbas 1 male 1.VII.2004 leg.
Mulas A. (UCA).
Zelotes sardus (Canestrini, 1873)
REFERENCES. (2b) Canestrini, 1873 (sub Prosthesima sarda); (2c) Canestrini, 1876 (sub Prosthesima
sarda); (3) Pavesi, 1876 (sub Prosthesima sarda);
(13) Garneri, 1902 (sub Prosthesima sarda); (73)
Brignoli & Murphy, 1984; (84) Pesarini, 1994; (111)
Senglet, 2004; (133) Trotta, 2009; (139) Trotta, 2011.
PROVINCE. (13, 2b, 2c, 84); Cagliari (13, 73);
Carbonia-Iglesias (3, 13, 139); Medio Campidano
(73, ined.); Nuoro (73, 111, ined.); Nuoro/Ogliastra
(13); Ogliastra (13, 111); Olbia-Tempio (73, 133);
Oristano (13, 111, ined.); Sassari (73, 111, ined.).
(ined.); mediterranean maquis (ined.); walnut-orchard (ined.); under Juniperus sp. (ined.); mining
covered by pioneer plants, 1 female 21.IV-5.V.2009,
1 female 19.V-4.VI.2009, 2 males 25.VIII8.IX.2009, 2 males 8-22.IX.2009, 1 male and 1 female 22.IX-6.X.2009 (ISE); m 200, garrigue, 5
females 21.IV-5.V.2009, 3 females 19.V-4.VI.2009,
2 females 5-19.V.2009, 1 female 16-29.VI.2009, 1
female 14-28.VII.2009, 1 male 25.VIII-8.IX.2009,
3 males 8-22.IX.2009, 7 males 22.IX-6.X.2009, 5
males 6-20.X.2009 (ISE) (all pitfall traps and leg.
Piccalinna, m 225, mining landfills covered by pioneer plants, 2 females 21.IV-5.V.2009, 2 females 5-
18.V.2009, 1 female 18.V-3.VI.2009, 2 females 1530.VI.2009, 1 male 10-25.VIII.2009, 2 males
25.VIII-8.IX.2009, 1 male 6-20.X.2009 (ISE); m
235, garrigue, 4 females 21.IV-5.V.2009, 2 females
5-18.V.2009, 1 female 3-15.VI.2009, 2 males 1025.VIII.2009, 14 males 25.VIII-8.IX.2009, 10 males
and 1 female 8-22.IX.2009, 6 males 22.IX6.X.2009, 1 male and 1 female 6-20.X.2009 (ISE);
m 260, mediterranean maquis, 1 female 315.VI.2009 (ISE) (all pitfall traps and leg. Sassu A.,
Verdinelli M.).
NUORO: Belvì, Antonitzò, m 683, cherry-orchard, 4 males 24-30.VIII.2007, 2 males 27.IX4.X.2007 (ISE); Canale e' Figu, m 576,
walnut-orchard, 1 male 24-30.VIII.2007 (ISE); Mannaritzò, m 706, cherry-orchard, 1 female 1118.IV.2007, 1 female 16-23.V.2007, 2 males
24-30.VIII.2007, 2 males 4-11.X.2007 (all pitfall
traps and leg. I.S.E.). Desulo, Gennargentu, mountain
refuge, m 1500, under Juniperus sp., pitfall traps, 1
male 11-25.VII.2006, 1 male 25.VII-9.VIII.2006
leg. Fiori F. (ISE). Fonni, Gennargentu, mountain refuge, m 1550, under Juniperus sp., pitfall traps, 1
male 19.VII-2.VIII.2006, 3 males 2-17.VIII.2006, 1
female 5-19.VI.2007, leg. Fiori F. (ISE). Siniscola,
Monte Albo, on the road to Lula, m 800, pitfall traps,
1 female 19-26.IV.2003, leg. Pantini P., Magnati F.
(MBG). Tonara, Su Azzime, m 682, walnut-orchard,
pitfall traps, 2 females 16-23.V.2007, 1 female 2128.VI.2007, 1 female 17-24.VII.2007, 1 female 2430.VIII.2007, 1 female 4-11.X.2007 leg. I.S.E. (ISE).
OLBIA-TEMPIO: Olbia, Isola Molarotto, 1 female 18.V.1966 leg. Gruppo Isole C.N.R. (MBG).
ORISTANO: Cabras, Isola Mal di Ventre, 5 females 17-18.IV.2006 leg. Mulas A. (UCA).
Zelotes tenuis (L. Koch, 1866)
REFERENCES. (133) Trotta, 2009; (137) Senglet,
2011.
PROVINCE. Cagliari (137); Medio-Campidano
(ined.); Nuoro (137, ined.); Ogliastra (137); OlbiaTempio (133); Sassari (137).
HABITAT. Cherry-orchard (ined.); dam leaf litter
(137); lagoon (137); hazelnut-orchard (ined.); walnut-orchard (ined.); under Juncus (137); mining
females 30.VI-13.VII.2009, 3 males and 2 females
15-30.VI.2009, 1 male 3-15.VI.2009, 2 females 1327.VII.2009 leg. Sassu A., Verdinelli M. (ISE).
NUORO: Belvì, Antonitzò, m 683, cherry-orchard, 4 males 21-28.VI.2007, 1 female 1724.VII.2007 (ISE); Canale e' Figu, m 576,
walnut-orchard, 2 males 21-28.VI.2007, 2 males and
1 female 17-24.VII.2007, 1 female 24-30.VIII.2007
(ISE); Mannaritzò, m 706, cherry-orchard, 1 female
21-28.VI.2007 (ISE); Su Enazzu, m 550, hazelnutorchard, 1 female 27.VII-8.VIII.2006, 1 male and 1
female 13-27.VII.2006 (MBG), 1 male and 1 female
13-27.VII.2006 (ISE), 1 female 8-24.VIII.2006
Family SPARASSIDAE
Micrommata ligurina (C.L. Koch, 1845)
REFERENCES. (6) Magretti, 1880; (10) Costa,
1885; (13) Garneri, 1902; (20) Krauss, 1955; (84)
Pesarini, 1994; (114) Grill et al., 2005; (139)
Trotta, 2011.
PROVINCE. (20, 84, 114); Cagliari (13); Carbonia-Iglesias (6, 13, 139); Nuoro (ined.); Ogliastra
(13); Oristano (10).
HABITAT. Grass and shurbland (114); walnut-orchard (ined.).
EXAMINED MATERIAL. NUORO: Tonara, Su Azzime, m 682, walnut-orchard, pitfall traps, 1 male
18-26.IV.2007 leg. I.S.E. (ISE).
Micrommata virescens (Clerck, 1757)
REFERENCES. (6) Magretti, 1880; (10) Costa,
1885; (13) Garneri, 1902; (84) Pesarini, 1994; (132)
PROVINCE. (10, 84); Oristano (13); Oristano/Nuoro (6); Sassari (13, 132).
Olios argelasius (Walckenaer, 1805)
REFERENCES. (8) Costa, 1884 (sub O. spongitar-
73
sis); (13) Garneri, 1902 (sub Sparassus spongitartis); (127) Loru et al., 2008; (132) Pantini & Sassu,
2009; (139) Trotta, 2011.
PROVINCE. (127); Cagliari (13, ined.); CarboniaIglesias (139); Medio Campidano (139, ined.);
Nuoro (8); Oristano (8, ined.); Sassari (132).
HABITAT. Citrus-grove (ined.); mediterranean
maquis (ined.); hazelnut-orchard (127).
Maria, Punta su Pisu, m 14, 1 male and 1 female
XII.2006 leg. Ruzzante-Wilkins J. (MBG). Uta,
S’Appassiu, Agris farm, citrus-grove, beating tray,
1 female, 29.VI.2000, 1 female 10.VIII.2000, 1 female 1.IX.2000, 2 males 21.IX.2000, 1 female
12.X.2000, 1 male 4.X.2000, 1 female 2.XI.2000,
1 female 13.XI.2000 leg. CRAS (ISE).
male 21.IV-5.V.2009 leg. Sassu A., Verdinelli M.
(ISE). Guspini, Montevecchio, Piccalinna, m 260,
mediterranean maquis, pitfall traps, 1 male 315.VI.2009 leg. Sassu A., Verdinelli M. (ISE).
ORISTANO: Cabras, San Giovanni in Sinis, 1
male, 26.IV.2003, leg. Pantini P., Magnati F. (MBG).
Family PHILODROMIDAE
Philodromus aureolus (Clerck, 1757)
1902; (84) Pesarini, 1994; (110) Muster & Thaler,
2004; (139) Trotta, 2011.
PROVINCE. (84); Cagliari (13); Carbonia-Iglesias (13, 139); Nuoro (8, 13); Ogliastra (13, 110).
Philodromus bistigma Simon, 1870
1902; (123) Muster et al., 2007; (132) Pantini &
Sassu, 2009.
PROVINCE. Cagliari (13, 123, ined.); MedioCampidano (ined.); Nuoro (123, ined.);
Nuoro/Ogliastra (123); Ogliastra (13, 123); Oristano (10, 123); Sassari (132, ined.).
HABITAT. Cherry-orchard (ined.); mediterranean
maquis (ined.); hazelnut-orchard (ined.).
74
EXAMINED MATERIAL. CAGLIARI: Pula, Is Cannoneris, m 715, 1 female, 18.VIII.2006, leg. Ruzzante-Wilkins (MBG).
male 4-16.VI.2009 leg. Sassu A., Verdinelli M. (ISE).
NUORO: Belvì, Mannaritzò, m 706, cherry-orchard, 1 female 21-28.VI.2007 (ISE); Su Enazzu,
m 550, hazelnut-orchard, 1 female 27.VII8.VIII.2006, 1 male 30.V-4.VII.2007 (MBG), 2
males and 1 female 30.V-4.VII.2007 (ISE) (all pitfall traps and leg. I.S.E.).
SASSARI: Olmedo, Pulpazos, Perlina farm, 1
male 8.V.2008 leg. R.A. Pantaleoni (ISE).
Philodromus blanckei (Wunderlich, 1995)
REFERENCES. (87) Wunderlich, 1995b (sub Artanes blanckei); (131) Muster, 2009.
PROVINCE. Nuoro (131, ined.); Ogliastra (87,
131); Olbia-Tempio (131); Oristano (131).
HABITAT. Under the bark (87), hazelnut-orchard
(ined.).
Enazzu, m 550, hazelnut-orchard, beating tray, 1 female 4.V.2005 leg. I.S.E. (MBG).
*Philodromus bosmansi Muster & Thaler, 2004
HABITAT. Citrus-grove (ined.).
EXAMINED MATERIAL. CAGLIARI: Uta, S’Appassiu, Agris farm, citrus-grove, beating tray, 1 female 29.VI.2000 leg. CRAS (MBG).
of this species for Italian and European fauna.
Philodromus buchari Kubcová, 2004
Punti, Baldinca, m 110, 1 male 15.VI.2008 leg.
Sassu A. (MBG).
*Philodromus buxi Simon, 1884
HABITAT. Alder-grove (ined.).
EXAMINED MATERIAL. NUORO: Desulo, Gennargentu, mountain refuge, m 1500, alder-grove,
bark traps, 3 females 25.VI-2.VII.2007 leg. Fiori
F. (ISE).
Philodromus dispar Walckenaer, 1826
REFERENCES. (13) Garneri, 1902; (139) Trotta, 2011.
PROVINCE. Cagliari (13); Carbonia-Iglesias (139).
Philodromus emarginatus (Schrank, 1803)
Philodromus fuscolimbatus Lucas, 1846
REFERENCES. (10) Costa, 1885 (sub P. constellatus).
PROVINCE. Carbonia-Iglesias (10); Olbia-Tempio (10).
Philodromus glaucinus Simon, 1870
1885; (13) Garneri, 1902; (123) Muster et al., 2007.
PROVINCE. Cagliari (13); Carbonia-Iglesias (10,
11, 13); Medio Campidano (123); Oristano (13);
Sassari (123).
Philodromus histrio (Latreille, 1819)
1994.
PROVINCE. (13, 84); Sassari (ined.).
Mannu, Nali farm, orchard, bark traps, 1 male and
1 female 20-27.V.1998 leg. Ist. Ent. SS (ISE).
Philodromus lividus Simon, 1875
1902; (110) Muster & Thaler, 2004; (114) Grill et
al., 2005; (127) Loru et al., 2008; (132) Pantini &
Sassu, 2009; (139) Trotta, 2011.
PROVINCE. (114, 127); Cagliari (13, 110); Carbonia-Iglesias (10, 13, 139); Nuoro (ined.); Ogliastra (13, 110); Sassari (132, ined.).
HABITAT. Low shurbs, grass and shurbland,
ined.); olive-grove (ined.).
male 29.VI.2005 leg. I.S.E. (MBG). Tiana, m 522,
hazelnut-orchard, beating tray, 1 male and 1 juv.
29.VI.2005, 1 female and 2 juv. 16.VI.2005 leg.
I.S.E. (ISE).
olive-grove, beating tray, 1 male 3.VI.1993 leg. Ist.
Ent. SS (ISE).
Philodromus margaritatus (Clerck, 1757)
REFERENCES. (2c) Canestrini, 1876 (sub P. elegans); (131) Muster, 2009; (139) Trotta, 2011.
PROVINCE. (2c); Medio Campidano (139);
Nuoro (131).
REMARKS. For the correct assignment of Canestrini’s citation (1876) see Muster (2009: 149).
Philodromus praedatus O.P.-Cambridge, 1871
REFERENCES. (94) Arnò et al., 1998; (110) Muster & Thaler, 2004; (132) Pantini & Sassu, 2009.
(110); Ogliastra (110); Sassari (94, 132, ined.).
(ined.); mediterranean maquis (ined.); vineyard
(ined.).
maquis, pitfall traps, 1 male, 16-29.VI.2009, leg.
SASSARI: Sorso, Pianu Mannu, Nali farm, orchard, beating tray, 1 female 17.VII.1997 leg. Ist.
Ent. SS (ISE). Torralba, vineyard, 1 male and 1 female 24.V.1999 leg. Ist. Ent. SS (ISE).
75
Philodromus pulchellus Lucas, 1846
REFERENCES. (2c) Canestrini, 1876; (13) Garneri,
1902; (84) Pesarini, 1994; (123) Muster et al., 2007;
(127) Loru et al., 2008; (132) Pantini & Sassu, 2009.
PROVINCE. (84, 127); Cagliari (2c, 13); Medio
Campidano (123); Nuoro (123, ined.); Olbia-Tempio (123); Oristano (123); Sassari (123, 132).
leg. I.S.E. (ISE).
Philodromus rufus Walckenaer, 1826
et al., 2008.
PROVINCE. (127); Nuoro (ined.); Sassari (94,
ined.).
Enazzu, m 550, hazelnut-orchard, beating tray, 2 females 1.VI.2005 leg. I.S.E. (MBG). Tiana, m 522,
hazelnut-orchard, beating tray, 1 female 1.VI.2005
leg. I.S.E. (ISE).
SASSARI: Sorso, Pianu Mannu, Nali farm, olivegrove, 1 male 29.IV.1993 (ISE); olive-grove, beating
tray, 1 female 21.V.1993, 2 females and 4 juv.
25.VI.1993 (ISE); orchard, beating tray, 1 female
26.VI.1997 (ISE) (all leg. Ist. Ent. SS). Torralba, vineyard, 1 female 18.VI.1994, 1 female 15.VII.1995, 1
male and 1 female 24.V.1999 leg. Ist. Ent. SS (ISE).
Thanatus formicinus (Clerck, 1757)
PROVINCE. (13).
Thanatus oblongiusculus (Lucas, 1846)
CHOROTyPE.Sibero-European.
REFERENCES. (10) Costa, 1885 (sub T. oblongiusculus); (84) Pesarini, 1994 (sub Paratibellus
oblongiusculus).
76
Thanatus vulgaris Simon, 1870
REFERENCES. (8) Costa, 1884; (13) Garneri, 1902;
(84) Pesarini, 1994; (132) Pantini & Sassu, 2009.
PROVINCE. (84); Cagliari (13); Medio-Campidano (ined.); Ogliastra (13); Sassari (8, 13, 132).
HABITAT. Under stones on the beach (8); mining
covered by pioneer plants, pitfall traps, 2 males,
19.V-4.VI.2009, 1 female 29.VI-14.VII.2009, 1 female 4-16.VI.2009 leg. Sassu A., Verdinelli M. (ISE).
Tibellus oblongus (Walckenaer, 1802)
1902.
PROVINCE. Cagliari (10, 13); Oristano (13); Sassari (10).
Family THOMISIDAE
Cozyptila blackwalli (Simon, 1875)
REFERENCES. (13) Garneri, 1902 (sub Oxyptila
blackwalli); (84) Pesarini, 1994 (sub Ozyptila
blackwalli Simon, 1875).
PROVINCE. (84); Cagliari (13); Sassari (ined.).
Mannu, Nali farm, olive-grove, beating tray, 1 female 1.VII.1993 leg. Ist. Ent. SS (ISE).
Diaea dorsata (Fabricius, 1777)
Heriaeus hirtus (Latreille, 1819)
REFERENCES. (10) Costa, 1885 (sub H. hirsutus);
(11) de Carlini, 1885 (sub Misumena savignyi); (13)
Garneri, 1902 (sub H. savignyi); (20) Krauss, 1955;
(71) Loerbroks, 1983; (84) Pesarini, 1994.
PROVINCE. (20, 84); Cagliari (13); CarboniaIglesias (11, 13); Nuoro (10, 71).
Misumena vatia (Clerck, 1757)
REFERENCES. (11) de Carlini, 1885; (13) Garneri,
PROVINCE. (84, 127); Carbonia-Iglesias (11, 13).
HABITAT. Hazelnut-orchard (127).
Monaeses paradoxus (Lucas, 1846)
REFERENCES. (114) Grill et al., 2005.
PROVINCE. (114).
Ozyptila atomaria (Panzer, 1801)
horticola); (84) Pesarini, 1994.
Ozyptila bicuspis Simon, 1932
HABITAT. Grassland along the mouth of the river
(116).
Ozyptila brevipes (Hahn, 1826)
brevipes); (84) Pesarini, 1994.
Ozyptila confluens (C.L. Koch, 1845)
REFERENCES. (3) Pavesi, 1876 (sub Oxyptila gut-
tulata); (13) Garneri, 1902 (sub Oxyptila grata); (20)
Sassu, 2009; (133) Trotta, 2009; (139) Trotta, 2011.
Medio Campidano (139, ined.); Nuoro (ined.); OlbiaTempio (3, 133); Oristano (13); Sassari (132, 133).
minning landfills covered by pioneer plants (ined.).
Arbus, Ingurtosu, Naracauli, m 170, mining landfills covered by pioneer plants, 3 males 620.X.2009 (ISE); m 200, garrigue, 2 males,
22.IX-6.X.2009 (ISE), 2 males and 1 female 620.X.2009 (MBG), 1 male and 3 females 620.X.2009 (ISE) (all pitfall traps and leg. Sassu A.,
Verdinelli M.). Guspini, Montevecchio, Piccalinna,
m 235, garrigue, pitfall traps, 5 males 22.IX6.X.2009, 5 males 6-20.X.2009 (ISE), 2 males and
1 female 6-20.X.2009 (MBG) (all leg. Sassu A.,
Verdinelli M.).
NUORO: Belvì, Antonitzò, m 683, cherry-orchard, 2 males 24-30.VIII.2007, 1 male 27.IX4.X.2007 (ISE); Canale e' Figu, m 576,
walnut-orchard, 1 female 26.IV-4.V.2007 (ISE); Su
Enazzu, m 550, hazelnut-orchard, 1 male 24.VIII5.IX.2006 (MBG), 2 males 21.IX-5.X.2006 l. (ISE)
(all pitfall traps and eg. I.S.E.). Tonara, Su Azzime,
m 682, walnut-orchard, pitfall traps, 1 female 1724.VII.2007, 2 males 4-11.X.2007 leg. I.S.E. (ISE).
Ozyptila pauxilla (Simon, 1870)
Ozyptila praticola (C.L. Koch, 1837)
*Ozyptila sanctuaria (O.P.-Cambridge, 1871)
(ined.).
77
Arbus, Ingurtosu, Naracauli, m 170, mining landfills covered by pioneer plants, pitfall traps, 1 female 6-20.X.2009 leg. Sassu A., Verdinelli M.
(ISE). Guspini, Montevecchio, Piccalinna, m 225,
mining landfills covered by pioneer plants, 1 female
5-18.V.2009 (ISE); m 235, garrigue, 1 female 1530.VI.2009, 5 males 6-20.X.2009 (ISE) (all pitfall
NUORO: Belvì, Mannaritzò, m 706, cherry-orchard, pitfall traps, 1 male 19-26.XI.2007 leg. I.S.E.
(ISE).
Pistius truncatus (Pallas, 1772)
REFERENCES. (85) Buchar & Thaler, 1995; (127)
Loru et al., 2008.
PROVINCE. (127); Nuoro (85); Olbia-Tempio
(ined.).
HABITAT. Hazelnut-orchard (127); forest of
EXAMINED MATERIAL. OLBIA-TEMPIO: Buddusò, near Nuraghe Loelle, forest of Quercus suber,
beating tray, 1 female 24.V.1999 leg. Ist. Ent. SS
(ISE). Telti, Rio Almiddina, 1 male 20.V.2008 leg.
Sassu A. (ISE).
Runcinia grammica (C.L. Koch, 1837)
REFERENCES. (8) Costa, 1884 (sub R. lateralis);
(11) de Carlini, 1885 (sub R. lateralis); (13) Garneri, 1902 (sub R. lateralis); (84) Pesarini, 1994
(sub R. lateralis); (114) Grill et al., 2005 (sub R. lateralis); (139) Trotta, 2011.
PROVINCE. (84, 114); Cagliari (8, 13); CarboniaIglesias (11, 13, 139); Sassari (ined.).
HABITAT. Shurblands and trees, agricultural land
(114); vineyard (ined.).
EXAMINED MATERIAL. SASSARI: Torralba, vineyard, 1 female 30.VII.1994 leg. Ist. Ent. SS (ISE).
Synema globosum (Fabricius, 1775)
REFERENCES. (8) Costa, 1884 (sub S. globosa);
(11) de Carlini, 1885 (sub Diaea globosa); (13) Garneri, 1902 (sub S. globosa); (20) Krauss, 1955; (46)
78
Thermes, 1972; (84) Pesarini, 1994; (94) Arnò et al.,
1998 (sub Synaema globosum); (112) Thaler & Knoflach, 2004; (114) Grill et al., 2005; (127) Loru et
al., 2008; (132) Pantini & Sassu, 2009 (sub Synaema
globosum); (133) Trotta, 2009; (139) Trotta, 2011.
PROVINCE. (13, 20, 84, 114, 127); Cagliari (139);
Carbonia-Iglesias (11, 46, 139); Medio Campidano
(139); Nuoro (ined.); Ogliastra (112, 139); OlbiaTempio (133, ined.); Oristano/Nuoro (8); Sassari
(94, 132, 133, ined.).
habitat (46); low shurbs, grass and shurbland,
ined.); grassland (ined.); vineyard (ined.).
Enazzu, m 550, hazelnut-orchard, beating tray, 1 female 19.V.2005 leg. I.S.E. (MBG). Bolotana, Badde
e'Salighes, m 850, grassland, 1 female 27.V.2007
leg. Sassu A. (ISE).
OLBIA-TEMPIO: Telti, Rio Almiddina, 1 male
male and 1 female 22.V.2008 leg. Sassu A. (ISE).
Torralba, vineyard, 1 male 28.V.1994 leg. Ist. Ent.
SS (ISE).
Tmarus staintoni (O.P.-Cambridge, 1873)
1885; (13) Garneri, 1902 (sub T. albus Nomina
dubia); (20) Krauss, 1955; (84) Pesarini, 1994; (94)
Arnò et al., 1998; (114) Grill et al., 2005; (132) Pantini & Sassu, 2009.
PROVINCE. (13, 20, 84, 114); Carbonia-Iglesias
(11); Oristano/Nuoro (8); Sassari (94, 132, ined.).
HABITAT. Avenue of lime trees (94); shurblands
and trees (114); vineyard (ined.).
EXAMINED MATERIAL. SASSARI: Sassari, Platamona, at sea level, 2 males and 1 female 22.V.2008
leg. Sassu A. (ISE). Torralba, vineyard, 1 male
PROVINCE. Cagliari (10); Nuoro (10, ined.).
HABITAT. Cherry-orchard (ined.); hazelnut-orchard (ined.).
EXAMINED MATERIAL. NUORO: Belvì, Mannaritzò, m 706, cherry-orchard, 1 male 11-18.IV.2007
(ISE); Su Enazzu, m 550, hazelnut-orchard, 1 male
26.IV-30.V.2007 (ISE) (all pitfall traps and leg.
I.S.E.).
Thomisus onustus Walckenaer, 1805
Tmarus piochardi (Simon, 1866)
REFERENCES. (114) Grill et al., 2005.
PROVINCE. (114).
Sassari (ined.).
HABITAT. Orchard (ined.); hazelnut-orchard
(127, ined.); olive-grove (ined.).
EXAMINED MATERIAL. CAGLIARI: Pula, Is Cannoneris, m 715, 1 female 18.VIII.2006 leg. Ruzzante-Wilkins (MBG).
NUORO: Tiana, m 522, hazelnut-orchard, beating tray, 1 female 1.VI.2005 (MBG), 1 female
29.VI.2005 (ISE) (all leg. I.S.E.).
SASSARI: Alghero, Mamuntanas, olivegrove,
chemical knockdown, 1 male 6.VIII.2009 (MBG),
1 male and 1 female 27.VIII.2009, 2 females,
7.IX.2009 (ISE), 2 females, 3.X.2009 (MBG), 2 females 27.X.2009 (ISE) all leg. Ist. Ent. SS. Sassari,
Lu Buttangaru, olive-grove, 3 females 30.IX.2008
farm, olive-grove, 1 female, 9.VII.1993, 1 female
6.VIII.1993 (ISE); orchard, 1 female 26.VI.1997
(ISE) (all beating tray and leg. Ist. Ent. SS).
Xysticus acerbus Thorell, 1872
Xysticus audax (Schrank, 1803)
REFERENCES. (13) Garneri, 1902 (sub X. pini);
PROVINCE. (84); Cagliari (13); Ogliastra (13).
Xysticus bifasciatus C.L. Koch, 1837
1994.
Xysticus bliteus (Simon, 1875)
REFERENCES. (5) Pavesi, 1878; (13) Garneri,
1902; (84) Pesarini, 1994 (sub Ozyptila blitea);
(132) Pantini & Sassu, 2009.
Olbia-Tempio (5, 13); Sassari (132).
plants (ined.).
Arbus, Ingurtosu, Naracauli, m 170, mining landfills covered by pioneer plants, pitfall traps, 2
males, 22.IX-6.X.2009 (ISE) 1 male 22.IX6.X.2009 (MBG), 1 male and 1 juv. 8-22.IX.2009
(ISE) (all leg. Sassu A., Verdinelli M.).
Xysticus bufo (Dufour, 1820)
REFERENCES. (3) Pavesi, 1876 (sub Oxyptila albimana); (13) Garneri, 1902 (sub Oxyptila bufo); (84)
Pesarini, 1994; (89) Wunderlich, 1995d (sub Psammitis bufo); (133) Trotta, 2009; (139) Trotta, 2011.
(3, 13, 133, 139); Medio Campidano (ined.); Nuoro
(ined.); Sassari (89, ined.).
(ined.); olive-grove (ined.); mining landfills covered by pioneer plants (ined.).
Arbus, Ingurtosu, Naracauli, m 170, mining landfills covered by pioneer plants, pitfall traps, 1
male, 22.IX-6.X.2009, leg. Sassu A., Verdinelli M.
(MBG). Guspini, Montevecchio, Piccalinna, m
235, garrigue, pitfall traps, 1 male 22.IX6.X.2009, 1 male 6-20.X.2009 leg. Sassu A., Verdinelli M. (ISE).
NUORO: Belvì, Mannaritzò, m 706, cherry-orchard, pitfall traps, 1 male 4-11.X.2007 leg. I.S.E.
(ISE). Gavoi, S'eremu e sa mela, m 1000, 1 female
30.XII.2006 leg. Mulas A. (MBG).
SASSARI: Alghero, Mamuntanas, olivegrove,
chemical knockdown, 1 male 13.X.2009 leg. Ist.
Ent. SS (ISE). Sassari, Lu Buttangaru, olive-grove,
79
chemical knockdown, 1 male 29.IX.2008 leg. Ist.
Ent. SS (ISE).
Xysticus caperatus Simon, 1875
REFERENCES. (10) Costa, 1885; (84) Pesarini,
1994.
Xysticus cor Canestrini, 1873
REFERENCES. (2b) Canestrini, 1873; (2c) Canestrini, 1876; (13) Garneri, 1902; (18) Simon, 1932;
PROVINCE. (2b, 2c, 13, 18, 84).
Xysticus cristatus (Clerck, 1757)
1994.
*Xysticus erraticus (Blackwall, 1834)
HABITAT. Hazelnut-orchard (ined.); alder-grove
(ined.).
Enazzu, m 550, hazelnut-orchard, pitfall traps, 1 female 30.V-4.VII.2007 leg. I.S.E. (ISE). Fonni, Gennargentu, mountain refuge, m 1550, alder-grove,
pitfall traps, 1 female 26.IX-10.X.2006, 7 males and
2 females 5-19.VI.2007, 1 male 19.VI-1.VII.2007
leg. Fiori F. (ISE).
Xysticus graecus C.L. Koch, 1837
REFERENCES. (84) Pesarini, 1994; (139) Trotta,
2011.
PROVINCE. (84); Nuoro (139).
Xysticus grallator Simon, 1932
80
1994; (139) Trotta, 2011.
PROVINCE. (84); Medio Campidano (139);
Olbia-Tempio (20).
*Xysticus ibex Simon, 1875
1 female and 2 juv. 29.VI.2005 (ISE), 1 male
1.VI.2005 (MBG); Su Enazzu, m 550, hazelnutorchard, 2 females and 1 juv. 16.VI.2005 (MBG)
(all beating tray and leg. I.S.E.). Fonni, Talesso,
1 male 21.V.2008 leg. Pantaleoni R.A. and Loru
L. (ISE).
Xysticus kochi Thorell, 1872
REFERENCES. (46) Thermes, 1972; (84) Pesarini,
1994.
CHOROTyPE. Centro-European.
Mannu, Nali farm, orchard, pitfall traps, 1 male 1018.II.1998 leg. Ist. Ent. SS (ISE).
1902; (20) Krauss, 1955; (84) Pesarini, 1994; (127)
Loru et al., 2008.
PROVINCE. (20, 84, 127); Cagliari (8, 13); Nuoro
(8, 13, ined.); Oristano (13); Sassari (ined.).
(ined.); hazelnut-orchard (127, ined.).
EXAMINED MATERIAL. NUORO: Belvì, Mannaritzò, m 706, cherry-orchard, pitfall traps, 1 male
18-26.IV.2007, 2 males 16-23.V.2007 (ISE); Su
Ajone, m 585, hazelnut-orchard, beating tray, 1 female and 6 juv. 29.VI.2005 (MBG), 1 female and 2
juv. 1.VI.2005 (ISE) (all leg. I.S.E.).
OLBIA-TEMPIO: Olbia, Isola Molara, 2 females 25.II.1966 (MBG); Isola Tavolara, 1 female
24.II.1966 (MBG) (all leg. Gruppo Isole C.N.R.).
SASSARI: Sorso, Pianu Mannu, Nali farm, orchard, pitfall traps, 1 male 15-22.IV.1998 leg. Ist.
Ent. SS (ISE).
Xysticus lanio C.L. Koch, 1835
1902; (84) Pesarini, 1994; (127) Loru et al., 2008;
(139) Trotta, 2011.
PROVINCE. (8, 13, 84, 127); Carbonia-Iglesias
(139); Nuoro (ined.).
Ajone, m 585, hazelnut-orchard, 1 male and 1 juv.
19.V.2005 (MBG), 1 male and 2 juv. 1.VI.2005,
Xysticus luctuosus (Blackwall, 1836)
Xysticus nubilus Simon, 1875
REFERENCES. (18) Simon, 1932; (20) Krauss,
1955; (84) Pesarini, 1994.
PROVINCE. (18, 20, 84).
Xysticus parallelus Simon, 1873
REFERENCES. (18) Simon, 1932.
PROVINCE. (18).
Xysticus sardiniensis (Wunderlich, 1995)
REFERENCES.(89) Wunderlich, 1995d (sub
Psammitis sardiniensis).
Family SALTICIDAE
Aelurillus monardi (Lucas, 1846)
REFERENCES. (13) Garneri, 1902 (sub A. Monardi); (84) Pesarini, 1994; (115) Hansen, 2005;
(132) Pantini & Sassu, 2009.
(13); Nuoro (115); Olbia-Tempio (115); Sassari
(115, 132).
Aelurillus v-insignitus (Clerck, 1757)
REFERENCES. (13) Garneri, 1902 (sub Aelurops
insignita); (84) Pesarini, 1994.
PROVINCE. (13, 84).
Asianellus festivus (C.L. Koch, 1834)
REFERENCES. (13) Garneri, 1902 (sub Aelurops festiva); (84) Pesarini, 1994 (sub Aelurillus
festivus).
Ballus armadillo (Simon, 1871)
1994; (127) Loru et al., 2008.
Nuoro (ined.).
m 585, hazelnut-orchard, 1 female 19.V.2005 (ISE),
1 female 29.VI.2005, 1 female 27.VII.2005 (MBG);
Su Enazzu, m 550, hazelnut-orchard, 1 female
19.V.2005 (ISE), 1 female 19.V.2005 (MBG) (all
beating tray and leg. I.S.E.).
Ballus chalybeius (Walckenaer, 1802)
REFERENCES. (13) Garneri, 1902 (sub B. depressus); (84) Pesarini, 1994 (sub B. depressus); (115)
Hansen, 2005 (sub B. depressus); (127) Loru et al.,
2008; (132) Pantini & Sassu, 2009.
PROVINCE. (84, 127); Nuoro (ined.); Ogliastra
(13); Sassari (115, 132, ined.).
1.VI.2005 leg. I.S.E. (ISE).
olive-grove, beating tray, 1 female 1.VII.1993 leg.
Ist. Ent. SS (ISE).
81
Bianor albobimaculatus (Lucas, 1846)
REFERENCES. (8) Costa, 1884 (sub Eris albobimaculata); (13) Garneri, 1902 (sub Eris albo-bimaculata); (20) Krauss, 1955; (84) Pesarini, 1994; (91)
Fontana et al., 1996; (115) Hansen, 2005.
PROVINCE. (20, 84, 91); Cagliari (8, 13); Nuoro
(115).
HABITAT. Damp area (91).
Carrhotus xanthogramma (Latreille, 1819)
REFERENCES. (115) Hansen, 2005; (127) Loru et
al., 2008.
PROVINCE. (127); Cagliari (115); Nuoro (ined.).
m 585, hazelnut-orchard, 1 male 1.VI.2005 (ISE);
Su Enazzu, m 550, hazelnut-orchard, 1 male and 2
females 19.V.2005 (MBG) (all beating tray and leg.
I.S.E.). Tiana, m 522, hazelnut-orchard, beating
tray, 1 female 4.V.2005 (MBG), 1 female 19.V.2005
(ISE) (all leg. I.S.E.).
Chalcoscirtus infimus (Simon, 1868)
REFERENCES. (115) Hansen, 2005; (116) Helsdingen, 2005; (132) Pantini & Sassu, 2009.
(116, ined.); Ogliastra (115, 116); Olbia-Tempio
(ined.); Oristano (ined.); Sassari (132, ined.).
HABITAT. Coastal dunes (ined.); garrigue (ined.);
in drift (116); walnut-orchard (ined.); forest of
Quercus suber (ined.); vineyard (ined.).
Arbus, Ingurtosu, Naracauli, m 200, garrigue, pitfall
traps, 1 male 5-19.V.2009, 1 female 4-16.VI.2009
leg. Sassu A., Verdinelli M. (MBG).
NUORO: Belvì, Canale e' Figu, m 576, walnutorchard, 1 male, 21-28.VI.2007, leg. I.S.E. (MBG).
ORISTANO: Cabras, Sinis, at sea level, coastal
dunes, 4 males 20.IV.2003 leg. Pantini P., Magnati
F. (MBG).
forest of Quercus suber, beating tray, 1 male
82
SASSARI: Torralba, vineyard, 1 male 6.V.1995,
1 female 20.V.1995 leg. Ist. Ent. SS (ISE).
Cyrba algerina (Lucas, 1846)
1994; (115) Hansen, 2005; (139) Trotta, 2011.
PROVINCE. (84); Cagliari (115, 139); Nuoro
(ined.); Oristano (13); Sassari (115, ined.).
EXAMINED MATERIAL. NUORO: Dorgali, Cala Sisine, 1 male 21.IV.2003 leg. Ferrario E. (MBG).
SASSARI: Sassari, Li Punti, Baldinca, m 110, 1
male 23.V.2008 leg. Sassu A. (ISE).
*Dendryphantes hastatus (Clerck, 1757)
EXAMINED MATERIAL. SASSARI: Sassari, Platamona, at sea level, 2 females, 22.V.2008, leg. Sassu
A. (ISE).
Euophrys frontalis (Walckenaer, 1802)
1994.
Euophrys gambosa (Simon, 1868)
REFERENCES. (83) Pons, 1993; (116) Helsdingen, 2005; (132) Pantini & Sassu, 2009.
(83, 116); Sassari (132, ined.).
HABITAT. Hypogean habitat (83); orchard
(ined.); garrigue (ined.); grassland along the mouth
of the river (116); olive-grove (ined.); mining landfills covered by pioneer plants (ined.).
19.V-4.VI.2009 leg. Sassu A., Verdinelli M.
(MBG). Guspini, Montevecchio, Piccalinna, m 225,
mining landfills covered by pioneer plants, 1 male
3-15.VI.2009 (MBG); m 235, garrigue, 1 male
18.V-3.VI.2009, 2 males 3-15.VI.2009 (MBG) (all
SASSARI: Sassari, Gioscari, olive-grove, chemical knockdown, 2 males and 1 female 14.X.2008,
farm, orchard, beating tray, 1 male 16.V.1997 leg.
Ist. Ent. SS (ISE).
*Euophrys herbigrada (Simon, 1871)
Figu, m 576, walnut-orchard, 1 male 16-23.V.2007,
1 female 28.VI-24.VII.2007, 1 female 24.VII30.VIII.2007 (MBG) (all pitfall traps and leg. I.S.E.).
Tonara, Su Azzime, m 682, walnut-orchard, 1 female
23.V-28.VI.2007, 1 female 24.VII-30.VIII.2007
Euophrys rufibarbis (Simon, 1868)
CHOROTyPE. Centralasiatic-European-Mediterranean.
REFERENCES. (3) Pavesi, 1876; (13) Garneri,
1902; (84) Pesarini, 1994.
(3, 13); Medio Campidano (ined.).
22.IX-6.X.2009, 1 male 6-20.X.2009 leg. Sassu A.,
Verdinelli M. (MBG). Guspini, Montevecchio, Piccalinna, m 235, garrigue, pitfall traps, 1 female 1530.VI.2009 leg. Sassu A., Verdinelli M. (MBG).
Euophrys sulphurea (L. Koch, 1867)
CHOROTyPE. E-Mediterranean.
REFERENCES.(10) Costa, 1885; (84) Pesarini,
1994.
(10); Sassari (ined.).
HABITAT. Olive-grove (ined.); walnut-orchard
(ined.).
83
Figu, m 576, walnut-orchard, pitfall traps, 1 male
16-23.V.2007 leg. I.S.E. (MBG).
SASSARI: Sassari, Gioscari, olive-grove, chemical knockdown, 1 female 6.VIII.2008 leg. Ist.
Ent. SS (MBG).
chemical knockdown, 1 male 6.VIII.2009, 1 male
30.IX.2009, 1 male 7.X.2009 leg. Ist. Ent. SS (ISE).
Torralba, vineyard, 1 male and 1 female 23.VII.1994,
1 female 1.X.1994, 1 female 29.VII.1995, 1 male
22.VII.1995, 1 male and 1 female 23.IX.1995 leg. Ist.
Ent. SS (ISE).
REFERENCES. (6) Magretti, 1880 (sub Attus terrestris); (13) Garneri, 1902; (84) Pesarini, 1994.
PROVINCE. (84); Nuoro (13); Oristano/Nuoro (6);
Sassari (13).
REFERENCES.(20) Krauss, 1955; (46) Thermes,
(115) Hansen, 2005; (127) Loru et al., 2008; (132)
Pantini & Sassu, 2009; (139) Trotta, 2011.
PROVINCE. (84, 127); Cagliari (46, 139); Carbonia-Iglesias (139); Medio Campidano (139); Nuoro
(ined.); Ogliastra (115); Olbia-Tempio (115); Sassari (20, 94, 132, ined.).
habitat (46); cherry-orchard (ined.); orchard (ined.);
hazelnut-orchard (127, ined.); walnut-orchard
(ined.); olive-grove (ined.); vineyard (ined.).
m 683, cherry-orchard, 1 male 11-18.IV.2007, 1 male
27.IX-4.X.2007 (ISE); Canale e' Figu, m 576, walnut-orchard, 1 male 22-29.II.2008 (ISE); Mannaritzò, m 706, cherry-orchard, 2 males 18-26.IV.2007
(ISE); Su Ajone, m 585, hazelnut-orchard, 1 female
19-27.VII.2005 (ISE) (all in bark traps and leg.
I.S.E.). Tonara, Su Azzime, m 682, walnut-orchard,
bark traps, 1 male 18-26.IV.2007 leg. I.S.E. (ISE).
chemical knockdown, 1 male 7.X.2009 leg. Ist. Ent.
SS (ISE). Sorso, Pianu Mannu, Nali farm, orchard,
bark traps, 1 female 26.VI-2.VII.1997, 1 male 29.XII.1997 leg. Ist. Ent. SS (ISE). Torralba, vineyard, 1 male 14.V.1994, 2 females 4.VI.1994, 1
female and 4 juv. 9.VII.1994, 1 male and 1 juv.
24.IX.1994, 1 female 6.V.1995, 1 male 17.VI.1995,
1 female 1.VII.1995, 4 males and 6 juv.
26.VIII.1995, 1 female 5.VIII.1995, 4 males 12
juv., 9.IX.1995, 1 male and 15 juv., 2.IX.1995, 2
males and 10 juv. 16.IX.1995, 10 males and 4 juv.
30.IX.1995, 8 males and 2 juv. 23.IX.1995, 13
males, 1 female and 3 juv. 7.X.1995, 1 male and 1
juv. 14.X.1995 leg. Ist. Ent. SS (ISE).
Euophrys terrestris (Simon, 1871)
Evarcha falcata (Clerck, 1757)
REFERENCES. (13) Garneri, 1902 (sub Hasarius
falcatus); (84) Pesarini, 1994.
Evarcha jucunda (Lucas, 1846)
REFERENCES. (8) Costa, 1884 (sub Hasarius jucundus); (13) Garneri, 1902 (sub Hasarius jucundus); (20) Krauss, 1955; (84) Pesarini, 1994; (114)
Grill et al., 2005; (115) Hansen, 2005; (127) Loru
et al., 2008; (139) Trotta, 2011.
PROVINCE. (84, 114, 127); Cagliari (115, ined.);
Carbonia-Iglesias (139); Nuoro (8, 13, 115, ined.);
Ogliastra (13); Olbia-Tempio (20, 115); Sassari
(ined.).
(ined.); shurblands and trees (114); vineyard (ined.).
EXAMINED MATERIAL. CAGLIARI: Uta, S’Appassiu, Agris farm, citrus-grove, beating tray, 1 female 11.IX.2000, 1 male 23.X.2000, 1 female
13.XI.2000 leg. CRAS (ISE).
NUORO: Belvì, Mannaritzò, m 706, cherry-orchard, 1 male and 1 female 16-23.V.2007 (ISE); Su
Enazzu, m 550, hazelnut-orchard, 1 male 521.IX.2006 (ISE) (all pitfall traps and leg. I.S.E.).
Tiana, m 522, hazelnut-orchard, beating tray, 1 male
19.V.2005 leg. I.S.E. (ISE).
Heliophanus apiatus Simon, 1868
Heliophanus auratus C.L. Koch, 1835
84
1994.
PROVINCE. (84); Oristano (13).
Heliophanus cupreus (Walckenaer, 1802)
1902; (20) Krauss, 1955; (30) Cerruti, 1968; (84)
Pesarini, 1994; (115) Hansen, 2005; (127) Loru et
al., 2008.
PROVINCE. (84, 127); Cagliari (13, 30); Nuoro
(ined.); Ogliastra (115); Oristano (10); Sassari (20).
HABITAT. Hypogean habitat (20, 30); hazelnutorchard (127, ined.).
4.V.2005 (ISE), 1 male 29.VI.2005 (MBG); Su
Enazzu, m 550, hazelnut-orchard, bark traps, 1 female, 16-23.VI.2005 (MBG), 1 female 1623.VI.2005 (ISE) (all leg. I.S.E.). Tiana, m 522,
hazelnut-orchard, beating tray, 1 female 19.V.2005,
1 female 1.VI.2005 leg. I.S.E. (MBG).
Heliophanus flavipes (Hahn, 1832)
1994; (115) Hansen, 2005.
PROVINCE. (84); Cagliari (13); Nuoro/Ogliastra
(13); Ogliastra (13); Olbia-Tempio (115).
Heliophanus kochii Simon, 1868
1994; (109) Logunov, 2004; (114) Grill et al., 2005;
(133) Trotta, 2009; (139) Trotta, 2011.
PROVINCE. (84, 114); Cagliari (13); CarboniaIglesias (13, 139); Medio Campidano (139); Ogliastra (13); Olbia-Tempio (109, 133); Sassari (ined.).
Murta Maria, 1 female 28.VI.1993 leg. Gusmini
(MBG).
Heliophanus melinus L. Koch, 1867
1994.
Heliophanus rufithorax Simon, 1868
1994; (115) Hansen, 2005.
PROVINCE. (10, 84); Cagliari (ined.); Nuoro
(115, ined.); Ogliastra (115); Olbia-Tempio (115);
HABITAT. Walnut-orchard (ined.); pinewood
Manna, m 140, pinewood, emergence cages, 1
male 21.IV.2009, 1 female 25.V.2009 leg. Ist. Ent.
SS (ISE).
NUORO: Belvì, Canale e' Figu, m 576, walnutorchard, pitfall traps, 1 female 21-28.VI.2007 leg.
I.S.E. (ISE).
SASSARI: Torralba, vineyard, 1 male 24.VI.1995
Heliophanus tribulosus Simon, 1868
REFERENCES. (78) Hansen, 1986.
PROVINCE. (78).
Icius congener Simon, 1871
REFERENCES. (20) Krauss, 1955 (sub I. nebulosus); (84) Pesarini, 1994.
PROVINCE. (20, 84).
Icius hamatus (C.L. Koch, 1846)
REFERENCES. (10) Costa, 1885 (sub I. striatus);
(13) Garneri, 1902 (sub I. striatus); (78) Hansen,
(115) Hansen, 2005; (127) Loru et al., 2008; (134);
Marras et al., 2011.
PROVINCE. (78, 84, 127, 134); Cagliari (13,
ined.); Nuoro (ined.); Oristano (10); Sassari (94,
115, ined.).
trees (94); cherry-orchard (ined.); orchard (134,
ined.); hazelnut-orchard (127, ined.); walnut-orchard (ined.); olive-grove (ined.); alder-grove
males and 5 females 19.VI.2000, 1 male and 3 females, 29.VI.2000, 2 females 31.VII.2000, 5 females 20.VII.2000, 2 males and 6 females
31.VIII.2000, 3 females 10.VIII.2000, 2 females
21.VIII.2000, 3 males and 2 females 21.IX.2000, 1
male and 1 female 11.IX.2000, 1 female 20.X.2000,
3 males and 5 females 23.X.2000, 3 males and 3 females 12.X.2000, 1 male and 2 females 4.X.2000,
1 male and 1 female 2.XI.2000, 1 male 23.XI.2000,
1 female 4.XII.2000 leg. CRAS (ISE).
NUORO: Belvì, Mannaritzò, m 706, cherry-orchard, bark traps, 1 male 18-26.IV.2007, 1 male
and 1 female 4-11.X.2007 (ISE); Su Ajone, m 585,
(ISE); Su Enazzu, m 550, hazelnut-orchard, bark
traps, 1 female 29.IX-6.X.2005 (MBG) (all leg.
1550, alder-grove, bark traps, 1 male 25.VII2.VIII.2006 leg. Fiori F. (ISE). Tiana, m 522, hazelnut-orchard, bark traps, 1 female 11-18.V.2005
(MBG), 1 female 23-29.VI.2005 (ISE), 2 females
23-29.VI.2005, 1 male 8-16.VI.2005 (MBG), 1 male
8-16.VI.2005 (ISE), 1 female 27.VII-5.VIII.2005, 1
female 19-27.VII.2005, 1 female 10-17.VIII.2005,
1 female 5-10.VIII.2005, 1 male 29.IX-6.X.2005, 1
female 6-12.X.2005 (MBG) (all leg. I.S.E.). Tonara,
Su Azzime, m 682, walnut-orchard, bark traps, 1
male 4-11.X.2007 leg. I.S.E. (ISE).
olive-grove, beating tray, 1 male, 29.III.1993, 1
male, 1 female and 8 juv. 29.IV.1993, 1 male
3.VI.1993, 1 male and 2 females 25.VI.1993, 1
male, 2 females and 3 juv. 9.VII.1993, 2 males, 1
female and 7 juv. 16.IX.1993, 1 male 9.IX.1993, 1
male, 1 female and 23 juv. 2.IX.1993, 1 male and 1
juv. 11.XI.1993 (ISE), olive-grove, 1 male and 1 female 6.V.1993 (ISE); orchard, bark traps, 2 males
24.XI-2.XII.1997 (ISE); orchard, 1 female
2.XII.1997 (ISE) (all leg. Ist. Ent. SS). Torralba,
vineyard, 1 male and 1 juv. 14.V.1994, 1 male, 2 females and 8 juv. 9.VII.1994, 1 male, 1 female and
1 juv. 3.VI.1995, 3 males and 1 female 24.VI.1995,
1 female 29.VII.1995, 1 male 22.VII.1995, 1 male
85
15.VII.1995, 1 male and 2 females 8.VII.1995, 1
female and 23 juv. 26.VIII.1995, 1 male, 2 females
and 17 juv. 16.IX.1995, 1 male and 18 juv.
2.IX.1995, 1 male and 14 juv. 14.X.1995, 2 males
and 2 females 7.X.1995, 2 males, 2 females and 7
juv. 21.X.1995 (ISE); vineyard, beating tray, 1 female 6.VIII.1994, 1 male and 11 juv. 10.IX.1994
Icius subinermis Simon, 1937
REFERENCES. (8) Costa, 1884 (sub I. Notebilis);
(13) Garneri, 1902 (sub I. Notebilis); (20) Krauss,
1955 (sub I. Notebilis); (84) Pesarini, 1994.
PROVINCE. (84); Nuoro (8, 13, ined.); Sassari (20).
EXAMINED MATERIAL. NUORO: Dorgali, on the
pathway to Su Gorroppu, m 200, 1 female
23.IV.2003 leg. Pantini P., Magnati F. (MBG).
Leptorchestes mutilloides (Lucas, 1846)
1994; (115) Hansen, 2005.
Macaroeris nidicolens (Walckenaer, 1802)
REFERENCES. (10) Costa, 1885 (sub Dendryphantes nidicolens); (84) Pesarini, 1994 (sub Eris
nidicolens); (132) Pantini & Sassu, 2009.
PROVINCE. (84); Olbia-Tempio (10, ined.); Sassari (132, ined.).
EXAMINED MATERIAL. OLBIA-TEMPIO: Telti, Rio
Almiddina, 1 male 20.V.2008 leg. Sassu A. (ISE).
chemical knockdown, 1 female 6.IX.2009 leg. Ist.
olive-grove, beating tray, 1 female 22.VII.1993 leg.
Ist. Ent. SS (ISE).
Marpissa muscosa (Clerck, 1757)
1994; (127) Loru et al., 2008.
86
Nuoro (ined.).
HABITAT. Hazelnut-orchard (127, ined.); walnut-orchard (ined.).
Enazzu, m 550, hazelnut-orchard, bark traps, 5
males, 5 females and 16 juv. 6-13.X.2004, 2 males
13-20.X.2004, 1 male, 2 females and 1 juv. 2027.X.2004, 2 males, 1 female and 7 juv. 1522.III.2005 leg. I.S.E. (MBG). Tonara, Su Azzime,
m 682, walnut-orchard, bark traps, 1 female 2430.VIII.2007 leg. I.S.E. (ISE).
Marpissa pomatia (Walckenaer, 1802)
REFERENCES.(13) Garneri, 1902; (84) Pesarini,
1994.
Mendoza canestrinii (Ninnii, 1868)
REFERENCES. (10) Costa, 1885 (sub Hyctia Canestrinii); (13) Garneri, 1902 (sub Hyctia Canestrinii); (84) Pesarini, 1994 (sub Mithion canestrinii).
PROVINCE. (13, 84); Oristano (10).
Menemerus semilimbatus (Hahn, 1829)
REFERENCES. (10) Costa, 1885 (sub M. semilimbactus); (11) de Carlini, 1885; (13) Garneri, 1902;
(84) Pesarini, 1994; (94) Arnò et al., 1998.
(11, 13); Nuoro (10); Ogliastra (13); Sassari (94).
Mogrus frontosus (Simon, 1871)
PROVINCE. (84).
REMARKS. Species known in Corse, Pesarini records it in his checklist without supplying any other
information, its presence in Sardinia needs further
confirmation.
Myrmarachne formicaria (De Geer, 1778)
REFERENCES. (13) Garneri, 1902 (sub Salticus
formicarius); (84) Pesarini, 1994; (139) Trotta,
2011.
PROVINCE. (13, 84); Carbonia-Iglesias (139).
Neaetha membrosa (Simon, 1868)
REFERENCES. (20) Krauss, 1955 (sub N. membranosa); (84) Pesarini, 1994.
HABITAT. Cherry-orchard (ined.); mining landfills covered by pioneer plants (ined.).
Arbus, Ingurtosu, Naracauli, m 170, mining landfills covered by pioneer plants, pitfall traps, 1 female 4-16.VI.2009, 1 female 29.VI-14.VII.2009
leg. Sassu A., Verdinelli M. (MBG). Guspini, Montevecchio, Piccalinna, m 225, mining landfills covered by pioneer plants, pitfall traps, 1 male
18.V-3.VI.2009, 1 male 3-15.VI.2009 leg. Sassu A.,
Verdinelli M. (MBG).
NUORO: Belvì, Mannaritzò, m 706, cherry-orchard, pitfall traps, 1 female 21-28.VI.2007 leg.
I.S.E. (ISE).
Pellenes arciger (Walckenaer, 1837)
REFERENCES. (10) Costa, 1885 (sub P. arcigerus); (13) Garneri, 1902 (sub P. arcigerus); (84) Pesarini, 1994; (100) Metzner, 1999.
PROVINCE. (84); Cagliari (10); Nuoro (10);
Ogliastra (13, 100); Sassari (ined.).
HABITAT. Olive-grove (ined.); Strand (100).
EXAMINED MATERIAL. SASSARI: Sassari, Gioscari, olive-grove, 1 male 20.VIII.2009 (MBG); Lu
Buttangaru, olive-grove, 2 males 29.IX.2009, 1 female 13.X.2009 (MBG) (all chemical knockdown
and leg. Ist. Ent. SS).
Pellenes geniculatus (Simon, 1868)
1994.
Pellenes nigrociliatus (Simon, 1875)
REFERENCES. (91) Fontana et al., 1996.
PROVINCE. Olbia-Tempio (91).
Philaeus chrysops (Poda, 1761)
1902; (84) Pesarini, 1994; (115) Hansen, 2005.
PROVINCE. (84); Cagliari (13, 115); Nuoro (10);
Oristano (10).
Phlegra bresnieri (Lucas, 1846)
CHOROTyPE. Afrotropico-Mediterranean.
REFERENCES. (7a) Costa, 1882 (sub Attus bresnieri); (8) Costa, 1884 [sub P. (Attus) Bresnieri];
(13) Garneri, 1902; (20) Krauss, 1955; (84) Pesarini, 1994; (100) Metzner, 1999; (115) Hansen,
2005; (120) Logunov & Azarkina, 2006; (132) Pantini & Sassu, 2009.
PROVINCE. (20, 84); Cagliari (8, 13); Medio
Campidano (120); Nuoro (ined.); Ogliastra (100,
115); Oristano (7a, 8, 13, 120); Sassari (132, ined.).
HABITAT. Cultivated land, olive and almond
trees, with grass Euphorbia and Cirsium (under
Granite rocks) (120); marshy grassland with Cirsium (under stone) (120); walnut-orchard (ined.);
olive trees with grass, sheep pasture and slightly
moist grass (120); olive-grove (ined.); stony pastures with stonewalls, Rubus fruticosus, Juniperus
communis (120); Strand (100).
EXAMINED MATERIAL. NUORO: Tonara, Su Azzime, m 682, walnut-orchard, pitfall traps, 1 female
16-23.V.2007 leg. I.S.E. (ISE).
SASSARI: Sassari, Gioscari, olive-grove, 1 male
22.IX.2008 l (MBG); Lu Buttangaru, olive-grove,
3 males and 1 female 29.IX.2009 (MBG) (all chemical knockdown and leg. Ist. Ent. SS).
Phlegra lineata (C.L. Koch, 1846)
87
1994.
PROVINCE. (13, 84).
Plexippus paykulli (Audouin, 1826)
REFERENCES. (13) Garneri, 1902 (sub Hasarius
Paykulli); (84) Pesarini, 1994.
Pseudeuophrys erratica (Walckenaer, 1826)
REFERENCES. (13) Garneri, 1902 (sub Euophrys
erratica); (84) Pesarini, 1994 (sub Euophrys erratica); (127) Loru et al., 2008.
PROVINCE. (84, 127); Cagliari (13); Nuoro
HABITAT. Hazelnut-orchard (127); walnut-orchard (ined.); alder-grove (ined.).
EXAMINED MATERIAL. NUORO: Fonni, Gennargentu, mountain refuge, m 1550, alder-grove,
bark traps, 1 female 27-4.X.2006 leg. Fiori F.
(ISE). Tonara, Su Azzime, m 682, walnut-orchard,
bark traps, 1 female 16-23.V.2007 leg. I.S.E. (ISE).
*Pseudeuophrys nebrodensis Alicata et Cantarella, 2000
HABITAT. Alder-grove (ined.); hazelnut-orchard
(ined.); under Juniperus sp. (ined.).
m 585, hazelnut-orchard, bark traps, 1 female 1825.V.2005, 1 female 23-29.VI.2005, 2 females 18.VI.2005, 1 female 8-16.VI.2005, 1 female
19-27.VII.2005 leg. I.S.E. (MBG). Desulo, Gennargentu, mountain refuge, m 1500, alder-grove, bark
traps, 2 females and 1 juv. 26.IX-3.X.2006, 1 female 5-13.IX.2006, 2 females 3-10.X.2006 (ISE);
under Juniperus sp., pitfall traps, 1 male 1024.V.2007 (ISE) (all leg. Fiori F.).
Pseudeuophrys vafra (Blackwall, 1867)
CHOROTyPE. European-mediterranean.
REFERENCES. (100) Metzner, 1999; (115) Hansen, 2005; (116) Helsdingen, 2005.
88
(116, ined.); Ogliastra (100, 115, 116).
HABITAT. Inside house (116); mediterranean
maquis (ined.); walnut-orchard (ined.); Strand (100).
Guspini, Montevecchio, Piccalinna, m 260, mediterranean maquis, pitfall traps, 2 males 15-30.VI.2009,
2 females 13-27.VII.2009 leg. Sassu A., Verdinelli
M. (MBG).
NUORO: Belvì, Canale e' Figu, m 576, walnutorchard, 1 female 17-24.VII.2007 leg. I.S.E. (MBG).
Pseudicius encarpatus (Walckenaer, 1802)
REFERENCES. (84) Pesarini, 1994; (94) Arnò et
al., 1998; (127) Loru et al., 2008.
PROVINCE. (84, 127); Nuoro (ined.); Sassari (94,
ined.).
HABITAT. Avenue of lime trees (94); cherry-orchard (ined.); orchard (ined.); hazelnut-orchard
(127); olive-grove (ined.).
EXAMINED MATERIAL. NUORO: Belvì, Antonitzò, m 683, cherry-orchard, 1 female 1724.VII.2007 (ISE); Mannaritzò, m 706,
cherry-orchard, 1 female 4-11.X.2007 (ISE) (all in
bark traps and leg. I.S.E.).
olive-grove, beating tray, 1 male 1.VII.1993 (ISE);
orchard, bark traps, 1 female 24-31.VII.1997, 1
male and 1 juv. 11-19.VI.1998 (ISE) (all leg. Ist.
Ent. SS).
Saitis barbipes (Simon, 1868)
REFERENCES. (115) Hansen, 2005.
Punti, Baldinca, m 110, 1 male 15.VI.2008 leg.
Sassu A. (ISE).
Salticus conjonctus (Simon, 1868)
1994; (115) Hansen, 2005.
PROVINCE. (20, 84); Olbia-Tempio (115).
Salticus mutabilis (Lucas, 1846)
REFERENCES. (13) Garneri, 1902 (sub Calliethera mutabilis); (84) Pesarini, 1994; (94) Arnò et
al., 1998; (115) Hansen, 2005; (132) Pantini &
Sassu, 2009; (133) Trotta, 2009.
(133, ined.); Ogliastra (115); Oristano (ined.), Sassari (94, 115, 132, ined.).
(ined.); olive-grove (ined.); vineyard (ined.).
EXAMINED MATERIAL. CARBONIA-IGLESIAS:
Sant’Antioco, 1 male 25.IV.2003 leg. Ferrario E.
(MBG).
ORISTANO: Cabras, San Giovanni in Sinis, 2
males and 1 juv. 26.IV.2003 leg Pantini P., Magnati
F. (MBG).
olive-grove, beating tray, 1 male 9.VI.1993 (ISE);
orchard, bark traps, 1 male 15-22.IV.1998, 1 female
11-19.VI.1998 (ISE) (all leg. Ist. Ent. SS). Torralba,
vineyard, 1 female 17.VI.1995 leg. Ist. Ent. SS (ISE).
Salticus scenicus (Clerck, 1757)
REFERENCES. (6) Magretti, 1880 (sub Epiblemum scenicum); (8) Costa, 1884 (sub Callethera
scenica); (13) Garneri, 1902 (sub Calliethera scenica); (84) Pesarini, 1994.
PROVINCE. (8, 84); Oristano (6, 13).
HABITAT. Common inside houses (8).
Salticus zebraneus (C.L. Koch, 1837)
PROVINCE. Nuoro (ined.); Sassari (94).
HABITAT. Avenue of lime trees (94); cherry-orchard (ined.).
EXAMINED MATERIAL. NUORO: Belvì, Antonitzò, m 683, cherry-orchard, bark traps, 1 male 2128.VI.2007 leg. I.S.E. (ISE).
Sitticus pubescens (Fabricius, 1775)
PROVINCE. (84).
Synageles albotrimaculatus (Lucas, 1846)
1994.
PROVINCE. (84); Carbonia-Iglesias (13); Sassari
(ined.).
EXAMINED MATERIAL. SASSARI: Torralba, vineyard, 1 female 24.VI.1995, 1 female 3.VI.1995 leg.
Ist. Ent. SS (ISE).
Synageles venator (Lucas, 1836)
1994.
*Talavera aequipes (O.P.-Cambridge, 1871)
Mannu, Nali farm, orchard, pitfall traps, 1 female
24-31.VII.1997 leg. Ist. Ent. SS (ISE).
Thyene imperialis (Rossi, 1846)
CHOROTyPE. Indo-Mediterranean.
REFERENCES. (8) Costa, 1884 (sub Thya imperialis); (13) Garneri, 1902; (20) Krauss, 1955; (84)
Pesarini, 1994; (114) Grill et al., 2005.
PROVINCE. (84, 114); Cagliari (8, 13); OlbiaTempio (20).
Yllenus univittatus (Simon, 1871)
CHOROTyPE. European (?).
1994;
PROVINCE. (84); Cagliari (10)
REMARKS. Species known only in France, Pesarini records it in his checklist probably on the basis
of Costa’s old citation (1885), its presence in Sardinia needs further confirmation.
89
CONSIDERATIONS ABOUT SARDINIAN
ARANEOFAUNA
In this work, 495 species are reported (30.6% of
those known in Italy) which belong to 229 genera
divided into 43 families. Among these, the Philodromidae Philodromus bosmansi is reported for the
first time in Europe, while the following are reported for the first time in Italy: the Gnaphosidae
Setaphis parvula, the Linyphiidae Centromerus
succinus and Hybocoptus corrugis and the Prodidomidae Zimirina brevipes. For the latter species
and for its family, this is the first sighting in Italy.
As well as these 6 species, a further 37 species are
reported for the first time in Sardinia (Table 2).
Regarding distribution (Fig. 3), Cagliari, Sassari and Nuoro are the provinces with the highest
number of species reported. In the Cagliari area,
the 218 species reported are related to 50 publications, however most of the data are linked to the old
works of Garneri (1902) and Costa (1884, 1885).
No specific studies have been carried out in the province of Cagliari but among the most recent contributions which report data about Sardinian
araneofauna, those by Argiolas et al. (1970), Thermes
(1972) and Trotta (2011) deserve a mention. A good
number of species were also reported on the basis of
newly examined material. Regarding the province of
Sassari (52 publications, 213 species) and the province of Nuoro (46 publications, 212 species),
newly examined material from studies in olive
groves, vineyards and fruit orchards in the Sassari
area and from studies in the mountainous environment of the Nuoro area was crucial. It was collected
by the Sassari section of the ISE and is published
in this catalogue. For the Sassari province we can
also mention the analysis of tree spiders collected
in a tree-lined street in Sassari (Arnò et al., 1998)
and the study of the spiders on the Island of Asinara
(Pantini & Sassu, 2009). We know less about the
other provinces: for Carbonia-Iglesias (39 publications, 141 species) Trotta’s recent contribution
(2011) is important as well as data from old publications; for Ogliastra area (43 publications, 133
species) contributions by Garneri (1902) and Helsdingen (2005) are useful; the newly examined material from studies in cork-oak forests and Trotta’s
publication (2009) on the islands around Sardinia
are important for Olbia-Tempio province (32 publications, 103 species); for Medio Campidano (16 pu-
90
Family and species
FILISTATIDAE
Pritha nana
THERIDIIDAE
Dipoena coracina
Rhomphaea nasica
Theridion hannoniae
LINyPHIIDAE
Family and species
zODARIIDAE
Zodarion gallicum
PRODIDOMIDAE
Zimirina brevipes *
GNAPHOSIDAE
Drassodes luteomicans
Leptodrassus femineus
Centromerus subalpinus
Micaria albovittata
Centromerus succinus *
Micaria cyrnea
Diplostyla concolor
Nomisia recepta
Hybocoptus corrugis *
Phaeocedus braccatus
Meioneta mollis
Setaphis parvula *
Micrargus subaequalis
Trachyzelotes pedestris
Nematogmus sanguinolentus
Zelotes dentatidens
Pocadicnemis juncea
Zelotes egregius
Syedra nigrotibialis
PHILODROMIDAE
Walckenaeria antica
Philodromus bosmansi *
Walck enaeria stylifrons
Philodromus buxi
ARANEIDAE
THOMISIDAE
Araneus marmoreus
Ozyptila sanctuaria
Araneus quadratus
Xysticus erraticus
LyCOSIDAE
Pirata tenuitarsis
zORIDAE
Xysticus ibex
SALTICIDAE
Dendryphantes hastatus
Zora manicata
AGELENIDAE
Textrix pinicola
LIOCRANIDAE
Agraecina lineata
Euophrys herbigrada
Pseudeuophrys nebrodensis
Talavera aequipes
Table 2. List of the species which appear to be newly reported for Sardinian araneofauna. Marked with an asterisk
are those newly reported for Italy.
91
blications, 101 species) the outstanding new material
examined from studies in a disused mining area and
Trotta’s recent contribution (2011) are fundamental
while for the province of Oristano (30 publications;
Chorotype
n. sp.
Cosmopolitan
%
12
2,40
Holarctic
59
11,90
Widely spread in the Palearctic Region
171
1
Subcosmopolitan
6
Paleartic
W-Paleartic
85
Asiatic-European
13
Centralasiatic-Europeo-Mediterranean
1
Sibero-European
Centralasiatic-European
Centralasiatic-Mediterranean
Turano-Europeo-Mediterranean
34
5
5
3
Turano-European
13
Europeo-Mediterranean
6
Turano-Mediterranean
5
1,20
100 species) reports mostly come from old works.
The chorotypes of the species discovered are
listed in table 3 and reproduced, in uniform groups
in figure 14. Together with a small percentage of
European
84
17,00
Centraleuropean
1
0,20
European
51
10,30
S-European
32
34,50
Mediterranean
109
22,00
0,20
W-Mediterranean
43
8,70
17,20
2,60
6,90
0,20
1,00
1,00
0,60
2,60
1,00
1,20
Mediterranean
60
6,50
12,10
Afrotropicals
6
1,20
Afrotropico-Mediterranean
1
0,20
Afrotropico-Indo-Mediterranean
Indo-Mediterranean
4
1
0,80
0,20
Endemic
54
10,90
Tyrrhenian endemic
12
2,40
Sardinian endemic
31
Italian endemic
Sardo-Corsican endemic
Table 3. List of chorotypes for the species present in Sardinia.
Figure 14. Chorological spectrum of Sardinian araneofauna.
1
10
0,20
2,00
6,30
92
Cosmopolitan or Sub-cosmopolitan species, more
than half of the species present in the Region have
a wide distribution divided into: prevalently Palearctic species (41.3%), Holartic and a small amount
of Afro-Tropical elements. The rest of them are divided into Mediterranean (20.8%) and Southern European (6.5%) elements. There is a remarkable
number of endemic species (10.9%) which involves
54 species. In figures 15-18 the Sardinian distributions of 38 of the 41 Sardinian and Sardo-Corsican
endemism are shown (only Zodarion sardum, Mogrus frontosus and Xysticus parallelus are generic
cited for Sardinia and no indicates localities)
while the distribution of the remaining 13 endemic species can be found in table 4.
CONCLUSIONS
It is possible that during our bibliographical research, some publications have been overlooked,
however we believe that the data presented provides
us with a detailed and updated view of what is
known about Sardinian araneofauna. Examination
of the material kept at the Museum of Bergamo and
that collected and kept at the Institute of Ecosystem
Study in Sassari has allowed us to insert a lot of
new data into the catalogue: as well as the 43 new
species reported on the island, we were able to confirm the presence of 211 other species in the region
and provide new information about their distribution throughout Sardinia.
The analysis of this material has also allowed
us to detect several species which were difficult
to attribute and can probably be considered as
new taxa belonging to the following genera: Harpactea, Rhode, Zodarion, Cybaeodes, Haplodrassus and Zelotes. Finally, the genitalia of
Phrurolithus corsicus, Zimirina brevipes, Drassodes luteomicans and Zelotes dentatidens are illustrated (Figs. 4-13).
The level of knowledge about the araneological
fauna of Sardinia can surely still be greatly improved. Compared with other Northern Italian regions (Table 5) such as Alto Adige (Noflatscher,
1996), Piedmont and Lombardy (Isaia et al., 2007)
and Veneto (Ballarin et al. 2011), which have recently been the object of similar catalogues, Sardinia
Species
Ariadna brignoli
Cteniza brevidens
Distribution
Described from Lazio (Isola di
Ponza), cited from Sardinia and
Sicily only for female specimens
Tyrrhenian endemic (see also the
remark on page 11)
Cteniza sauvagesi
Tyrrhenian endemic
Spermophorides
elevata
Tyrrhenian endemic
Dysdera ancora
Sardinia and Sicily
Dysdera ventricosa
Sardinia and Sicily. Only known
from its original description
Harpactea sardoa
Sardinia and Lazio
Gonatium
biimpressum
Rare species known from Sardinia, Corse (Bonifacio) and Lazio
(Isola di Ponza)
Syedra nigrotibialis
Rare species known from Sardinia, Corse and Lazio
Malthonica sicana
Sardinia and Sicily
Zodarion ruffoi
Sardinia and Apulia (Isole Tremiti)
Zelotes sardus
Pseudeuophrys
nebrodensis
Sardinia and Tuscany (Isola di
Capraia)
Sardinia and Sicily
Table 4. Distribution of the endemic species which are not
exclusive to the Sardinian-Corsican area.
Italian Region
Families (n.)
Species (n.)
Veneto
44
738
Lombardy
Alto Adige
Piedmont
Valle d’Aosta
Sardinia
39
35
43
29
43
679
606
546
261
495
Table 5. Number of species and families known in some
Italian regions.
93
Figure 15. Distribution of Sardinian and Sardo-Corsican endemism of the family Nemesiidae (top left) and Dysderiidae
(top right, down left, down right).
94
Figure 16. Distribution of some Sardinian and Sardo-Corsican endemism of the family Leptonetidae (top left), Pholcidae
(top right), Liocranidae (down left) and zodaridae (down right) (Zodarion sardum is not reported because is generic cited
for Sardinia and no indicates localities).
95
Figure 17. Distribution Sardinian and Sardo-Corsican endemism of the family Agelenidae (top left) and Linyphiidae (top
right, down left, down right).
96
Figure 18. Distribution of some Sardinian and Sardo-Corsican endemism of the family Theridiidae (left) and Philodromidae plus Thomisidae (right) (Xysticus parallelus is not reported because is generic cited for Sardinia and no indicates
localities).
appears to be the region with the fewest species. In
our opinion, this situation is not due to the paleobio-geographical events and/or environmental
characteristics of the island but to a lack of specific research, which is demonstrated by the absence
of several species with a wide distribution. On this
topic, it is interesting to make a comparison with
the fauna of nearby Corsica (Table 6). In fact, the
number of species in the Sardinian-Corsican area
totals 700 species and the number present on each
island is nearly the same, but actually the similarity between the two faunas is not very high (Sorensen’s index of similarity = 0.63).
On the whole, what strikes us most is the
lack of research into the araneocenosis of the
main Sardinian environments carried out with
adequate timings and methods. Most of the data
comes from general faunal or taxonomic publication, many of which are now outdated. To
prove this, there are 94 species, slightly less
than 20% of the total, whose latest citation
comes from Garneri (1902) or previously, a
good 110 or more years ago. We hope therefore
that further research into Sardinian araneofauna
will allow us to update and complete this catalogue.
97
Family
Sardi- Corsinian
can Shared
species species species Total
(n.)
(n.)
(n.)
Family
Sardi- Corsinian
can Shared
species species species Total
(n.)
(n.)
(n.)
Ctenizidae
3
1
1
3
Oxyopidae
3
2
2
3
Nemesiidae
4
7
3
8
zoropsidae
2
1
1
2
Filistatidae
2
2
2
2
zoridae
2
4
1
5
Sicaridae
1
1
1
1
Agelenidae
18
16
10
24
Scytodidae
2
2
2
2
Hahniidae
1
1
0
2
Leptonetidae
5
1
0
6
Dictynidae
8
5
3
10
Pholcidae
7
6
4
9
Amaurobiidae
3
4
2
5
Segestriidae
4
4
2
6
Titanoecidae
1
2
1
2
Dysderidae
12
12
2
22
Miturgidae
7
8
6
9
Oonopidae
5
5
2
8
Anyphaenidae
2
2
2
2
Palpimanidae
1
0
0
1
Liocranidae
8
8
6
10
Mimetidae
3
5
3
5
Clubionidae
9
9
5
13
Eresidae
2
1
1
2
Corinnidae
4
5
4
5
Oecobiidae
3
2
2
3
zodariidae
7
8
4
11
Uloboridae
4
4
4
4
Prodidomidae
1
0
0
1
Nesticidae
1
1
1
1
Gnaphosidae
43
48
32
59
Theridiidae
54
59
41
72
Sparassidae
3
3
3
3
Theridiosomatidae
0
1
0
1
Philodromidae
20
20
15
25
Mysmenidae
1
1
0
2
Thomisidae
36
34
21
49
Linyphiidae
58
81
31
108
Salticidae
60
71
44
87
Tetragnathidae
11
11
9
13
TOTAL
495
522
319
698
Araneidae
36
31
24
43
Lycosidae
35
31
21
45
Pisauridae
3
2
1
4
Table 6. Comparison between Sardinian and Corsican araneofauna. The data for Corsican fauna comes from “Fauna
Europaea” (Helsdingen, 2011).
98
ACKNOWLEDGEMENTS
The authors wish to thank Roberto A. Pantaleoni
for pressing and encouraging us even during the
most critical drafting phases of the catalogue, for
his patience and for allowing us to examine the vast
amount of material of the ISE Institute.
We would also like to thank Fulvio Gasparo and
Arthur Decae for their valuable advice. Thanks also
go to Aldo Mulas, Gianpaolo Ruzzante and Paolo
Fontana for their collected material. Heartfelt
thanks to Elena Pelizzoli for her illustrations and
Emanuele Biggi for the cover photos.
Our thanks also go to Marcello Verdinelli,
Maria Leonarda Fadda, Laura Loru, Alessandro
Molinu, Francesca Fiori, Piera Marras and Tiziana
Nuvoli who, together with the staff of the ISE and
the Department of Plant Protection of the University of Sassari contributed greatly to the collection
of new material.
REFERENCES
Alicata P., 1966. Le Harpactea (Araneae, Dysderidae)
della fauna italiana e considerazioni sulla loro origine. Atti dell'Accademia Gioenia di Scienze Naturali, 18: 190-221.
Alderweireldt M. & Jocqué R., 1992. A review of the nebulosa-group of Pardosa Koch 1847 in Africa, a
complex with some highly variable species (Araneae
Lycosidae). Tropical zoology, 5: 73-113.
Altara E., 1968. Ricerche biologiche in Sardegna. Sottoterra, Rivista del Gruppo speleologico bolognese CAI
e Speleo Club Bologna Esagono ENAL, 7: 23-27.
Argiolas M., Bartolo G. & Puddu S., 1970. Il Monte del
Castello di Quirra e le sue grotte. Ed. Tipografia
I.B.S., Cagliari, 56 pp.
Arnedo M.A., Gasparo F. & Opatova V., 2009. Systematics and phylogeography of the Dysdera erythrina
species complex (Araneae, Dysderidae) in Sardinia.
zooKeys, 16: 319-345.
Arnò C., Luciano P. & Pantaleoni R.A., 1998. La tassocenosi a ragni sui tigli di un'alberata cittadina a Sassari. In: Bologna M.A., Carpaneto G.M. & Cignini
B. (eds.) 1998, Atti 1° Convegno Nazionale sulla
Fauna Urbana. Roma, 12 aprile 1997. Fratelli Palombi editori, Roma, 87-89.
Ausserer A., 1871. Beiträge zur Kenntniss der Arachniden-Familie der Territelariae Thorell (Mygalidae
Autor). Verhandlungen zoologisch-botanische Gesellschaft in Wien, 21: 117-224.
Ballarin F., Pantini P. & Hansen H., 2011. Catalogo ragionato dei ragni (Arachnida, Araneae) del Veneto.
Memorie del Museo Civico di Storia Naturale di Verona - 2. Serie. Sezione Scienze della Vita 21, 151 pp.
Bonnet P., 1945. Bibliographia araneorum; analyse methodique de toute la litterature araneologique
jusqu‘en 1939. Douladore, Tolouse, 1: 832 pp.
Bartolo G., Bruscu E. & Dore M., 1973. Sa Fossa de Perd’e Cerbu. A cura dello Speleo Club di Cagliari. Ed.
Fossataro, Cagliari, 20 pp.
Bartolo G., Dore M. & Trincas G., 1972. Sa Fossa de su
Fenu Trainu (Terza voragine della Sardegna). A cura
dello Speleo Club di Cagliari. Ed. Fossataro, Cagliari,
39 pp.
Baum S., 1974. zum "Cribellaten-Problem": Die Genitalstrukturen der Oecobiinae und Urocteinae
(Arach.: Aran.: Oecobiidae). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg, 16
(N.F.): 101-153.
Bolzern A., Hänggi A. & Burckhardt D., 2008. Funnel
web spiders from Sardinia: Taxonomical notes on
some Tegenaria and Malthonica spp. (Araneae: Agelenidae). Revue suisse de zoologie, 115: 759-778.
Bosmans R., 1997. Revision of the genus Zodarion
Walckenaer, 1833, part II. Western and Central Europe, including Italy (Araneae: zodariidae). Bulletin of the British Arachnological Society, 10:
265-294.
Bosmans R., 2008. A new contribution to the knowledge
of the genus Typhochrestus Simon in Europe and
North Africa (Araneae: Linyphiidae). Revista Iberica
de Arachnologia, 16: 83-92.
Bosmans R. & Van Keer J., 1999. The genus Enoplognatha Pavesi, 1880 in the mediterranean region (Araneae: Theridiidae). Bulletin of the British
Arachnological Society, 11: 209-241.
Brignoli P.M., 1967a. Oonopidae di Sardegna e Puglie
(Araneae). Rendiconti dell'Istituto lombardo di
Scienze e Lettere (B), 101: 360-368.
Brignoli P.M., 1967b. Su alcuni Leptonetidae della Sardegna (Araneae). Rendiconti dell'Istituto lombardo
di Scienze e Lettere (B), 101: 352-359.
Brignoli P.M., 1969a. Note sugli Scytodidae d'Italia e
Malta (Araneae). Fragmenta Entomologica, 6: 121166.
Brignoli P.M., 1969b. Osservazioni su alcuni casi di rigenerazione delle zampe in Loxosceles rufescens
(Doufur) (Araneae, Scytodidae). Fragmenta Entomologica, 6: 115-119.
Brignoli P.M., 1969c. Secondo contributo alla conoscenza dei Leptonetidae della Sardegna (Araneae).
Archivio zoologico italiano, 54: 11-31.
Brignoli P.M., 1971a. Contributo alla conoscenza degli
Agelenidae italiani (Araneae). Fragmenta entomologica, 8: 57-142.
Brignoli P.M., 1971b. Note sui Pholcidae d'Italia. Fragmenta Entomologica, 7: 79-101.
Brignoli P.M., 1971c. Note sui ragni cavernicoli italiani.
Fragmenta Entomologica, 7: 121-229.
Brignoli P.M., 1973. Ragni d'Italia XX. Note sugli Hahniidae (Araneae). Fragmenta Entomologica, 8: 265274.
Brignoli P.M., 1974. Ragni d'Italia XXI. Settimo contributo alla conoscenza dei ragni cavernicoli di Sardegna e descrizione di una nuova specie di Corsica
(Araneae). Revue suisse de zoologie, 81: 387-395.
Brignoli P.M., 1975. Ragni d'Italia XXIII. Nuovi dati su
alcuni Haplogynae. Bollettino della Società Entomologica Italiana, 107: 170-178.
Brignoli P.M., 1976a. Beiträge zur Kenntnis der Scytodidae (Araneae). Revue suisse de zoologie, 83: 125191.
Brignoli P.M., 1976b. Ragni d'Italia XXIV. Note sulla
morfologia dei genitali interni dei Segestridae e cenni
sulle specie italiane (Araneae). Fragmenta Entomologica, 12: 19-62.
Brignoli P.M., 1977. Ragni d'Italia XXVII. Nuovi dati su
Agelenidae, Argyronetidae, Hahniidae, Oxyopidae e
Pisauridae, cavernicoli ed epigei (Araneae). Quaderni
del Museo di Speleologia "V. Rivera" L'Aquila, 2: 3118.
Brignoli P.M., 1979a. Contribution à la connaissance des
Uloboridae paléarctiques (Araneae). Revue Arachnologique, 2: 275-282.
Brignoli P.M., 1979b. Ragni d'Italia XXIX. Dysderidae
nuovi o interessanti. Bollettino della Società Entomologica Italiana, 111: 17-26.
Brignoli P.M., 1979c. Ragni d'Italia XXX. Nuovi dati
corologici ed ecologici su alcune specie di araneidae.
(Araneae). Fragmenta Entomologica, 15: 17-41.
Brignoli P.M., 1979d. Ragni d'Italia XXXI. Specie cavernicole nuove o interessanti (Araneae). Quaderni
del Museo di Speleologia "V. Rivera" L'Aquila, 10:
3-48.
Brignoli P.M., 1980. Ragni d'Italia XXXIII. Il genere Robertus (Araneae, Theridiidae). Fragmenta Entomologica, 15: 259-265.
Brignoli P.M., 1982. Contribution à la connaissance des
Filistatidae paléartiques (Araneae). Revue Arachnologique, 4: 65-75.
Brignoli P.M., 1984. zur Problematik der mediterranen
Pisaura - Arten (Arachnida, Araneae, Pisauridae).
zoologischer Anzeiger, Jena, 213 (1/2): 33-43.
Brignoli P.M. & Gaddini A., 1979. Nuovi dati su alcuni
Anyphenidae, Liocranidae e Gnaphosidae italiani
(Araneae). Bollettino dell'Associazione Romana di
Entomologia, 34: 10-15.
Brignoli P.M. & Murphy J., 1984. Nuovi dati su alcuni
Gnaphosidae italiani (Araneae). Bollettino dell'Associazione Romana di Entomologia, 38 (1983): 1-4.
Buchar J., Knoflach B. & Thaler K., 2006. On the identity of Arctosa variana C.L. Koch and Arctosa similis
99
Schenkel, with notes on related species (Araneae: Lycosidae). Bulletin of the British Arachnological Society, 13: 329-336.
Buchar J. & Thaler K., 1995. zur Variation der Kopulationsorgane von Pistius truncatus (Pallas) (Araneida,
Thomisidae) in Mitteleuropa. Linzer biologische Beiträge, 27: 653-663.
Burger M., Graber W., Michalik P. & Kropf C., 2006. Silhouettella loricatula (Arachnida, Araneae, Oonopidae): A Haplogyne Spider With Complex Female
Genitalia. Journal of Morphology, 267: 663-677.
Canestrini G. & Pavesi P., 1868. Araneidi italiani. Atti
della Società Italiana di Scienze Naturali, 11: 738-872.
Canestrini G. & Pavesi P., 1870. Catalogo sistematico
degli Araneidi italiani. Archivio per la zoologia,
l'Anatomia e la Fisiologia, 2: 1-44.
Canestrini G., 1873. Nuove specie italiane di Aracnidi.
Atti della Società Veneto-Trentina di Scienze Naturali, 2: 45-52.
Canestrini G., 1876. Osservazione aracnologiche. Atti
della Società Veneto-Trentina di Scienze Naturali, 3:
206-232.
Cannas V.M., 1982. Visioni di un paesaggio carsico. Sadali. Storia e aspetti naturali del suo territorio. Ettore
Gasperini ed., Cagliari, 46-49.
Casale A., 1972. Visione d'insieme del complesso ecologico e faunistico della Grotta del Bue Marino (Cala
Gonone, Dorgali, NU). Bollettino della Società Sarda
di Scienze Naturali, 10: 111-136.
Casale A., Grafitti G. & Onnis C., 2004. Sardaphaenops adelphus, il nuovo coleottero ipogeo dedicato
a tutti gli speleologi sardi. Anthèo, Bolletino
Gruppo speleo-archeologico Giovanni Spano, Cagliari, 8: 30-36.
Cerruti M., 1968. Materiali per un primo elenco degli Artropodi speleobii della Sardegna. Fragmenta Entomologica, 5: 207-257.
Colombo M. & Manunza B., 2009. First record of Cyrtarachne ixoides (Simon, 1870) (Araneae: Araneidae)
from Sardinia. Revista Ibérica de Aracnología, 17:
67-70.
Costa A., 1882. Notizie ed osservazioni sulla geofauna
sarda - Memoria Prima. Atti della Reale Accademia
delle Scienze Fisiche e Matematiche di Napoli, 9:
1-42.
sarda - Memoria Seconda. Atti della Reale Accademia delle Scienze Fisiche e Matematiche di Napoli,
1 serie 2: 1-109.
sarda - Memoria Terza. Atti della Reale Accademia
delle Scienze Fisiche e Matematiche di Napoli, 1
serie 2: 1-64.
sarda - Memoria Quarta. Atti della Reale Accademia
100
serie 2: 1-31.
sarda - Memoria Quinta. Atti della Reale Accademia
serie 2: 1-24.
de Carlini A., 1885. Artropodi dell'isola di S. Pietro. Bollettino della Società Entomologica Italiana, 17: 192196.
Decae A.E. & Di Franco F., 2005. Nemesia sanzoi Fage,
1917 (Araneae, Nemesiidae): first description of the
male and some additional information on the female, distribution and biology of the species. Bulletin of the British Arachnological Society, 13:
133-137.
Deeleman-Reinhold C.L. & Deeleman P.R., 1988. Revision
des Dysderinae (Araneae, Dysderidae), les especes mediterraneennes occidentales exceptees. Tijdschrift voor
Entomologie, 131: 141-269.
Di Pompeo P., Kulczycki A., Legittimo C.M. & Simeon
E., 2011. New records for Europe: Argiope trifasciata
(Forsskal, 1775) from Italy and Malta (Araneae, Araneidae). Bulletin of the British Arachnological Society, 15: 205-208.
Dresco E., 1956. Tegenaria henroti, espece nouvelle des
grottes de Sardaigne. Fragmenta Entomologica, 6:
167-181.
Dresco E. & Hubert M., 1969. Araneae Speluncarum Italiae I. Fragmenta Entomologica, 2: 115-119.
Fontana R., Cantarella T., Sala L. & Tongiorgi P., 1996.
A colletion of Italian salticids (Araneae), mainly from
the Tuscan-Emilian Apennines. Revue suisse de zoologie, hors série: 211-222.
Fresi G., Arnò C. & Pantaleoni R. A., 2002. I Ragni negli
ecosistemi: prime ricerche in Sardegna. Notiziario
Forestale, Cagliari, 20: 30-33.
Garneri G.A., 1902. Contribuzione alla fauna sarda.
Aracnidi. Bollettino della Società zoologica italiana,
serie 2, 3: 57-103.
Gasparo F., 1999. Ridescrizione di Stalita patrizii Rower,
1956, specie tipo del nuovo genere Sardostalita (Ara
Dysderidae). Bollettino del Museo regionale di
Scienze naturali di Torino, 16: 59-76.
Gozo A., 1908. Gli Aracnidi di caverne italiane. Bollettino
della Società Entomologica Italiana, 38: 109-139.
Grafitti G., 1979. Primi dati sulla fauna di Sa Ucca de Su
Tintirriolu. Bollettino del Gruppo speleologico Sassarese, 5: 32-37.
Grafitti G., 1984. Ricerche faunistiche nelle grotte di
Monte Majore. Bollettino del Gruppo speleologico
Sassarese, 8: 36-44.
Grafitti G., 1993. La fauna della Grotta Sa Rocca Ulari
(Borutta, Prov. di Sassari). Bollettino del Gruppo speleologico Sassarese, 14: 17-27.
Grafitti G., 2001. Note speleofaunistiche. In: De Waele
J., Onnis C. & Robin y. 2001. Lovettecannas, dove
le dolomie incontrano i graniti. Speleologia, Rivista
della Società speleologica Italiana, 45: 26-29.
Grafitti G., 2003. Ricerche faunistiche nella Grotta Is
zuddas (Santadi-Sardegna sud-occidentale). Sardegna Speleologica, 20: 64-68.
Grafitti G., 2007a. La fauna cavernicola. In: Mucedda
M. & Grafitti G. (eds.) 2007. Le grotte di Bonu
Ighinu nel Comune di Mara. Ed. Segnavia, Sassari,
65-75.
Grafitti G., 2007b. La fauna delle grotte. In: Piras V. (ed.)
2007. Gruttas e Pelciones. Grotte e caverne di terra e
di mare della Planargia e del Montiferru occidentale.
Geologia, geomorfologia, biologia. Ed. Grafiche Editoriali Solinas, Nuoro, 219-227.
Grafitti G. & Merella G., 2008. Ricerche faunistiche in
grotte del Sulcis - Iglesiente. Atti del XX Congresso
Nazionale di Speleologia, Iglesias 27-30 aprile 2007.
Memorie dell’Istituto Italiano di Speleologia, serie
II, 21: 83-95.
Grasshoff M., 1959. Dysdera-Arten von Inseln des Mittelmeergebietes (Arach., Araneae). Senckenbergiana
biologica, 40: 209-220.
Grasshoff M., 1983. Larinioides Caporiacco 1934, der
korrekte Name für die sogennante Araneus cornutus-Gruppe. Senckenbergiana biologica, 64: 225229.
Grill A., Knoflach B., Cleary D.F.R. & Kati V., 2005.
Butterfly, spider, and plant communities in different
land-use types in Sardinia, Italy. Biodiversity and
Conservation, 14: 1281-1300.
Grimm U., 1985. Die Gnaphosidae Mitteleuropas (Arachnida, Araneae). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg, 26 (NF): 5-318.
Grimm U., 1986. Die Clubionidae Mitteleuropas: Corinninae und Liocraninae (Arachnida, Araneae). Abhandlungen des Naturwissenschaftlichen Vereins in
Hamburg, 27 (NF): 3-91.
Hansen H., 1985. Contributo alla conoscenza dei Salticidae italiani (Arachnida: Araneae). Bollettino del
Museo civico di Storia naturale di Venezia, 34(1983):
241-322.
Hansen H., 1986. Die salticidae der coll. Canestrini (Arachnidae: Araneae). Lavori - Società Veneziana di
Scienze Naturali, 11: 97-120.
Hansen H., 2005. Arachnida Araneae Salticidae + CDROM. In: Ruffo S. & Stoch F. (eds) 2005, Checklist
e distribuzione della fauna italiana. Memorie del
Museo civico di Storia naturale di Verona, 2 Serie,
Sezione Scienze della Vita, 16: 77-78.
Helsdingen P.J. van, 1969. A reclassification of the species of Linyphia Latreille based on the functioning of
the genitalia (Araneida, Linyphiidae), I. zoologische
Verhandelingen, 105: 3-303.
Helsdingen P.J. van, 2005. New spider species records
for Sardinia (Arachnida: Araneae). Nieuwsbrief SPINED, 20: 13-20.
Helsdingen P.J. van, 2012. Araneae. In: Fauna Europaea,
Database European spiders and their distribution - Distribution - Version 2012.1, http://www.faunaeur.org.
Isaia M., 2005. Check-list delle specie di ragni (Arachnida, Araneae) della Val d’Aosta con una nuova segnalazione per la fauna Italiana. Revue Valdotaine
d’Histoire Naturelle, 59:25-43.
Isaia M., Pantini P., Beikes S. & Badino G., 2007. Catalogo ragionato dei ragni (Arachnida, Araneae) del
Piemonte e della Lombardia. Memorie dell’Associazione naturalistica piemontese, 9, 161 pp.
Knoflach B., 1996. Die Arten der Steatoda phalerataGruppe in Europa (Arachnida: Araneae, Theridiidae).
Mitteilungen der Schweizerischen Entomologischen
Gesellschaft - Bulletin de la Société Entomologique
Suisse, 69: 377-404.
Knoflach B., 1999. The comb-footed spider genera Neottiura and Coleosoma in Europe (Araneae, Theridiidae).
Mitteilungen der Schweizerischen entomologischen
Gesellschaft - Bulletin de la Société Entomologique
Suisse, 72: 341-371.
Knoflach B., 2004. Diversity in the copulatory behaviour
of comb-footed spiders (Araneae, Theridiidae). Denisia 12, zugleich Kataloge der OÖ. Landesmuseen,
Neue Serie 14: 161-256.
Knoflach B. & Pfaller K., 2004. Kugelspinnen - eine Einführung (Araneae, Theridiidae). Denisia 12, zugleich
Kataloge der OÖ. Landesmuseen, Neue Serie 14:
111-160.
Knoflach B., Rollard C. & Thaler K., 2009. Notes on Mediterranean Theridiidae (Araneae) - II. zooKeys, 16:
227-264.
Knoflach B. & Thaler K., 2000. Notes on the mediterranean Theridiidae (Araneae). I. Memorie della Società
Entomologica Italiana, 78: 411-442.
Krauss O., 1955. Spinnen von Korsika, Sardinien und
Elba (Arach., Araneae). Senckenbergiana biologica,
36 (5/6): 371-394.
Kulczycki A., Legittimo C.M., Simeon E. & Di Pompeo P., 2012. New records of Steatoda nobilis (Thorell, 1875) (Araneae, Theridiidae), an introduced
species on the Italian mainland and in Sardinia.
Bulletin of the British Arachnological Society, 15:
269-272.
Lana E., Casale A., Giachino P.M. & Grafitti G., 2006.
Attività biospeleologica anni 2004-2005. Grotte, Bollettino del Gruppo speleologico piemontese CAIUGET, Torino, (49) 145: 38-47.
Letardi A. & Pantaleoni R.A., 2007. Survey towards an
inventory of Italian Neuropterida. In: Pantaleoni
R.A., Letardi A. & Corazza C. (ed). Proceedings of
the IX International Symposium on Neuropterology,
Ferrara 20-23 June 2005 [with a Tribute to Maria Matilde Principi]. Annali del Museo civico di Storia naturale di Ferrara, 8: 87-93.
101
Loerbroks A., 1983. Revision der Krabbenspinnen-Gattung Heriaeus Simon (Arachnida: Araneae: Thomisidae). Verhandlungen des Naturwissenschaftlichen
Vereins in Hamburg, 26 (NF): 85-139.
Logunov D., 2004. Notes on the collection of Salticidae
(Araneae) from the Museum of Natural History "Enrico Gaffi" in Bergamo, Italy. Revista Ibérica de
Aracnologia, 9: 271-275.
Logunov D. & Azarkina G.N., 2006. New species and
records of Phlegra from Africa (Araneae, Salticidae).
Revue suisse de zoologie, 113: 727-746.
Loru L., Marras P.M., Guidone L., Sassu A., Verdinelli
M., Tavella L. & Pantaleoni R.A., 2008. Importanza
dei predatori generalisti in noccioleti piemontesi e
sardi. In: Scortichini M. (ed.) 2008. La corilicoltura
biologica in Italia. Ed. CRA Centro di Ricerca per la
Frutticoltura, Roma, 111-136.
Lugetti G. & Tongiorgi P., 1965. Revisione delle specie
italiane dei generi Arctosa C.L. Koch e Tricca
Simon con note su una Acantholycosa delle Alpi
Giulie (Araneae, Lycosidae). Redia, 49 (19641965): 165-229.
Lugetti G. & Tongiorgi P., 1969. Ricerche sul genere Alopecosa Simon (Araneae, Lycosidae). Atti della Società Toscana di Scienze Naturali Memorie, serie B
76: 1-100.
Magretti P., 1878. Rapporto su di un'escursione nella Sardegna compiuta nel dicembre 1877. Atti della Società
Italiana di Scienze Naturali, 21: 451-463.
Magretti P., 1880. Una seconda escursione zoologica all'isola di Sardegna. Atti della Società Italiana di
Marras P.M., Loru L., Sassu A., Sanna F. & Pantaleoni
R.A., 2011. Influence of ant-exclusion on generalist
predators in a citrus orchard. IOBC/wprs Bulletin Vol
62, 2011: 345-350.
Metzner H., 1999. Die Springspinnen (Araneae, Salticidae) Griechenlands. Andrias, 14: 3-279.
Minelli A., Ruffo S. & La Posta S. (eds.), 1993-1995.
Checklist delle specie della fauna italiana. Edizioni
Calderini, Bologna.
Muccedda M., Grafitti G. & Dore G., 2002. Grotte della
provincia di Sassari. La Celere Editrice, Alghero:
208 pp.
Muster C., 2009. Phylogenetic relationships within Philodromidae, with a taxonomic revision of Philodromus subgenus Artanes in the western Palearctic
(Arachnida: Araneae). Invertebrate Systematics, 23:
135-169.
Muster C., Bosmans R. & Thaler K., 2007. The Philodromus pulchellus-group in the Mediterranean: taxonomic revision, phylogenetic analysis and
biogeography (Araneae: Philodromidae). Invertebrate Systematics, 21: 39-72.
Muster C. & Thaler K., 2004. New species and records
of Mediterranean Philodromidae (Arachnida, Ara-
102
neae): I. Philodromus aureolus group. Denisia, 14:
305-326.
Nentwig W., Blick T., Gloor D., Hanggi A. & Kropf C.,
2011. Araneae Spiders of Europe, Version 6.2011.
Web site at http://www.araneae.unibe.ch/
Noflatscher M.T., 1996. Aranei - Spinnen, Webspinnen.
In: Hellrigl K. (ed.) 1996, Die Tierwelt Südtirols. Veröffentlichungen des Naturmuseums Südtirol, Bozen,
211-228.
Pantini P., in press. Italian araneofauna: the state of our
knowledge. XXXVIII Congresso della Società Italiana di Biogeografia. Biogeographia vol. XXXI.
Pantini P. & Isaia M., 2008. New records for the Italian
spider fauna (Arachnida, Araneae). Arthropoda Selecta, 17: 133-144.
Pantini P. & Sassu A., 2009. I ragni dell’Isola dell’Asinara (Sardegna NW) (Arachnida, Araneae). Annali
del Museo civico di storia naturale di Genova, 100:
619-647.
Pavesi P., 1876. Le prime crociere del Violante, comandato dal Capitano-Armatore Enrico d'Albertis. Risultati aracnologici. Annali del Museo civico di storia
naturale di Genova, 8: 407-451.
Pavesi P., 1878. Nuovi risultati aracnologici delle Crociere del "Violante". Aggiunto un catalogo sistematico degli Aracnidi di Grecia. Annali del Museo
civico di storia naturale di Genova, 11: 337-396.
Pesarini C., 1994. Arachnida Araneae. In: Minelli A.,
Ruffo S. & La Posta S. (eds.) 1994. Checklist delle
specie della fauna italiana. Edizioni Calderini, Bologna, 23, 1-42.
Pesarini C., 2000. Contributo alla conoscenza della fauna
araneologica italiana (Araneae). Memorie della Società Entomologica Italiana, 78: 379-393.
Platnick N.I., 2012. The world spider catalog, version
12.5. American Museum of Natural History, online
at http://research.amnh.org/iz/spiders/catalog. DOI:
10.5531/db.iz.0001.
Platnick N.I. & Di Franco F., 1992. On the relationships
of the spider genus Cybaeodes (Araneae, Dionycha).
American Museum Novitates, 3053: 1-9.
Pons G., 1993. Noves dades sobre biospeleologia de Sardenya: (Arachnida, Araneae). Endins (Palma de Mallorca), 19: 43-47.
Puddu S., 1970. La fauna. In: Bartolo G. & Di Paola A.
1970. Ussassai. Gairo. Osini. Attuali conoscenze speleologiche. A cura dello Speleo Club di Cagliari. Ed.
Sarda Fossataro, Cagliari, 79-93.
Puddu S., 1971. La fauna. In: Bartolo G. & Ferrara R.
1971. Il Promontorio di S. Elia e le sue grotte. A cura
dello Speleo Club di Cagliari. Ed. Sarda Fossataro,
Cagliari, 67-68.
Puddu S., 1980. La fauna. In: Bartolo G., Dore M. &
Lecis A. 1980. Is Angurtidorgius (La cavità più vasta
della Sardegna). A cura dello Speleo Club Cagliari.
Ed. GIA, Cagliari, 41-56.
Puddu S., 1983. Attuali conoscenze speleofaunistiche. In:
Bartolo G., Coroneo M. & Lecis A. (eds.) 1983. Su
Marmori (Perla di Ulassai). A cura dello Speleo Club
Cagliari. Ed. Guido Bartolo, Cagliari, 33-40.
Puddu S. & Pirodda G., 1974. Catalogo sistematico ragionato della fauna cavernicola della Sardegna. Rendiconti del Seminario della Facoltà di Scienze
dell’Università di Cagliari, 43 (3-4), (1973): 151-205.
Raven R.J., 1985. The spider infraorder Mygalomorphae (Araneae): Cladistics and systematics. Bulletin
of the American Museum of Natural History, 182:
1-180.
Roewer C.F., 1953. Cavernicole Arachniden aus Sardinien. Notes biospéléologiques, 8: 39-49.
Roewer C.F., 1956. Cavernicole Arachniden aus Sardinien. II. Fragmenta Entomologica, 2 (9): 97-104.
Ruffo S. & Stoch F., 2005. Checklist e distribuzione della
fauna italiana. Memorie del Museo Civico di Storia
Naturale di Verona, 2a serie, Sezione Scienze della
Vita, 16, 307 pp.
Ruffo S. & Vigna Taglianti A., 2002. Generalità sulla
fauna italiana. In: Minelli A., Chemini C., Argano R.
& Ruffo S. (eds) 2002, La fauna in Italia, Touring
Editore, Milano e Ministero dell’Ambiente e della
Tutela del Territorio, Roma, 24-28.
Sanna B., Puddu S. & Bartolo G., 1972. Su Mannau. Ed.
Fossataro, Cagliari, 97-114.
Senglet A., 2004. Copulatory mechanism in Zelotes,
Drassyllus and Trachyzelotes (Araneae, Gnaphosidae), with additional faunistic and taxonomic data on
Southwest Europe. Mitteilungen der Schweizerischen
entomologischen Gesellschaft - Bulletin de la Société
Entomologique Suisse, 77: 87-119.
Senglet A., 2011. New species in the Zelotes tenuis-group
and new or little known species in other Zelotes
groups (Gnaphosidae, Araneae). Revue suisse de
zoologie, 118: 513-559.
Simon E., 1884. Les arachnides de France. La famille des
Theridionidae (suite). Paris, Enciclopedie Roret, L.
Mulo, tomo 5 (2 parte), 180-420.
Simon E., 1914. Les arachnides de France. Synopsis générale et catalogue des espèces françaises de l'ordre
des Araneae. Paris, Enciclopedie Roret, L. Mulo,
tomo 6 (1 parte), 1-308.
des Araneae (suite). Paris, Enciclopedie Roret, L.
Tanasevitch A.V., 2011. On linyphiid spiders (Araneae)
from the Eastern and Central Mediterranean kept at
the Muséum d’histoire naturelle, Geneva. Revue
suisse de zoologie, Genève, 118: 49-91.
Thaler K. & Knoflach B., 1998. Zoropsis spinimana (Dufour), eine für Österreich neue Adventivart (Araneae,
zoropsidae). Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, 85: 173-185.
Thaler K. & Knoflach B., 2003. zur Faunistik der Spinnen (Araneae) von Österreich: Orbiculariae p.p.
(Araneidae, Tetragnathidae, Theridiosomatidae,
Uloboridae). Linzer biologische Beiträge, 35: 613655.
Thaler K. & Knoflach B., 2004. Fauna Austriaca: Webspinnen - zur Einführung (Arachnida, Araneae). Denisia 12, zugleich Kataloge der OÖ. Landesmuseen,
Neue Serie 14: 357-380.
Thaler K., van Harten A. & Knoflach B., 2004. Pirate
spiders of the genus Ero C.L. Koch from southern
Europe, yemen, and Ivory Coast, with two new species (Arachnida, Araneae, Mimetidae). Denisia, 13:
359-398.
Thermes G., 1972. Primo contributo alla conoscenza
della fauna araneologica della Sardegna (Fauna ipogea ed epigea). Bollettino della Società Sarda di
Tongiorgi P., 1966. Italian Wolf Spiders of the Genus
Pardosa (Araneae, Lycosidae). Bulletin of the Museum of Comparative zoology, 134: 275-334.
Tongiorgi P., 1974. Some notes on Pardosa occidentalis
Simon (Araneae, Lycosidae). Bulletin of the British
Arachnological Society, 3: 62-63.
Tongiorgi P. & Sola C., 1991. Pardosa nigristernis Denis,
1966. A North African component of the Italian and
Spanish arachnofauna. XIIIe Colloque Européen
d’Arachnologie Neuchâtel 2-6 sept. 1991, Bulletin de
la Société neuchâteloise des Sciences Naturelles, 116:
241-244.
Trotta A., 2005. Introduzione ai Ragni italiani (Arachnida
Araneae). Memorie della Società Entomologica Italiana, 83: 3-178.
Trotta A., 2009. Ricerche zoologiche della nave oceanografica “Minerva” (C.N.R.) sulle isole circumsarde.
XXXII. Araneae. Annali del Museo Civico di Storia
Naturale "Giacomo Doria", 100: 481-500.
Trotta A., 2011. Second contribution to the knowledge
of the spiders of Sardinia (Arachnida, Araneae). In:
103
Nardi G., Whitmore D., Bardiani M., Birtele D.,
Mason F., Spada I. & Cerretti P. (eds) 2011. Biodiversity of Marganai and Montimannu (Sardinia). Research in the framework of the ICP Forest network.
Conservazione Habitat Invertebrati, 5: 137-161.
Vigna Taglianti A., Audisio P.A., Belfiore C., Biondi M.,
Bologna M.A., Carpaneto G.M., De Biase A., De Felici S., Piattella E., Racheli T., zapparoli M. & zoia
S., 1992. Riflessioni di gruppo sui corotipi fondamentali della fauna W-Paleartica ed in particolare italiana.
Biogeographia, 16: 159-179.
Vigna Taglianti A., Audisio P.A., Biondi M., Bologna
M.A., Carpaneto G.M., De Biase A., Fattorini S.,
Piattella E., Sindaco R.,Venchi A. & zapparoli M.,
1999. A proposal for a chorotype classification of the
Near East fauna in the framework of the Western Paleartic region. Biogeographia, 20: 31-59.
Wunderlich J., 1973. Beschreibung einiger bisher unbekannter Arten der Gattung Zodarion Walckenaer aus
Südeuropa (Arachnida: Araneae: zodariidae). Senckenbergiana biologica, 54: 171-176.
Wunderlich J., 1995a. Sardinion perplexum n. gen. n. sp.,
eine bisher unbekannte Spinnenart von Sardinien
(Arachnida: Araneae: Theridiidae). Beiträge zur Araneologie, 4 (1994): 687-690.
Wunderlich J., 1995b. zur kenntnis der Endemiten, zur
evolution und zur Biogegraphie der Spinnen Corsicas und Sardiniens, mit neubeschreibungen (Arachnida: Araneae). Beiträge zur Araneologie, 4
(1994): 353-383.
Wunderlich J., 1995c. zur kenntnis mediterraner Arten
der Gattung Enoplognatha Pavesi 1880, mit einer
neubeschreibung (Arachnida: Araneae: Theridiidae).
Beiträge zur Araneologie, 4 (1994): 703-713.
Wunderlich J., 1995d. zur kenntnis West-Palaarktischer
Arten der Gattungen Psammitis Menge 1875, Xysticus C.L. Koch 1835 und Ozyptila Simon 1864 (Arachnida: Araneae: Thomisidae). Beiträge zur
Araneologie, 4 (1994): 749-774.
Wunderlich J., 1995e. zur taxonomie und Biogeographie der Arten der Gattung Oecobius Lucas 1846,
mit neubeschreibungen aus der mediterraneis und
von der arabischen Halbinsel (Arachnida: Araneae:
Oecobiidae). Beiträge zur Araneologie, 4 (1994):
585-608.
Wunderlich J., 2011. Extant and fossil spiders (Araneae).
Beiträge zur Areneologie, 6: 1-640.
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INDEX
ABSTRACT
3
ARANEOLOGICAL RESEARCH
IN SARDINIA
4
INTRODUCTION
MATERIALS AND METHODS
LIST OF SPECIES
Family Ctenizidae
Family Nemesiidae
Family Filistatidae
Family Sicariidae
Family Scytodidae
Family Leptonetidae
Family Pholcidae
Family Segestriidae
Family Dysderidae
Family Oonopidae
Family Palpimanidae
Family Mimetidae
Family Eresidae
Family Oecobiidae
Family Uloboridae
Family Nesticidae
Family Theridiidae
Family Mysmenidae
Family Linyphiidae
3
8
11
11
11
12
12
13
13
14
15
15
18
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19
19
19
20
20
21
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Family Tetragnathidae
Family Araneidae
Family Lycosidae
Family Pisauridae
Family Oxyopidae
Family zoropsidae
Family zoridae
Family Agelenidae
Family Hahniidae
Family Dictynidae
Family Amaurobiidae
Family Titanoecidae
Family Miturgidae
Family Anyphaenidae
Family Liocranidae
Family Clubionidae
Family Corinnidae
Family zodariidae
Family Prodidomidae
Family Gnaphosidae
Family Sparassidae
Family Philodromidae
Family Thomisidae
Family Salticidae
CONSIDERATIONS ABOUT
SARDINIAN ARANEOFAUNA
CONCLUSIONS
ACKNOWLEDGEMENTS
REFERENCES
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Catalogue of the spiders (Arachnida Araneae

Transcription

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