Wildflowers of Mount Ashland - Southern Oregon Digital Archives
Transcription
Wildflowers of Mount Ashland - Southern Oregon Digital Archives
Wildflowers of Mount Ashland E. Noelle Jordan ACKNOWLEDGMENTS Many individuals and organizations are deserving of special thanks. The United States Forest Service, Medford District, namely Mr. Wayne Rolle, provided financial backing and botanical expertise. The University of Washington Press, Stanford University Press, and University of California Press provided the line drawings. Drawings are from University of Washington Press unless otherwise indicated. Bob Vos and Elizabeth Wasserman kindly donated the photographs that are credited to them. Some photographs were obtained from various internet sites and are credited to the appropriate individuals. All other photographs were taken by the author. Dr. Steve Jessup provided guidance, great reference material, and much needed assistance in identification of specimens. Dr. Frank Lang provided direction, focus, and his valuable time and patience in trying the key for the first time. Lori Hammer, Matt MacKinnon, John Taylor, Chris van Schaak, and Elizabeth Wasserman assisted with collecting and identification of specimens. Mr. Jim Duncan, and Mr. Wayne Rolle provided partial floral lists for Mt. Ashland. Mr. Steve Johnson of the U. S. Forest Service, Ashland District, provided statistics, various information, photographs, and maps. i TABLE OF CONTENTS Section Part 1: Background Information Part 2: The Key System and How It Works Part 3: Family Descriptions Flower Descriptions Appendix A: Glossary Appendix B: Illustrated Glossary Appendix C: Bibliography Index by Common Name Index by Scientific Name ii PART 1 Background Information "Drink in the beauty, and think and wonder at the meaning of what you see." Rachel Carson Sense of Wonder (1956) Background Information Introduction The term "wildflower" is vague, often defined differently by different people. What one person considers a weed, another considers a wildflower. What one considers a wildflower, another considers a shrub. In this field guide, the term "wildflower" is used to indicate herbaceous plants that have showy flowers, and all flowering shrubs. An herbaceous plant is simply a non-woody plant, and a shrub is any plant that is woody throughout, shorter than 1.5 m tall, or branched from the base. This guide does not include conifers, ferns, mosses, lichens, grasses, sedges, or rushes. This key and guide are designed for nature enthusiasts who have recently been introduced to the world of plants and flowers. It is my hope that the beginner botanist will find this guide simple and straight-forward to use. But I have also designed this guide in hopes that you will learn a few new words to improve your observational skills, not only of wildflowers, but of all things in the natural world. Most of all, I hope to share with you some of the beauty and wonder of the wildflowers on Mount Ashland. Because this field guide covers a fairly small area, I had the luxury of being extremely thorough. Within these pages you will find descriptions and anecdotes for over 200 species of wildflowers and flowering shrubs. Why Wildflowers? Wildflowers brighten all seasons. Spring and summer see the bright faces of the blossoms, the autumn air is full of the blowing seeds; and winter is speckled with vivid fruits and berries. Wildflowers have touched peoples' lives in many ways through the millennia. They are symbols of love, happiness, and all the joys of summertime. They act as timepieces - people have come to know that certain flowers appear at certain times of the year, or that flowers progress through their life cycles at certain times. And flowering plants have earned our respect as sources of food, medicine, dyes, oils, spices, fibers, perfumes, and tools. Indeed, without the flower, where would we be? It seems important that we spend some time becoming familiar with the very things that bring us life and happiness. Why Mount Ashland? Mount Ashland, the tallest peak of the Siskiyou Crest, is a special place providing sweeping vistas of fields abloom and of other peaks in the distance. The dry hillsides, wet meadows, and forested areas offer three types of habitats to explore, and new sets of flowers in each. The Pacific Crest Trail that courses along the south slope of Mount Ashland offers hours of hiking and naturalizing pleasure. Whether you plan to picnic, camp, hike, bike, drive, or simply meander along Mount Ashland, flowers will greet you everywhere. Ski Ashland brings hundreds of skiers to Mount Ashland every winter. I wonder if they are curious to see what lies blanketed beneath their snow. The Siskiyou Crest comprises the mountain peaks and ridges of the Siskiyou Mountains in the Rogue River and Klamath National Forests. Mount Ashland is joined by McDonald Peak, Siskiyou Peak, Red Mountain, Dutchman's Peak, Observation Peak, and several more. The Siskiyou Mountains are unique in that they are one of the few east-west mountain ranges in the 1 Background Information United States. The Siskiyou Mountains connect the Cascade Mountains to the Coastal Mountain Range, and to the greater Klamath Ranges. Many of the flowers on Mount Ashland can easily be viewed from the Pacific Crest Trail and from the forest service roads. There is no need to venture far in order to soak in the beauty and serenity of these places. If, however, you are feeling spritzy and wish to leave the trail or the roads, please walk lightly. These subalpine ecosystems are fragile, and could be in danger of being loved to death. Some of the flowers you will see on Mount Ashland can be seen nowhere else in the world! So, enjoy, revel, ponder, and play. But please ...take only pictures, and leave only footprints! How to Get There Take 1-5 to exit 6 to reach Mount Ashland. Go straight at the end of the ramp, and turn right onto Mount Ashland Road (County Road 1151). The road is paved up to the ski area, and just beyond the ski area parking lot, the road turns to gravel and becomes U.S. Forest Service (USFS) Road 20. At mile 10 the road forks - the right fork, Road 300, will take you to the summit, the left fork, USFS Road 20, will take you to Grouse Gap and beyond to Siskiyou Peak. If you have more time and are up an adventure, hike into the area on the Pacific Crest National Scenic Trail, more commonly called the Pacific Crest Trail, or PCT. You can find the trail at several different places. To hike the entire mountain, pick up the trail on the road close to the junction of 1-5. For a shorter hike (an easy 3.5 miles one way), take the trail from mile 7.1 along County Road 1151 (just beyond the signs for Bull Gap) to Grouse Gap Shelter. For a leisurely stroll, you can locate the trail at USFS Road 40S15 just below the picnic and camping area, and follow it to Grouse Gap Shelter (approximately 1.5 miles). The Pacific Crest National Scenic Trail snakes 2,650 miles from the U.S./Canada border to the U.S./Mexico border. The north section follows the Cascade Mountains through Washington and most of Oregon. In southern Oregon, it winds through the eastern Siskiyous and into the Klamath Mountains in California. The section of the trail through the Siskiyou Mountains is jointly managed by the Ashland Ranger District and the Klamath National Forest of northern California. The PCT is open to hikers, campers, and horseback riders; bikes and motor vehicles are prohibited. For maps and more information, you can contact the following organizations: Ashland Ranger District of the Rogue River National Forest, 541/482-3333, Oak Knoll Ranger District of the Klamath National Forest, 530/468-1238, Pacific Crest Trail Association. 916/349-2109, or at www.pcta.org. You may also want to pick up The Pacific Crest Trail Volume 2: Oregon and Washington (Schaffer and Selters 1990). 2 Background Information Area Covered By This Field Guide Welcome to the eastern portion of the Siskiyou Crest! With this field guide in hand, you will have the pleasure and excitement of discovering some of the most colorful beauties in southern Oregon. The flowers in this field guide can be found on Mount Ashland above 6,200 feet in elevation. However, these flowers are representative of those found all along the eastern section of the Siskiyou Crest wherever you find granite-diorite soils. Mount Ashland Area Covered by this Field Guide and Major Landmarks (Source: USGS Mt. Ashland Quadrangle 1983 edition, 7.5 minute series) Map Legend: Boundary of Survey Area, 1900 m (6200 ft) USFS Road 20 _ USFS Road 300 to summit — — Grouse Gap Shelter X Rabbit Ears Summit Q Picnic Area USFS Road 40S30, Grouse Gap Road Pacific Crest Trail (PCT) A Ski Ashland Lodge | The Knoll 3 Background Information Management of Mount Ashland Mount Ashland straddles two National Forests and therefore is under management of two districts (and two different regions) within the U.S. Forest Service (USFS). The north slope of Mount Ashland lies within the Rogue River National Forest and is an extremely important area in the Ashland Watershed. The north slope is managed by the USFS Ashland Ranger District, part of the Pacific Northwest region which includes Washington, Oregon and parts of California. The south slope lies within the Klamath National Forest and is managed by the USFS Oak Knoll Ranger District, part of the Pacific Southwest Region, which includes parts of California, parts of Arizona, and Nevada. Most of Mount Ashland has been designated as a Late Successional Reserve (LSR) in accordance with the Northwest Forest Plan. LSRs are designated areas that are managed to protect and enhance late successional and old growth forest ecosystems in order to preserve habitat for the Spotted Owl and other old-growth obligate species. Surprisingly, this does not mean that the area cannot be logged or managed in other ways. LSRs can be logged as long as the activity is certified to maintain or enhance the old-growth qualities. The land on which the ski area was built is owned by the City of Ashland and is leased to Ski Ashland who operates and manages the ski area in accordance with the rules and regulations set forth by the USFS. At the summit, there is a television transmitter for KTVL, and a National Oceanographic and Atmospheric Administration (NOAA) Weather Radio (162.475 MHz) that broadcasts continuous weather information through the National Weather Service Program Office in Medford, Oregon. Topography and Geology of Mount Ashland From a distance, the mountain peaks in southwest Oregon appear to be masses of bare rock, but upon closer inspection, we find an embarrassment of riches right at our back door. The steep, craggy slopes of Mount Ashland are cloaked in open dry meadows, wet meadows, coniferous forest, and rocky outcrops. Flowers will be found in all of these habitats. Many flowers specialize in one particular habitat, but some will be found in several - or all - of the habitats. The highest point in Oregon west of the Cascade Range, Mount Ashland (7,533 feet) is considered part of the Ashland batholith, an ancient granitic intrusion. The dominant rock types constructing this peak are quartz diorite and granite. Much of the rock is now very weathered and decomposed, resulting in loose, gravelly to cobbly sandy loam soils. The topsoil depth, which varies with aspect and slope, are generally between 5 and 10 inches, while the overall soil depth ranges from two to four feet. The soils are moderately well-drained to excessively drained. Nowhere is it documented that glaciers covered any area in the Siskiyou Mountains. However, Mount Ashland reveals evidence of ancient small montane glaciers. Glacial features on Mount Ashland are found primarily on the upper northern slopes where there is a steepwalled cirque and a lateral moraine. This area is commonly called "the bowl" by downhill and cross-country skiers. Two other small cirques, many small remnants of moraines, and two Ushaped valleys are found to the west and north of the bowl. The steep, fragile slopes of Mount Ashland are riddled with unstable terrain. If observant, you will notice areas scarred by severe erosion and landslides. Ridges are generally 4 Background Information more stable, and are characterized by gentle slopes, rock outcrops, and bouldery ground. Northfacing slopes are dominated by coniferous forests, while the summit, ridges, south and west sides of the mountain are mostly non-forested plant communities, alternating from meadow to shrubland. Mount Ashland produces the headwaters of Ashland Creek, Grouse Creek, and Cottonwood Creek. The East and West Forks of Ashland Creek, the primary source of drinking water for the nearby City of Ashland, originate on the north slopes of Mount Ashland. Ashland Creek runs through Lithia Park in the City of Ashland and feeds into Bear Creek, a tributary of the Rogue River. Grouse Creek originates on the south slope, and Cottonwood Creek flows down the southeast slope of Mount Ashland, both emptying into the Klamath River. For more information on the topography and geology of Mount Ashland, try the following resources: Final Environmental Impact Statement: Mount Ashland Ski Area (USFS 1991), and Roadside Geology of Oregon (Alt and Hyndman 1978). The Climate Of Mount Ashland The general climate in and around Ashland, Oregon is characterized by hot, dry summers and mild, wet winters. This type of regime is often referred to as a Mediterranean climate. The average annual precipitation in the City of Ashland is about 50 cm (20 inches), while the average annual precipitation on Mount Ashland from 1961-1990 was 150-200 cm (60-80 inches). Snow begins falling on Mount Ashland around mid to late November and accumulates through the end of April during a good snow year. Snow depth on the south slopes can reach 23210 cm (9-84 inches), depending on the amount of precipitation, with snow drifting sometimes much deeper. Snow depth on the peak can vary from 55-560 cm (22-220 inches), and at the ski lodge from 37-384 cm (15-151 inches). The average temperature during the winter months is 3.33° C (26°F). The summit and open ridges are exposed to severe winter winds. Summer temperatures on Mount Ashland reach 29-32°C (85-90°F). Summer precipitation is rare, and usually occurs as thunderstorms. For more information on the climate of Mount Ashland, check the website for the Western Regional Climate Center at www.wrcc.dri.edu, or call the U.S. Forest Service, Ashland Ranger District at 541/482-3333. Subalpine Plant Adaptations An adaptation is defined as a change in structure, function or form that improves the chance of survival for an organism within a given environment. Darwin used the term largely to refer to favorable characters which enable organisms to adjust to their surroundings, and therefore, persist, while organisms lacking such characters perished. It follows, then, that adaptations arise from a mutual relationship between an organism - plants, in this case - and its environment. Plants are special because they are completely at the mercy of the physical 5 Background Information environment. Although over time plants can migrate as a result of seeds blowing on the wind or hitching a ride on animal fur, an individual plant cannot pick up and move to a more favorable location. For many reasons, mountains probably more than any other place offer the greatest possibilities for wildflowers. Differences in elevation, abundant water, clear air, light, different soil types, and temperature mingle together to create a richness irresistible to flowers and those who love them. The flowers on Mount Ashland above 1900 m (6200 ft) are, strictly speaking, in the subalpine zone, that is at or below the tree line. On higher mountains, the alpine zone starts just above the tree line. Plants in the alpine and subalpine zones face special challenges compared to plants in lower elevations. They must grow, reproduce and set seed during a very short growing season. They must withstand cold temperatures, brisk winds and snow, and survive frost, intense sun, and occasional seasonal drought that is exacerbated by porous soils. We can say that flowers in the subalpine zone are both hearty and fragile. How do they survive? They have developed adaptations - such as matted growth habits, hairs, small waxy leaves, and red coloration - that allow these beautiful wonders not only to survive, but to thrive in their subalpine habitat. As you read through the descriptions for the plants in this book, you will notice that the majority on Mount Ashland are perennials (plants that require three or more years to complete their life cycle). The deep roots of perennials grasp firmly onto the loose soil or rocks and stretch to find water during the dry summers. Large perennial roots, rhizomes, or bulbs store large amounts of carbohydrates necessary to sustain the plant through long winters. Perennials often reproduce vegetatively because seed set requires a very specific set of circumstances. Annuals - plants that complete their life cycle in one year - are rare for several reasons. They generally cannot afford to bloom and go to seed in the short capricious season that the mountains call summer. The annuals that grow on Mount Ashland, including Torrey's Blue-Eyed Mary (Collinsia torreyi), Staining Collomia (Collomia tincturia), and Twinleaf Bedstraw (Galium biflorum), tend to be diminutive plants that can be easily missed if you are not looking closely. They huddle close to the ground on short, delicate stalks so as not to expend energy creating tissue; most of their energy is expended in ovule and seed development. On Mount Ashland, you will also find that many plants growing along the high ridgelines hug the ground and spread in a matting fashion. These plants survive by becoming dwarfs; they are smaller than their lower elevation counterparts. This growth form yields several advantages to the plant. It brings protection from the winter winds that blast over them, while allowing them to take advantage of the warmth radiated from the soil. The short stems easily conduct water to their flowers. Usually, these plants have masses of flowers that help to advertise to their insect pollinators. Another frequent characteristic of these flowers is hairs. Hairs of all types - bristly, silky, sticky, wavy - work for the flowers in three ways. They trap heat to prevent the plant from freezing, and they are usually white to deflect the harsh sunshine. In this way, the plants can regulate their own temperature and save water. Finally, they deter insects and other animals from eating the plants. You'll find hairy leaves, hairy stems, hairy sepals, and even some hairy petals. Subalpine plants must be frugal; leaves are often small, hairy or waxy. This is especially obvious in flowering shrubs. These sclerophyllous leaves - 6 Background Information leaves that are stiff and waxy - can withstand nightly freezing with complete cessation of water conduction without suffering damage to the plant. You may notice that many flowers, including Pussy Paws (Calyptridium umbellatum), Western Burnet (,Sanguisorba occidentalis) and Oregon Willow-herb (Epilobium oregonense) have red stems. The red coloring is a result of a cellular pigment called anthocyanins. The anthocyanins act as ultraviolet filters, protecting * \£ the plant tissues from radiation caused by the brutal sun. Great sources for additional reading on this topic include Timberline: Mountain and Arctic Forest Frontiers (Arno and Hammerly 1984), Arctic and Alpine Vegetation: Plant Adaptations to Cold Summer Climates (Billings 1974), and Adaptive Plant Evolution in the Alpine Environment of the Greek Mountains (Papanicolaou and Kokkini 1982). Special Species On Mount Ashland Although all the plants on Mount Ashland are special, there are a few flowers that warrant special mention. These species are rare either because they are endemics to this peak (that is, they grow nowhere else in the world) and therefore, have very restricted ranges, or because they are growing at the western-most point of their range, and are rare in this area. Pay special attention and watch for these species.... Mt. Ashland Lupine (Lupinus aridus var. ashlandensis) and Henderson's Horkelia (Horkelia hendersonii): Mount Ashland Lupine is a beautiful flower that is endemic to Mount Ashland! It can be found on the south and west slopes above 2100 m (6800'), and around the summit. The total population consists of 5,000 - 6,000 individuals and covers an area of about 24 ha (60 acres). Currently, there are six populations of Henderson's Horkelia recorded in the United States. They are all found in the Siskiyou Mountains of southern Oregon and northern California, within the Klamath National Forest. In Oregon, the populations are found on Mount Ashland, McDonald Peak, Split Rock, Dutchman Peak- Jackson Gap, and Observation Peak. In California, one population has been recorded at Dry Lake Lookout, eight miles south of the Oregon-California state line. The Mount Ashland population is the largest. The U.S. Forest Service considers Mount Ashland Lupine and Henderson's Horkelia sensitive species. Essentially, that means that the U.S. Forest Service assumes an obligation to maintain viability of these species. From 1980 to 1996, both flowers were Category 2 candidates for listing under the U.S. Endangered Species Act. This category included species for which sufficient data was not available to confirm vulnerability. In 1996, this category of species was discontinued, and both species were dropped. In September 1999, the U.S. Fish and Wildlife Service received a formal petition from the Rogue Group of the Sierra Club of Ashland to list Mount Ashland Lupine and Henderson's Horkelia as endangered or threatened. In June 2000, the U.S. Fish and Wildlife Service announced that the two flowers are candidates, and a comprehensive review of both flowers would be conducted to determine if they should be protected under the Endangered Species Act. As of this writing, no final decision has been made. Jayne's Canyon Buckwheat (Eriogonum diclinum): This special flower is endemic to the Klamath Mountains in northern California and southern Oregon. That is, if you want to see these flowers, you must come to the Klamath Mountains because they grow nowhere else in the 7 r Background Information world. Currently, botanists are aware of about 5 populations of Jayne's Canyon Buckwheat in Oregon, and the population on Mount Ashland is one of the largest. The U.S. Forest Service considers this a sensitive species, assuming an obligation to maintain its viability. American Sawwort (Saussurea americana): At the edge of its range and rare in the Siskiyou Mountains, American Sawwort is of great local concern because of its contribution to local and regional floral diversity. There are four known populations in the Siskiyou Mountains, and the population on Mount Ashland is one of them. However, it is more common farther north in the Cascades and in Northeast Oregon. This flower is not afforded any legal or U.S. Forest Service protective status. What's In A name? As we get to know the flowers, they become our friends. And like friends, we want to know their names. Common names are very interesting because they arise from discoverers, myths and legends, folklore, healing properties, and the colors, shapes, and smells of the flowers. For example, on Mount Ashland, there is Sneezeweed, Serviceberry, Miner's Lettuce, and Trip Vine. However useful these names are in certain regional areas, common names can lead us astray. This is why I have included the scientific names, as well as the derivation of the Latin and Greek words. The scientific names can tell us a great deal more about a flower, and add to a fond relationship with it. In deciphering the scientific names, we often learn that they are the true sources for many of the popular, common names. The system of naming plants with scientific names was developed by the Swedish botanist, Carl von Linne (1707-1778), also known as Carl Linneaus. Linneaus chose Latin because it was at that time the international academic form of communication. Today, Latin and Greek are the languages most drawn on for these names, but in some cases the words are latinized versions of other languages. The complete scientific name for a plant has two parts, and is therefore, called a binomial. The first part is the genus (plural = genera), the second part is the specific epithet. Together, they create a unique name that often contains clues for the plant's identification. In most texts, the binomial scientific name appears in italics. Generic names are always capitalized, and the specific epithet is always in lower case letters. The binomials are immediately followed by the last name of the author, or the accepted abbreviation of the author's name, who published the species (plural = species) name. Often there will be two names, one in parentheses and one without. The name in parentheses is the original author who placed the species in a different genus. The name without parenthesis is the author who placed the species in the current genus. The scientific name of a plant is never complete without the name of the publishing author(s) attached. Aside from using the scientific name as the ultimate reference for identification, the Latin and Greek words can provide a lot of interesting information. For starters, the general habitat of a plant is often indicated. For example, palustris means "of the swamp" and arvense means "afield." Knowing this will make it easier if you want to know where to look to find a particular plant. The Latin and Greek words can also tell you what a plant might look like. For example, lanatum means "woolly," and chrysophyllus means "yellow leafed." If you were looking for plants 8 Background Information with these words in the scientific name, you would know to look for a plant with many fuzzy hairs, or a plant with yellow leaves. As you learn about the flowers in this field guide, give the scientific names a try. You might be surprised at how easy it is. For a deeper understanding of the scientific names, try the following resources: Dictionary of Word Roots and Combining Forms (Borror 1988), and Botanical Latin: History, Grammar, Syntax, Terminology, and Vocabulary (Stearn 1992). To learn more about botanical exploration in the West and many of the plant authors, try Botanical Exploration of the TransMississippi West, 1790-1850 (McKelvey 1991). Using the Metric System Throughout the key and in the written descriptions, I have used the metric system for measurement because it is the conventional system in science. To make the transition from the English system to the metric system easier, here are a few tips. The whole system is based on tenths. In this book, the millimeter is the smallest unit of measurement. A millimeter (mm) is approximately the depth of a dime. Ten millimeters make one centimeter (cm), which is approximately the width of an adult male's small fingernail, or a little less than a lA inch. Ten centimeters make one decimeter (dm), which is roughly equivalent to four inches. Thirty centimeters is approximately the length of one foot. 9 PART 2 The Key System and How It Works "My soul can find no staircase to Heaven unless it be through Earth's loveliness." Michelangelo The Key System and How It Works EXPLANATION OF BASIC TERMS All the terms described in this section are for the purposes of this field guide. For illustrations of these terms, refer to Appendix B. Flower Descriptions: Radial, Bilateral, and Indistinguishable Radial Flowers are radially symmetrical meaning that the petals, or parts that look like petals, extend from the center like spokes of a wheel. If these flowers were pies, each piece of the pie would look more or less similar. Most of the flowers you will find on Mount Ashland are radial. Some examples are Common Blue Flax (Linum lewisii), Oregon Sunshine (Eriophyllum lanatum), and Alpine Shooting Star (Dodecatheon alpinum). Some plants, like Ballhead Ipomopsis (Ipomopsis congesta var. congesta) and Pussy Paws (Calyptridium umbellatum), have flowers that grow in tight bunches. It is important to isolate one single flower to determine whether the flower is radial or not. Bilateral Flowers are bilaterally symmetrical, meaning that there is only one way to divide the flower so that the parts would be mirror-images of each other. The petals, or parts that look like petals, generally do not look similar to each other. Usually, these flowers have upper and lower lips. Some examples of bilateral flowers found on Mt. Ashland include Coyote Mint {Monardella odorotissima), Meadow Larkspur (Delphinium nuttallianum), and Mt. Ashland Lupine (Lupinus aridus var. ashlandensis). Indistinguishable flowers have no visible petal-like parts, or the parts are so small that they are not easily seen. Be careful with this category: a flower may be very small but still have discernable parts. If you use this category and cannot find your specimen, reexamine the flower, perhaps with a small 5x or 1 Ox hand lens. It may be an irregular flower that is difficult to distinguish. Plant Descriptions: Herbaceous Flowers, Shrubs, and Vines Herbaceous wildflowers are plants that are not woody and not trailing or climbing. They can be vigorous and grow quite tall, but there will be no woody features. When in doubt, check the very base of the stem for a woody appearance. If it does not look or feel like wood, the plant is herbaceous. Shrubs are any woody plant that is less than 3 meters tall. Usually, shrubs have more than one stem or branch from the base of the plant. Some plants, such as Coyote Mint {Monardella odorotissima) and Sulfur Flower (Eriogonum umbellatum), have woody stems and are, therefore, technically shrubs. But since many people might overlook this feature, they are considered herbaceous wildflowers. A vine is a trailing or climbing plant, sometimes with thorns and/or tendrils. Leaf Arrangements: Basal, Alternate, Opposite or Whorled Basal leaves grow only at the base of the plant; leaves growing along the stem are cauline leaves, or stem leaves. Alternate leaves grow singly at different points along the stem. Opposite leaves appear at the same point on the stem in pairs, directly across from each other. Whorled leaves appear at the same point on the stem in groups of 3 or more. 11 The Key System and How It Works When determining whether the leaves are alternate or opposite, it is best to look at the largest leaves toward the middle of the stem. Try to avoid using the leaves at the top of the stem as they are sometimes crowded, and it is difficult to determine their arrangement. If there is only one leaf on the stem it is considered alternate. Leaf Descriptions: Entire, Toothed, Lobed, or Compound There are two very basic leaf types: simple and compound. If the leaf blade is made of one undivided unit, it is a simple leaf. If the blade is divided into leaflets, the leaf is compound. In order to determine whether a leaf is simple or compound, look for next years bud. The buds are always at the base of the leaf. In this field guide, the margins of simple leaves are described as entire, toothed, or lobed. A leaf that has an unbroken margin is referred to as entire. An example of a plant with entire leaves is Pussy Ears (Calochortus tolmiei). Toothed leaves have serrated edges. That is, the leaf margin is broken into pointed segments. These pointed segments, or teeth, can be very shallow so as to be barely noticeable, or they can be so deep that the leaf may appear to be compound. An example of a plant with toothed leaves is Nettle-leaf Giant Hyssop (Agastache urticifolia). A leaf that has a margin broken into wide, round segments is referred to as lobed. An example of a plant with lobed leaves is Sticky Currant (Ribes viscosissimum). Compound leaves are made up of several to many leaflets. A palmately compound leaf has leaflets that all emerge from the same point, similar to the fingers on your hand. A pinnately compound leaf has leaflets that emerge from different points along the central vein, similar to a feather. Brewer's Lupine (Lupinus breweri) has palmately compound leaves, and Showy Jacob's Ladder (Polemonium californicum) has pinnately compound leaves. When keying plants, it is important to remember that leaflets can also be entire, toothed or lobed. If the plant you are trying to identify has compound leaves, follow the key break for compound leaves, regardless of whether the leaflets are entire, toothed, or lobed. Basic Floral Structure There are four major parts of a flower: sepals, petals, stamens, and pistil. A complete flower will have all four parts; an incomplete flower could have only one, two, or three of these parts. The sepals are the usually green leaf-like appendages on the outmost portion of the flower. Collectively, the sepals are referred to as the calyx. Their purpose is to protect the flower while it is in the bud stage, but the calyx usually persists while the flower is in bloom. The petals are what most people notice first when they look at a flower. The petals are usually the showy, colorful part of the flower. Collectively they are referred to as the corolla, and their primary purpose is to attract pollinators. The reproductive structures are found in the center of the flower. The stamens, which are the male reproductive parts, consist of the pollen-bearing anther and the filament. The pistil, or the female reproductive part, consists of the stigma, the style, and the ovary. The stigma is often sticky so that the pollen will adhere to it. Pollen tubes grow down through the elongated style and into the ovary. Inside the ovary are the ovules, which will become seeds after fertilization. Most flowers have both the male and female reproductive parts, and are therefore called perfect flowers. However, some flowers have only one reproductive part, and is called an imperfect flower. Pistillate flowers contain only pistils, and staminate flowers contain only 12 The Key System and How It Works stamens. These unisexual flowers can grow on the same plant or on different plants (one plant with only pistillate flowers and a second plant with only staminate flowers). If they grow on the same plant, the plant is called monoecious. If pistillate flowers grow on one plant and staminate flowers grow on a separate plant, the plants are called dioecious. Floral Inflorescences Flowers on the same plant grow in several different arrangements. The general term for the arrangement of flowers is the inflorescence. The inflorescences you will encounter on Mount Ashland include umbels, heads, corymbs, spikes, racemes, panicles, cymes, paired or whorled in the axils, and solitary flowers. Examples of plants with these inflorescences: Swamp White Heads (Sphenosciadium capitellatum) - umbel Orange Agoseris (Agoseris aurantiaca) and White Clover (Trifolium repens) - head Yarrow (Achillea millefolium) - corymb (of heads) White-flowered Bog Orchid (Platanthera leucostachys) - spike White-veined Wintergreen (Pyrola picta) - raceme Jessica's Stickseed (Hackelia micrantha) - panicle Common Blue Flax (Linum lewisii) - cyme Changeable Phacelia (Phacelia mutabilisa) - one-sided coiled cyme Wedded Honeysuckle (Lonicera conjugialis) - paired in axils Stinging Nettle (Urtica dioica) - whorled in axils White Pasque Flower (Anemone occidentalis) - solitary flower For definitions of these inflorescences, refer to the glossary in Appendix A. The Illustrated Glossary in Appendix B has illustrations of each. 13 The Key System and How It Works IDIOSYNCRASIES OF THE KEY Every key system has its share of quirks and quibbles, and sometimes you wonder just what the author was thinking. In this section, I'll try to explain most of quirks of this key. First of all, this key is designed for beginners, so I assumed that there may be some confusion about certain things including the Sunflower Family (Asteraceae), compound leaves, and perianth parts to name a few. Sunflower Family (Asteraceae): I know this pains many experienced botanists, but this key assumes that the user is not aware that plants in the Sunflower Family have a head of flowers. It assumes that the user will think that one head of flowers is one flower. Therefore, Oregon Sunshine (Eriophyllum lanatum) and Orange Agoseris (Agoseris aurantiaca) are radial flowers with more than 7 petals. Compound Leaves: The key was developed so that a plant with compound leaves can be keyed out as a plant with compound leaves, or with lobed leaves. Accordingly, Big-leaf Lupine {Lupinus polyphyllus) can be keyed out as a plant with compound leaves and as a plant with deeply lobed leaves. Perianth Parts: I assumed that the user would not be able to tell the difference between petals and showy sepals, especially if the plant has no petals. That is why the choices under "Flower Descriptions" include "petals or petal-like parts." Therefore, plants like Drummond's Anemone (Anemone drummondii), which has no petals but showy sepals, is keyed out as if the sepals are petals. And Golden Chinquapin (Chrysolepis chrysophyllus), which has no petals but small green stamens, is keyed out as if the sepals are petals. Leafless Stems: In some cases, the key assumes that plants with leafless stems are plants with "no apparent leaves," even though the plant may have basal leaves. This, again, is for the neophyte who may not realize that the leaves surrounding the flowering stem are, in fact, basal leaves. I apologize if these idiosyncrasies are difficult or insulting to experienced botanists. In my opinion, this is the best way to introduce beginners to wildflowers without confusing them to the point of no return. 14 The Key System and How It Works THE KEY SYSTEM AUD HOW IT WORKS This key was based on the system invented by Lawrence Newcomb and published in 1977 in Newcomb's Wildflower Guide, created for the northeastern United States. Identification begins by answering several easy questions about the plant you are looking at. After answering these questions, you will have a three-digit number. Find this number in the Locator Key, which is a basic dichotomous key. This is a series of two-parted statements that describe the plants. The plant you are looking at will fit the description of only one statement in the first series. Then, move to the next series (indicated by the next level of indentation), and your plant will fit the description of only one of these statements. You continue using this pattern until the key indicates a species name. Then, turn to the page indicated for a thorough species description, photograph, and line drawing. The Questions Looking at the flower... • Is the flower radial or bilateral, or are the flower parts indistinguishable? • If the flower is radial, how many petals (or petal-like parts) does it have? Looking at the overall plant... • Is the plant an herbaceous flower, a shrub, or a vine? • If the plant is an herbaceous flower, does it have leaves? • If the plant is herbaceous and has leaves, are they all at the base of the plant, arranged alternately on the stem, or arranged opposite one another in pairs or whorls? Looking at the leaves... • Are the leaves simple or compound? • If the leaves are simple, are the leaf margins entire (even and unbroken), or are they toothed, or lobed? If you need further explanation of any of these terms, turn to Appendix B where the terms are illustrated. Determining the Three-Digit Number Each plant is classified in three ways. The first is by the flower description: the arrangement and number of petals or petal-like parts. The second is by plant description herbaceous flower, shrub, or vine - and if the plant is an herbaceous flower, by the arrangement of its leaves. The third is by leaf description. The combination of these three classifications determines the plant group number to which your plant belongs. These three classifications and their corresponding numbers are on page 17. For example, if your flower is bilateral (number 1 under flower description), and is an herbaceous wildflower with alternate leaves (number 3 under plant description), and has toothed leaves (number 3 under leaf description), your group number is 133. Another example: Common Blue Flax will be found in group number 532 because it is a radial flower with 5 petals (5—), an herbaceous flower with alternate leaves (53-), and its leaves are entire (532). After determining the three-digit number, turn to the Locator Key where your search will be narrowed until the exact species is known. 15 The Key System and How It Works puTTirrr IT ALL TOGETHER 1. Answer the following questions: Flower Description a. Is the flower radial or bilateral, or are the flower parts indistinguishable? b. If the flower is radial, how many petals or petal-like parts does it have? (Note: Because there is some variation among flowers of the same species, it helps to look at more than one of the same flower. Some flowers lose petals, some leaves get chewed on by insects or small mammals, etc.) Plant Description c. Is the plant a wildflower, a shrub, or a vine? d. If it is a wildflower, does it have leaves? e. If the wildflower has leaves, are they all at the base of the plant, or are they arranged on the stem in an alternate fashion, opposite or whorled? (Note: If there is only one stem on the leaf, it is considered alternate.) Leaf Description f. Are the leaves simple or compound? g. If simple, are the margins entire (even and unbroken), toothed, or lobed? 2. Use the chart on the next page to determine the three-digit number for the plant. 3. Find the three-digit number in the Locator Key and follow the dichotomous key to determine the plant. a. The dichotomous key is a series of two-parted statements that describe a plant characteristic. Series are indicated by the level of indentation. b. Your specimen will match only one of the statements in the first series. Drop down to the next level of indentation, and match your specimen to the correct statement in this series. c. Continue until you arrive at the species name and page number. 4. Turn to the page number indicated to confirm that you have the correct plant. You can use the written description, photograph, and drawing for confirmation. 16 The Key System and How It Works CHART FOR DETERMINING THE THREE-DIGIT NUMBER FLOWER DESCRIPTION PLANT DESCRIPTION LEAF DESCRIPTION Bilateral flowers Radial flowers with 2 petals or petal-like parts Radial flowers with 3 petals or petal-like parts Radial flowers with 4 petals or petal-like parts Radial flowers with 5 petals or petal-like parts Radial flowers with 6 petals or petal-like parts Radial flowers with 7 or more petals or petal-like parts Parts Indistinguishable No apparent leaves Herbaceous Basal leaves only Wildflowers Alternate leaves Opposite or Whorled Leaves Shrubs Vines No Apparent Leaves Leaves Entire Leaves Toothed or Lobed Leaves Compound 1 2 3 4 5 6 7 8 1 2 3 4 5 6 1 2 3 4 17 The Key System and How It Works LOCATOR KEY BILATERAL FLOWERS 111 Bilateral Wildflowers, No Apparent Leaves Plant green and photosynthetic Mt. Ashland Lupine (Lupinus aridus var. ashlandensis) p. 124 Plant not green Plant 10 cm or shorter Inflorescence a tight spike with many bracts California Ground Cone (Boschiaka strobilacea) p. 170 Inflorescence not a spike Flowers in wet or moist soil Naked Broomrape (Orobanche uniflora) p. 172 Flowers in dry, bare soil Clustered Broomrape (Orobanche fasciculata) p. 170 Plant 13 cm or taller Lower lip of flower white with purple spots Spotted Coralroot (Corallorhiza maculata) p. 162 Lower lip of flower not clearly spotted Sepals and upper 2 petals clearly striped Striped Coralroot (Corallorhiza striata) p. 164 Sepals and upper 2 petals not clearly striped Western Coralroot (Corallorhiza mertensiana) p. 164 122 Bilateral Wildflowers, Basal Leaves Only, Leaves Entire Flowers white or cream Leaves dark green with prominent white veins Leaves solid green Flowers yellow, pink, purple Leaves very hairy Leaves not obviously hairy, or not hairy at all 123 Rattlesnake Plantain (Goodyera oblongifolia) p. 166 Macloskey's Violet {Viola macloskeyi) p.240 '. Primrose Monkey Flower (Mimulus primuloides) p.230 Calypso Orchid (Calypso bulbosa) p. 162 Bilateral Wildflowers, Basal Leaves Only, Leaves Toothed or Lobed Leaves palmately lobed Flowers deep purple, with long spur Yellowtinge Larkspur (.Delphinium decorum) p. 198 Flowers lavender, not spurred Mt. Ashland Lupine (Lupinus aridus var. ashlandensis) p. 124 Leaves crenate or with rounded teeth, flowers white or cream Macloskey's Violet (Viola macloskeyi) p.240 124 Bilateral Wildflowers, Basal Leaves Only, Leaves Compound Leaves palmately compound Flowers with long spur Flowers not spurred Leaves pinnnately compound 132 Yellowtinge Larkspur (Delphinium decorum) p. 198 Mt. Ashland Lupine (Lupinus aridus var. ashlandensis) p. 124 Steer's Head (Dicentra uniflora) p. 172 Bilateral Wildflowers, Alternate Leaves, Leaves Entire Plants green and photosynthetic Plants with colorful bracts beneath or in front of the flowers Flowers primarily bright red (perhaps with some yellow) Plant with many dense, gray hairs Frosted Indian Paintbrush (Castilleja pruinosa) p.224 Plant glabrous, or sparsely hairy Plant of moist places Giant Red Indian Paintbrush (Castilleja miniata) p.224 Plant of dry places Coast Indian Paintbrush (Castilleja affinis) p.222 Flowers primarily pink and white Flower extended beyond bract, large white balloon Siskiyou Mountains Owl's Clover (Orthocarpus cuspidatus) p.230 Flower not or barely extended beyond bract Mountain Owl's Clover (Orthocarpus imbricatus) p.232 18 The Key System and How It Works Plants with no bracts, or with green bracts Stamen fused with style and stigma forming a column Lower petal spurred Flowers white White-flowered Bog Orchid (Platanthera leucostachys) p. 168 Flowers green Spur short and blunt Green-flowered Bog Orchid (Platanthera hyperborea) p. 166 Spur long and pointed Sparse-flowered Bog Orchid (.Platanthera sparsiflora) p. 168 Lower petal not spurred Basal leaves with obvious white veins Rattlesnake Plantain (Goodyera oblongifolia) p. 166 Basal leaves solid green Inflorescence with large bracts Hooded Ladies' Tresses (Spiranthes romanzoffiana) p. 168 Inflorescence with very small bracts Broad-leaved Twayblade (Listera convallarioides) p. 166 Stamens and styles separate Flowers yellow Plant tall with long raceme/spike Dalmatian Toadflax (Linaria genistifolia) p.226 Plant small with solitary flowers Mountain Violet ( Viola purpurea) p.242 Flowers bicolored - white and deep purple/blue Plant very glandular, sepals wide and blunt Torrey's Blue-eyed Mary (Collinsia torreyi) p.226 Plant minimally glandular; sepals narrow, sharp Blue-eyed Mary (Collinsiaparviflora) p.226 Plants not green Plant with slender stem Entire plant white or cream Phantom Orchid (Cephalanthera austinae) p. 162 Plant mostly purplish-red Lower petal of flower white with purple spots Lower lip with 2 purple spots Western Coralroot (Corallorhiza mertensiana) p. 164 Lower lip with 4 or more purple spots Spotted Coralroot (Corallorhiza maculata) p. 162 Lower petal of flower not clearly spotted Striped Coralroot (Corallorhiza striata) p. 164 Plant without obvious stem, many brown bracts California Ground Cone (Boschniaka strobilacea) p. 170 Bilateral Wildflowers, Alternate Leaves, Leaves Toothed or Lobed Leaves serrate or crenate Flowers pale pink to white Leafy Lousewort (Pedicularis racemosa) Flowers yellow Leaves purple-tinged, in dry areas Mountain Violet {Violapurpurea) Leaves solid-green, in wet areas Stream Violet {Violaglabella) Leaves lobed Leaves palmately lobed Leaf lobes toothed Flowers with long spur Plant 60 cm or taller Mountain Larkspur (Delphinium glaucum) Plant shorter than 60 cm Upper and lower leaf surface glabrous Leaf lobes < 4mm wide Anderson's Delphinium (Delphinium andersonii) Leaf lobes < 7mm wide Meadow Larkspur (Delphinium nuttalianum) Lower leaf surface minutely hairy Yellowtinge Larkspur (Delphinium decorum) Flowers not spurred, large hood-like petal Columbian Monkshood (Aconitum columbianum) Leaf lobes entire Plant 60 cm or taller, in wet areas Big-leaf Lupine (Lupinus polyphyllus) Plant shorter than 60 cm, in dry areas Sickle-keel Lupine (.Lupinus albicaulis) Leaves pinnately lobed Flowers red, orange, yellow Plant cob-webby/woolly, flowers pale yellow Cobwebby Paintbrush (Castilleja arachnoidea) Plant not cob-webby/woolly, flowers red/orange Frosted Indian Paintbrush (Castilleja pruinosa) Flowers not red, orange, yellow Flower extended beyond bract Siskiyou Mountains Owl's Clover (Orthocarpus cuspidatus) p.232 p.242 p.240 p. 198 p. 198 p.200 p. 198 p. 190 p. 126 p. 124 p.224 p.224 p.230 19 The Key System a n d How It Works Flower not or barely extended beyond bract 134 Mountain Owl's Clover (Orthocarpus imbricatus) p.232 Bilateral Wildflowers, Alternate Leaves, Leaves Compound Leaflets 3 Inflorescence with obvious colorful bracts Flowers strongly bicolored, white and pink-purple Flower extended beyond bract Siskiyou Mountains Owl's Clover (Orthocarpus cuspidatus) p.230 Flower not or barely extended beyond bract Mountain Owl's Clover (Orthocarpus imbricatus) p.232 Flowers yellowish-green Cobwebby Paintbrush (Castilleja arachnoidea) p.224 Inflorescence with no bracts, or very small bracts Inflorescence dense, head-like Leaflets narrow, finely-toothed Long-Stalk Clover (Trifolium longipes) p. 126 Leaflets as wide as long, heart-shaped White Clover (Trifolium repens) p. 128 Inflorescence solitary flowers in leaf axils Bird's Foot Trefoil {Lotus purshianus var. purshianus) p. 122 Leaflets more than 3 Flowers yellow Shelton's Violet (Viola sheltonii) p.242 Flowers pink, purple, or blue Flowers spurred Upper and lower leaf surface glabrous Anderson's Delphinium (Delphinium andersonii) p. 198 Lower leaf surface minutely hairy Yellowtinge Larkspur {Delphinium decorum) p. 198 Flowers not spurred Leaves pinnately compound Leafy Pea {Lathyrus polyphyllus) p. 122 Leaves palmately compound Plant 30 inches or taller Big-leaf Lupine {Lupinus polyphyllus) p. 126 Plant shorter than 30 inches Sickle-keel Lupine {Lupinus albicaulis) p. 124 142 Bilateral Wildflowers, Opposite or Whorled Leaves, Leaves Entire Flowers yellow, white, or brownish-greenish Only two stem leaves, leaves rounded Broad-leaved Twayblade {Listera convallarioides) More than two stem leaves Leaves glabrous Seep Monkey Flower {Mimulus guttatus) Leaves very hairy Musk Monkey Flower {Mimulus moschatus) Flowers pink, purple, lavender, blue Plants woody at the base Inflorescence short raceme Davidson's Penstemon {Penstemon davidsonii) Inflorescence dense, head-like Coyote Mint {Monardella odoratissima) Plants not woody Leaves narrow, at least 4x longer than wide Flowers strongly bicolored: upper lip white, lower lip blue or purple Sepals sharply pointed Blue-Eyed Mary {Collinsiaparviflora) Sepal tips rounded or blunt Torrey's Blue-Eyed Mary {Collinsia torreyi) Upper lip and lower lip more or less same color Siskiyou Penstemon {Penstemon anguineus) Leaves wide, no more than 2x longer than wide Flowers with 4 petals, almost radial Thyme-leaf Speedwell (Veronica serpyllifolia) Flowers with 5 petals or lobes, clearly bilateral Plant more than 10 cm tall Azure Penstemon {Penstemonparvulus) Plant less than 10 cm tall Dwarf Purple Monkey Flower {Mimulus nanus) 143 p.228 p.228 p.234 p. 142 p.226 p.226 p.232 p.236 p.234 p.228 Bilateral Wildflowers, Opposite or Whorled Leaves, Leaves Toothed or Lobed Stamens 2 Axillary racemes Flowers solitary in axils Stamens more than 2 20 p. 166 American Brooklime ( Veronica americana) p.236 Thyme-leaf Speedwell {Veronica serpyllifolia) ^>.22)6^ The Key System and How It Works Stamens strongly exserted Stem square, glabrous Stem round, glabrous except at nodes Stamens not exserted Flowers 2 cm long or longer Flowers 1.5 cm long or shorter Leaves glabrous Leaves hairy 144 Nettle-leaf Giant Hyssop (Agastache urticifolia) p. 140 Sitka Valerian (Valeriana sitchensis) p.240 Woodland Penstemon (Nothochelone nemorosus) p.230 Torrey's Blue-Eyed Mary (Collinsia torreyi) p.226 Rough Hedge Nettle (Stachys ajugoides var. rigida) p. 142 Bilateral Wildflowers, Opposite or Whorled Leaves, Leaves Compound Main axis of leaf ends as a tendril Main axis of leaf without tendril 152 Shrubs with Bilateral Flowers, Leaves Entire Shrub erect, 15 cm or taller Flowers pink, white, or lavender Flowers orange-yellow, or maroon Flowers orange-yellow trumpet shaped Flowers maroon, bilabial Shrub low, mat-forming, less than 10 cm tall 153 .....Leafy Pea (Lathyrus polyphyllus) p. 122 Sitka Valerian (Valeriana sitchensis) p.240 Coyote Mint (Monardella odoratissima) p. 142 Orange Honeysuckle (Lonicera ciliosa) p. 104 Wedded Honeysuckle (Lonicera conjugialis) p. 104 Davidson's Penstemon (Penstemon davidsonii) p.234 Shrubs with Bilateral Flowers, Leaves Toothed or Lobed Low, mat-forming shrub with lavender flowers 154 Brewer's Lupine (Lupinus breweri) p. 124 Shrubs with Bilateral Flowers, Leaves Compound Low, mat-forming shrub with lavender flowers Brewer's Lupine (Lupinus breweri) p. 124 RADIAL FLOWERS WITH 2 PETALS OR PETAL-LIKE PARTS 222 Radial Flowers, 2 Petals, Basal Leaves Flowers creamy white or pink with red anthers 232 Radial Flowers, 2 Petals, Alternate Leaves, Leaves Entire Leaves, stems, sepals with black glands, very sticky Sticky oil foul smelling Sticky oil pleasant smelling Leaves, stems, sepals without black glands, fairly sticky 233 Mountain Tarweed (Madia glomerata) p.82 Slender Tarweed (Madia gracilis) p.82 Greene's Goldenbush (Ericameria greenei) p.72 Radial Flowers, 2 Petals, Alternate Leaves, Leaves Toothed or Lobed Flowers yellow, plant sticky, foul-smelling 242 Pussy Paws (Calyptridium umbellatum) p. 186 Mountain Tarweed (Madia glomerata) p.82 Radial Flowers, 2 Petals, Opposite or Whorled Leaves, Leaves Entire Flowers yellow, plant sticky, foul-smelling Mountain Tarweed (Madia glomerata) p.82 21 The Key System and How It Works 243 Radial Flowers, 2 Petals, Alternate Leaves, Leaves Toothed or Lobed Flowers yellow, plant sticky, foul-smelling Mountain Tarweed (Madia glomerata) p.82 RADIAL FLOWERS WITH 3 PETALS OR PETAL-LIKE PARTS 312 Radial Flowers, 3 Petals, No Apparent Leaves Flowers maroon with 3 long-tapered petal-like sepals 322 Wild Ginger (Asarum caudaturri) p.60 Radial Flowers, 3 Petals, Basal Leaves Only, Leaves Entire Leaves linear, grass-like Many small flowers in tight terminal cluster 1 or 2 fairly large flowers Petals hairy Petals not hairy Leaves not grasslike Flowers bright yellow Flowers deep maroon with long-tapered petals 332 Pussy Ears (Calochortus tolmiei) p. 144 Blue-eyed Grass (Sisyrinchium bellum) p. 140 Sulfur Flower (Eriogonum umbellatum) p. 182 Wild Ginger (Asarum caudatum) p.60 Radial Flowers, 3 Petals, Alternate Leaves, Leaves Entire At least upper half of stem very sticky Leaves, stems, sepals with black glands, very sticky Sticky oil foul smelling Sticky oil pleasant smelling Leaves, stems, sepals without black glands, fairly sticky Upper half of stem not sticky Leaves linear, grass-like Petals hairy Petals not hairy. Flowers deep blue or purple Flowers white-yellowish Leaves elliptic or lanceolate, not grass-like Flowers green; large, stout plant Flowers deep red; dainty plant 333 Pacific Onion (Allium validum) p. 142 Mountain Tarweed (Madia glomerata) p.82 Slender Tarweed (Madia gracilis) p.82 Greene's Goldenbush (Ericameria greenei) p.72 Pussy Ears (Calochortus tolmiei) p. 144 Blue-eyed Grass (Sisyrinchium bellum) p. 140 Yellow-flowered Iris (Iris chrysophylla) p. 140 Radial Flowers, 3 Petals, Alternate Leaves, Leaves Toothed or Lobed Plant sticky and foul-smelling Plant not sticky Petals red or maroon Petals pink, white, or green Leaves entire to shallowly toothed, margins may be wavy Leaves fern-like with many segments 334 Mountain Tarweed (Madia glomerata) p.82 Sheep Sorrel (Rumex acetosella) p. 184 Willow Dock (Rumex salicifolius) p. 186 Yarrow (Achillea millefolium) p.60 Radial Flowers, 3 Petals, Alternate Leaves, Leaves Compound Leaves fern-like, petals white or cream 22 Willow Dock (Rumex salicifolius) p. 186 Sheep Sorrel (Rumex acetosella) p. 184 Yarrow (Achillea millefolium) p.60 V The Key System and How It Works Radial Flowers, 3 Petals, Opposite or Whorled Leaves, Leaves Entire Flowers yellow Flowers not yellow Flowers large (25 mm or more wide) Flowers small (less than 5 mm across) 343 Mountain Tarweed (Madia glomerata) p.82 Western Trillium (Trillium ovatum) p. 150 Low Mountain Bedstraw (Galium bifolium) p.216 Radial Flowers, 3 Petals, Alternate Leaves, Leaves Toothed or Lobed Flowers yellow Mountain Tarweed (Madia glomerata) p.82 RADIAL FLOWERS WITH 4 PETALS OR PETAL-LIKE PARTS 411 Radial Flowers, 4 Petals, No Apparent Leaves White flowers enclosed in brownish bracts 422 California Ground Cone (Boschniaka strobilacea) p. 170 Radial Flowers, 4 Petals, Basal Leaves Only, Leaves Entire Plant woody at base Plant not woody Plant decumbent or prostrate Plant erect Petals lavender or purple, reflexed Petals green or colorless 432 Pioneer Rockcress (Arabisplatysperma) p.96 Pussy Paws (Calyptridium umbellatum) p. 186 Alpine Shooting Star (Dodecatheon alpinum) p.190 English Plantain (Plantago lanceolata)p.\14 Radial Flowers, 4 Petals, Alternate Leaves, Leaves Entire Flowers with 8 stamens (Evening Primrose Family) Flowers white Petals notched Tall Annual Willow-herb (Epilobium brachycarpum) p. 158 Petals notched more than Vi petal length Petals notched less than Vi petal length Milkflower Willow-herb (Epilobium lactiflorum) p. 160 Petals entire Spreading Ground Smoke (Gayophytum diffusum) p. 160 Flowers purple, lavender, or pink Inflorescence long raceme, at least 30 cm Fireweed (Epilobium angustifolium) p. 156 Inflorescence short raceme, no longer than 15 cm Leaf axils leafy Fringed Willow-herb (Epilobium ciliatum) p. 158 Leaf axils not leafy Hornemann's Willow-herb (Epilobium hornemannii) p.158 Flowers with 2, 3, 4, 5 or 6 stamens, or stamens indistinguishable Flowers with 6 stamens, 4 long, 2 short (Mustard Family) Fruits heart-shaped Shepherd's Purse (Capsella bursa-pastoris) p.98 Fruits long and linear Flowers bright yellow-orange Western Wallflower (Erysimum capitatum) p. 100 Flowers white, cream, pale pink, or lavender Fruits strongly reflexed (pointed downward) Holboell's Rockcress (Arabis holboellii) p.96 Fruits erect to spreading Plant more or less woody at base Seeds with wide wing Pioneer Rockcress (Arabis platysperma) p.96 Seeds with narrow wing Lyall's Rockcress (Arabis lyallii) p.96 Plant not woody Fruits strongly appressed, erect, flowers cream Tower Rockcress (Arabis glabra) p.94 Fruits spreading, flowers pink/lavender..Spreading Pod Rockcress (Arabis xdivaricarpa) p.94 23 The Key System and How It Works Flowers with 2, 3, 4, or 5 stamens, or stamens indistinguishable Flowers yellow Leaves, stems, sepals with black glands, very sticky Sticky oil foul smelling Mountain Tarweed (Madia glomerata) Sticky oil pleasant smelling Slender Tarweed (Madia gracilis) Leaves, stems, sepals without black glands, fairly sticky..Greene's Goldenbush {Ericameria greenei) Flowers white or pale pink Plant green and photosynthetic Plant erect, many round fruits Virginia Pepperweed (Lepidium virginicum) Plant decumbent to prostrate, red anthers Pussy Paws (Calyptridium umbellatum) Plant not green California Ground Cone (Boschniaka strobilacea) 433 Spreading Pod Rockcress {Arabis xdivaricarpa) p.94 Shepherd's Purse (Capsella bursa-pastoris) p.98 Holboell's Rockcress (Arabis holboellii) p.96 Mountain Tarweed {Madia glomerata) p.82 Pineapple Weed {Chamomilla suaveolens) p.70 Fringed Willow-herb {Epilobium ciliatum) p. 158 Pineapple Weed {Chamomilla suaveolens) p.70 Western Burnet {Sanguisorba occidentalis) p.214 Fendler's Meadow Rue {Thalictrumfendleri) p.202 Yarrow {Achillea millefolium) p.60 Mountain Tansy Mustard {Descurainea incisa) p.98 Baneberry {Acteae rubra) p. 192 Fendler's Meadow Rue {Thalictrum fendleri) p.202 Nuttall's Toothwort {Cardamine nuttallii) p.98 Radial Flowers, 4 Petals, Opposite or Whorled Leaves, Leaves Entire Leaves Opposite Petals notched Flowers white, cream, or yellow Plant very sticky Calyx inflated, petals white or cream Calyx not inflated, petals yellow Plant not sticky or barely sticky 24 Mountain Tansy Mustard (Descurainia incisa) p.98 Western Wallflower (Erysimum capitatum) p. 100 Radial Flowers, 4 Petals, Alternate Leaves, Leaves Compound Petals yellow, green, or no petals Plant pineapple scented Plant not highly aromatic Inflorescence tight spike Inflorescence open panicle Petals white, cream, pale pink, or purplish Stem leaves sessile Petals white or cream, leaves fern-like Petals bright yellow Stem leaves stalked (petioled) Stamens and/or pistils obvious and exserted Flowers white Flowers green or purplish Stamens and/or pistils inconspicuous 442 p. 100 p. 186 p. 170 Radial Flowers, 4 Petals, Alternate Leaves, Leaves Toothed or Lobed Flowers with 6 stamens, 4 long and 2 short (Mustard Family) Flowers yellow Flowers 5-6 mm or narrower Flowers 10-11 mm or wider Flowers white, cream, pink, or lavender Lower stem leaves sagittate and clasping Fruits long, linear, and slender Fruits heart-shaped Lower stem leaves oblong to lanceolate Flowers with 5 or 8 stamens, or stamens indistinguishable Plant with many black glands, very sticky, foul-smelling Plant without black glands, pleasant smelling or no scent Plant pineapple scented • Plant not pineapple scented 434 p.82 p.82 p.72 Palmer's Cacthfly {Silene bernardina) p.108 Mountain Tarweed {Madia glomerata) p.82 The Key System and How It Works Leaves 2-5 cm, oval to lance-shaped Milkflower Willow-herb (Epilobium lactiflorum) p. 160 Leaves less than 1 cm, linear, purple-tipped Oregon Willow-herb (.Epilobium oregonense) p. 160 Flowers lavender, purple, or pink Leaf axils leafy Fringed Willow-herb (Epilobium ciliatum) p. 158 Leaf axils not leafy Hornemann's Willow-herb (Epilobium hornemannii) p. 158 Petals entire Petal tips pointed, petals white Milk Kelloggia (Kelloggia galioides) p.216 Petal tips rounded Petals all white, 8 stamens Spreading Ground Smoke (Gayophytum diffusum) p. 160 Petals blue or blue striped, 2 stamens Thyme-leaf Speedwell (Veronica serpyllifolia) p.236 Leaves Whorled Plant without hooked hairs Sweet Scented Bedstraw (Galium triflorum) p.216 Plant with hooked hairs Goosegrass (Galium aparine) p.214 443 Radial Flowers, 4 Petals, Opposite or Whorled Leaves, Leaves Toothed or Lobed Plants obviously hairy Hairs glandular, sticky Hairs stinging Plants not obviously hairy Inflorescence solitary flowers in leaf axils Inflorescence a raceme Racemes axillary Raceme terminal 452 Mountain Tarweed (Madia glomerata) p.82 Stinging Nettle (Urtica dioica) p.238 Common Speedwell (Veronica arvensis) p.236 American Brooklime (Veronica americana) p.236 Fringed Willow-herb (Epilobium ciliatum) p. 158 Shrub with Radial Flowers, 4 Petals, Leaves Entire Flowers pale pink to white, fruits erect and wide 453 Pioneer Rockcress (Arabis platysperma) p.96 Shrub with Radial Flowers, 4 Petals, Leaves Toothed or Lobed Petals maroon or deep red Oregon Boxwood (Paxistima myrsinites) p. 112 RADIAL FLOWERS WITH 5 PETALS OR PETAL-LIKE PARTS 511 Radial Flowers, 5 Petals, No Apparent Leaves Inflorescence a tight spike with many brown bracts Inflorescence not a spike 1-3 flowers in a cluster, in wet or moist soil More than 3 flowers in a cluster, in dry, bare soil 522 California Ground Cone (Boschiaka strobilacea) p. 170 Naked Broomrape (Orobanche uniflora) p. 172 Clustered Broomrape (Orobanche fasciculata) p. 170 Radial Flowers, 5 Petals, Basal Leaves Only, Leaves Entire Leaves noticeably hairy Primrose Monkey Flower (Mimulus primuloides) p.230 Leaves glabrous Inflorescence a raceme or panicle Leaves with distinct white veins, plant of dry forest areas White-veined Wintergreen (Pyrola picta) p. 120 Leaves solid green, plant of wet places Oregon Saxifrage (Saxifraga oregana) p.220 Solitary, terminal flowers White Marsh Marigold (Caltha leptosepala var. biflora) p. 196 25 The Key System and How It Works 523 Radial Flowers, 5 Petals, Basal Leaves Only, Leaves Toothed or Lobed Flowers or flowering heads wider than 1 cm Petals, or petal-like structures yellow Petals white Flowers or flowering heads smaller than 1 cm Petals entire, white Petals thread-like Petals not thread-like Leaf margins regularly, sharp-toothed Leaf margins irregularly, shallowly lobed Petals pinnately lobed, green Leaf margins round-toothed, dry forest areas Leaf margins sharp-toothed, streamside 524 Naked-stem Hawksbeard (Crepis pleurocarpa) p.72 White Marsh Marigold (Caltha leptosepala var. biflora) p. 196 Brook Saxifrage (Saxifraga odontoloma) p.222 Trail Plant (Adenocaulon bicolor) p.62 Brewer's Mitrewort (Mitella breweri) p.220 Five-point Bishop's Cap (Mitella pentandrd) p.220 Radial Flowers, 5 Petals, Basal Leaves Only, Leaves Compound Plant tall, 20 cm or taller Inflorescence an open cyme Inflorescence an umbel Plant short, 15 cm or shorter Leaflets 3 Leaflets entire Leaflets toothed Leaflets 7-11 532 Lace Flower (Tiarella trifoliata var. unifoliata) p.222 Bigleaf Avens (Geum macrophyllum) p.206 Gray's Lovage (Ligusticum grayi) p.54 Three-leafed Wood Sorrel (Oxalis trilliifolia) p. 172 Wood Strawberry (Fragaria vesca) p.204 Sticky Cinquefoil (Potentilla glandulosa) p208 Radial Flowers, 5 Petals, Alternate Leaves, Leaves Entire Flowers white or cream Plants not green Plant cone-like, with many brown bracts California Ground Cone (Boschniaka strobilacea) p. 170 Plant not cone-like Inflorescence nodding, petals free Pinesap (Monotropa hypopytis) p. 118 Inflorescence erect, petals fused Woodland Pinedrops (Pterospora andromedea) p. 120 Plants green and photosynthetic Leaves narrow, 5x longer than wide Plant obviously hairy or bristly, glandular or not Hairs bristly, not glandular Nutlets 4 Back of nutlet rough Pinewoods Cryptantha (Cryptantha simulans) p.92 Back of nutlet smooth, mottled Torrey's Cryptantha (Cryptantha torreyana) p.92 Nutlet 1 Weak-stem Cryptantha (Cryptantha flaccida) p.92 Hairs soft or glandular Inflorescence a panicle of heads Plant strongly scented Wright's Cudweed (Gnaphalium canescens) p.76 Plant not scented Pearly Everlasting (Anaphalis margariticea) p.64 Inflorescence not as above Crushed leaves stain fingers Staining Collomia (Collomia tinctoria) p. 176 Crushed leaves do not stain fingers Slender Phlox (Phlox gracilis) p. 178 Plant glabrous, or not obviously hairy Leaves with papery stipules at base Inflorescence solitary flowers in leaf axils Petals opening completely Davis's Knotweed (Polygonum davisiae) p. 182 Petals erect Douglas's Knotweed (Polygonum douglasii) p. 184 Inflorescence a dense terminal cluster Western Bistort (Polygonum bistortoides) p.l82| Leaves without stipules at base, plant close to the ground Spreading Phlox (Phlox diffusa) p. 178 26 The Key System a n d How It Works Leaves wide, less than 5x longer than wide Inflorescence one-sided, coiled, uncoiling as flowers bloom Leaves lobed, at least some of them Lobed leaves with 3 lobes Changeable Phacelia (Phacelia mutabilis) p. 136 Lobed leaves with 5-9 lobes Vari-Leaf Phacelia (Phacelia heterophylla) p. 136 Leaves entire Silverleaf Phacelia (Phacelia hastata) p. 134 Inflorescence not coiled Petals fused, flowers urn-shaped Pinemat Manzanita (Arctostaphylos nevadensis) p.l 14 Petals free Leaves succulent, oblanceolate Cream Stonecrop (Sedum oregonense) p.l 14 Leaves leathery but not succulent Basal leaves white-veined White-veined Wintergreen (Pyrola picta) p. 120 Basal leaves solid green One-sided Wintergreen (Orthilia secunda) p. 118 Flowers yellow, peach, brown, pink, blue, purple, Flowers yellow, peach, brown Petals fused Inflorescence an elongated spike Woolly Mullein (Verbascum thapsus) p.234 Inflorescence not an elongated spike Flowers yellow Plant sticky, bush-like Greene's Goldenbush (Ericameriagreenei) p.72 Plant not sticky, not very leafy Naked-Stem Hawksbeard (Crepis pleurocarpa) p.72 Flowers brown or peach Petals large and obvious Large-flowered Collomia (Collomia grandijlora) p. 174 Petals minute and inconspicuous Western Coneflower (Rudbeckia occidentalism p.84 Petals free Leaves succulent Leaves oblanceolate Cream Stonecrop (Sedum oregonense) p.l 14 Leaves lanceolate or elliptic Basal leaves many, clumped, red Spear-leaf Stonecrop (Sedum lanceolatum) p.l 12 Worm-leaf Stonecrop (Sedum stenopetalum) p.l 14 Basal leaves few, if any... Leaves fleshy, or leathery, but not succulent Plant covered with tiny, barbed hairs Bushy Blazing Star (Mentzelia dispersa) p. 154 Plant glabrous Leaves ovate Plantain-leaf Buttercup (Ranunculus alismifolius) p.200 Leaves round or heart-shaped Popular Buttercup (Ranunculus populago) p.200 Flowers pink, blue, purple Inflorescence a corymb of heads American Sawwort (Saussurea americana) p.84 Inflorescence a simple head, raceme, panicle, or flowers solitary in leaf axils Petals obviously fused, plant of wet places Bog Laurel (Kalmia polifolia) p.l 18 Petals free, or not obviously fused Flowers 2 cm wide or more Common Blue Flax (Linum lewisii) p. 154 Flowers 1 cm wide or narrower Flowers 7-20 in a tight terminal head Tiny Trumpet (Collomia linearis) p. 174 Flowers many in an open panicle Jessica's Stickseed (Hackelia micrantha) p.94 Radial Flowers, 5 Petals, Alternate Leaves, Leaves Toothed or Lobed Flowers white or cream Stamens 5, or stamens indistinguishable Flowers in tight heads (Sunflower Family) Flowers cream/pale yellow; plant of dry, open places Flowers white; plant of moist woods Flowers in various inflorescences, not heads Stamens exserted Inflorescence nodding Inflorescence erect Dusty Ma.\dzr\s(Chaenactis douglasii) p. 68 Trail Plant (Adenocaulon bicolor) p.62 White Waterleaf (Hydrophyllum fendleri) p. 134 27 • The Key System a n d How It Works Inflorescence coiled in bud Leaves hastate and entire Silverleaf Phacelia (Phacelia hastata) p. 134 | Leaves all more or less pinnately lobed Tall Phacelia (Phaceliaprocera) p. 136 Inflorescence not coiled in bud Stem glabrous Yampah (Perideridia oregana) p.58 Stem with white, woolly hairs Ballhead Ipomopsis (Ipomopsis congesta) p. 178 Stamens not exserted Flowers 10 mm or wider Large-flowered Boykinia (Boykinia major) p.218 Flowers 5 mm or narrower Small-flower Nemophila (Nemophilaparviflora) p. 134 Stamens more than 5 Flower 1.5 cm or narrower Petals notched or lobed Small-flower Woodland Star (Lithophragmaparviflorum) p.218 Petals entire Petals thread-like Lace Flower (Tiarella trifoliata) p.222 Petals as wide as long, not thread-like One-sided Wintergreen (Orthilia secunda) p. 118 Flower 2 cm or wider Stamens 10 Richardson's Geranium (Geranium richardsonii) p. 130 Stamens greater than 20 Thimbleberry (Rubus parviflorus) p.212 Flowers brown, yellow, orange, red, pink, purple, or blue Flowers brown, yellow, orange, or red Petals free Flower nodding with 5 spurs, red Western Columbine (Aquilegia formosa) p. 196 Flower opening toward sky, no spurs, yellow Bigleaf Avens (Geum macrophyllum) p.206 Petals fused, at least at base Leaves toothed, or serrate Inflorescence one large, terminal head Western Coneflower (Rudbeckia occidentalis) p.84 Inflorescence not as above Plant woody at base Whitney's Bristleweed (Hazardia whitneyi) p.78 Plant not woody Shasta Tansy Aster (Machaeranthera canescens) p.80 Leaveslobed Inflorescence open raceme, flowers large Scarlet Gilia (.Ipomopsis aggregata) p. 176 Inflorescence panicle of heads, flowers small Dusty Maidens (Chaenactis douglasii) p.68 Flowers pink, purple, or blue Inflorescence or individual flowers nodding Stamens and/or style greatly exserted Petals reflexed or recurved Flowers pale blue, raceme Pale Bellflower (Campanula scouleri) p. 102 Flowers bright blue, panicle California Harebell (Campanulaprenanthoides) p. 100 Petals straight, not opening completely White Waterleaf (Hydrophyllum fendleri) p. 134 Stamens and/or style not exserted Small Flower Nemophila (Nemophila parviflora) p. 134 Inflorescence or individual flowers erect Petals free Oregon Checkerbloom (Sidalcea oregana) p. 156 Petals fused Heads radiate Chicory (Cichorium intybus) p.70 Heads ligulate American Sawwort (Saussurea americana) p.84 534 Radial Flowers, 5 Petals, Alternate Leaves, Leaves Compound Leaves pinnately compound Inflorescence more or less flat-topped Inflorescence a corymb (of heads) Yarrow (Achillea millefolium) p.60 Inflorescence a simple or compound umbel (Carrot Family) Plant stout, stem about 12 mm diameter or larger Stem smooth to the touch, glabrous Angelica (Angelica arguta) p.52 Stem rough to the touch, short stififhairs Swamp White Heads (Sphenosciatum capitellatum) p.58/ 28 The Key System a n d How It Works Plant not stout, stem 6 mm diameter or less Plant smells like anise or licorice when crushed Flower bright yellow Western Sweet Root (Osmorhiza occidentalism p.56 Flower white or cream Western Sweet Cicely (Osmorhiza chilensis) p.56 Plant does not smell like anise or licorice Leaflets short, lobed Gray's Lovage (Ligusticum grayi) p.54 Leaflets long, narrow, almost thread-like Yampah (Perideridia oregana) p.58 Inflorescence rounded, not flat-topped Stamens greater than 5, usually 10 or more Flowers bright yellow or pale yellow Leaves glabrous Bigleaf Avens (Geum macrophyllum) p.206 Leaves hairy Sticky Cinquefoil {Potentilla glandulosa) p.208 Flowers white, pink, green, or purplish Leaves silky with long Henderson's Horkelia (Horkelia hendersonii) p.206 Leaves glabrous or short-hairy, but not silky Flowers white Baneberry (Acteae rubra) p. 192 Flowers green or purplish Fendler's Meadow Rue (Thalictrum fendleri) p.202 Stamens 5 or less, or stamens not obvious Inflorescence an umbel, simple or compound (Carrot Family) Flowers yellow Leaves solid green California Lomatium (Lomatium californicum) p.54 Leaves with purple coloring Northern Sanicle (Sanicula graveolens) p.58 Flowers white or cream Anthers purple Piper's Desert Parsley (Lomatiumpiperi) p.54 Anthers not purple California Indian Potato (Orogenia fusiformis) p.56 Inflorescence not an umbel Leaflets entire Flowers blue or lavender Showy Jacob's Ladder (Polemonium californicum) p. 180 Flowers red or orange Scarlet Gilia (Ipomopsis aggregata) p. 176 Leaflets toothed or lobed Inflorescence or individual flowers nodding Flowers with both stamens and pistil White Waterleaf (Hydrophyllum fendleri) p. 134 Flowers with only stamens or pistils Fendler's Meadow Rue (Thalictrum fendleri) p.202 Inflorescence or individual flowers erect Inflorescence a one-sided cyme, coiled in bud Lowermost leaves with 3 leaflets Changeable Phacelia (Phacelia mutabilis) p. 136 Lowermost leaves with 5-9 leaflets Vari-leaf Phacelia (Phacelia heterophylla) p. 136 Inflorescence a head, not coiled Dusty Maidens (Chaenactis douglasii) p.68 Leaves not pinnately compound Stamens 10 or fewer, or stamens indistinguishable Petals notched or lobed Small-flower Woodland Star (Lithophragmaparviflorum) p.218 Petals entire Leaflets heart-shaped Three-leaf Wood Sorrel (Oxalis trilliifolia) p. 172 Leaflets not heart-shaped Inflorescence an umbel Cow Parsnip (Heracleum lanatum) p.52 Inflorescence an open cyme Richardson's Geranium (Geranium richardsonii) p. 130 Stamens greater than 10 Sepals longer than petals Sticky Cinquefoil (Potentilla glandulosa) p.208 Sepals shorter than petals Sulphur Cinquefoil (Potentilla recta) p.208 Radial Flowers, 5 Petals, Opposite or Whorled Leaves, Leaves Entire Petals tips notched, lobed, or fringed Flowers bright yellow Flowers white, cream, or pale pink Klamath Weed (.Hypericum perforatum) p. 138 29 The Key System a n d How It Works Flowers axillary (and terminal) Sepals green, free Petals 0, 5 green sepals Curled Starwort (Stellaria crispa) p. 110 Petals white, 5 green sepals Common Chickweed (Stellaria media) p. 110 Sepals with red-brown markings, fused Palmer's Catchfly (Silene bernardina) p. 108 Flowers in raceme Uppermost leaves fused, completely encircling stem Miner's Lettuce (Claytoniaperfoliata) p. 188 Uppermost leaves free or, if fused, not completely encircling stem Basal leaves more or less heart-shaped Heart-leaf Spring Beauty (Claytonia cordifolia) p. 186 Basal leaves not heart-shaped Stem and leaves always green; petals striped Candy Flower (Claytonia sibirica) p. 188 Stem and leaves often brownish; petals plain Red-stem Spring Beauty (Claytonia rubra) p. 188 Petals entire Plant woody at base Flowers 1 cm or wider Stamens 10, leaves elliptic Bog Laurel (Kalmiapolifolia) p.l 18 Stamens 5, leaves needle-like Spreading Phlox (Phlox diffusa)p.l7S Flowers narrower than 1 cm Plant smells minty when crushed Coyote Mint (Monardella odoratissima) p. 142 Plant not fragrant Creeping Snowberry (Symphoricarpos mollis) p. 106 Plant not woody Flowers white or cream Sepals hairy Hairs glandular, sticky Slender Phlox (Phlox gracilis) p. 178 Hairs not glandular, sticky Upper leaves glabrous .• Milk Kelloggia (Kelloggia galioides) p.216 Upper leaves hairy Nutlets 4 Back of nutlet rough Pinewoods Cryptantha (Cryptantha simulans) p.92 Back of nutlet smooth, mottled Torrey's Cryptantha (Cryptantha torreyana) p.92 Nutlet 1 Weak-stem Cryptantha (Cryptantha flaccida) p.92 Sepals glabrous Leaves opposite Inflorescence tight cluster of many flowers Ballhead Sandwort (Arenaria congesta) p. 108 Inflorescence open and few-flowered Large-leaf Sandwort (Moehringia macrophylla) p. 108 Leaves whorled Western Starflower (Trientalis latifolia) p. 190 Flowers yellow, purple, or pink Petals reflexed Little Prince's Pine (Chimaphila menziesii) p. 116 Petals not reflexed Flower yellow Tinker's Penny (Hypericum anagalloides) p. 138 Flower purple Red Sand Spurry (Spergularia rubra) p. 110 543 Radial Flowers, 5 Petals, Opposite or Whorled Leaves, Leaves Toothed or Lobed Flowers or inflorescence nodding Plant more or less shrubby, woody at base Leaves not evergreen; stamens 5 Leaves evergreen; stamens 10 Inflorescence sticky; leaf veins not bordered Inflorescence not sticky; leaf veins bordered white Plant not shrubby, not woody Flowers red or orange Flowers white, yellow, pale blue, or pale purple Leaves toothed 30 Creeping Snowberry (Symphoricarpos mollis) p. 106 Pipsissewa (Chimaphila umbellata) p. 116 Little Prince's Pine (Chimaphila menziesii) p. 116 Scarlet Gilia (Ipomopsis aggregata) p. 176 The Key System a n d How It Works Flowers yellow to white Flowers pink Leaves lobed Flowers or inflorescence erect Flowers 19 mm wide or wider Leaves opposite Leaves whorled "lowers 12 mm wide or smaller Plant woody at base Plant not woody Stem square or ridged Stem round Leaves glabrous Leaves short-hairy 544 Richardson's Geranium (Geranium richardsonii) p. 130 Columbia Windflower (Anemone deltoidea) p. 192 Mahala Mat (Ceanothus prostratus) p.202 Nettleleaf Giant Hyssop (Agastache urticifcolia) p. 140 Sitka Valerian (Valeriana sitchensis) p.240 California Valerian (Valeriana califomica) p.238 Radial Flowers, 5 Petals, Opposite or Whorled Leaves, Leaves Compound Leaves opposite Stamens 10 Stamens 3 Leaves whorled Fruiting heads erect Fruiting head size of large marble Fruiting head size of golf ball Fruiting heads nodding 552 Sticky Cinquefoil (Potentilla glandulosa) p.208 Twinflower (Linnaea borealis) p. 102 Small-flower Nemophila (Nemophilaparviflora) p. 134 Richardson's Geranium (Geranium richardsonii) p. 130 California Valerian (Valeriana califomica) p.238 Drummond's Anemone (Anemone drummondii) p. 194 White Pasque Flower (Anemone occidentalis) p. 194 Lyall's Anemone (Anemone lyallii) p. 196 Shrub with Radial Flowers, 5 Petals, Leaves Entire Flowers urn-shaped or bell-shaped, and nodding Shrub erect Flowers 1-4 per inflorescence Leaves thin and tissuey Leaves more or less thick Flowers more than 4 per inflorescence Inflorescence terminal Inflorescence axillary Shrub low-lying, creeping Inflorescence terminal Inflorescence axillary Flowers not as above Plant erect Petals or petal-like parts green Leaf tips pointed Leaf tips rounded Petals or petal-like parts white, pink or lavender Stamens 10 Stamens 4 Plant low, mat-forming Leaves very narrow, needle-like Leaves wider, elliptic Thinleaf Huckleberry (Vaccinium membranaceum) p.120 Western Blueberry (Vaccinium uliginosum ) p. 122 Greenleaf Manzanita (Arctostaphylospatula) p.l 16 Snowberry (Symphoricarpos albus) p. 106 Pinemat Manzanita (Arctostaphylos nevadensis) p. 114 Creeping Snowberry (Symphoricarpos mollis) p. 106 Golden Chinquapin (Chrysolepis chrysophyllus) p. 128 Bush Chinquapin (Chrysolepis sempervirens) p. 128 Bog Laurel (Kalmiapolifolia) p.l 18 Coyote Mint (Monardella odoratissima) p. 142 Spreading Phlox (Phlox diffusa) p. 178 Davidson's Penstemon (Penstemon davidsonii) p.234 31 The Key System a n d How It Works 553 Shrub with Radial Flowers, 5 Petals, Leaves Toothed or Lobed Flowers urn-shaped or bell-shaped Leaves opposite Flowers in pairs Flowers in bunches of 8 or more Shrub erect Shrub creeping Leaves alternate Shrub spiny and/or bristly Sepals green or purple, petals purple Sepals red and reflexed, petals white Shrub without spines Leaves small, less than 3 cm wide Bract at base of flowers lobed Bract at base of flowers entire Leaves large, 6 cm wide or more Flowers not urn-shaped or bell-shaped Stamens 5 Leaves evergreen Leaves alternate, finely toothed Leaves opposite, coarsely toothed Leaves deciduous Inflorescence dome-shaped Inflorescence flat-topped Stamens 10 or more Petals reflexed, flowers nodding Petals straight, flowers erect Flowers 2 cm or wider Flowers less than 2 cm in width Sepals green Sepals cream to pale yellow 554 Wax Currant {Ribes cereum) p. 130 Red Flowering Currant {Ribes sanguineum) p. 132 Sticky Currant {Ribes viscosissimum) p. 132 Tobacco Brush {Ceanothus velutinus) p.202 Mahala Mat {Ceanothus prostrates) p.202 Red Elderberry {Sambucus racemosa) p. 106 Blue Elderberry {Sambucus mexicana) p. 104 Little Prince's Pine {Chimaphila menziesii) p. 116 Serviceberry {Amelanchier alnifolia) p.204 Bitter Cherry {Prunus emarginata) p.210 Ocean Spray {Holodiscus discolor) p.208 Wood Rose {Rosa gymnocarpa) p.210 Black-cap Raspberry {Rubus leucodermis) p.212 Pennsylvania Blackberry {Rubus pensilvanicus) p.212 Blue Elderberry {Sambucus mexicana) p. 104 Red Elderberry {Sambucus racemosa) p. 106 Cascade Mountain Ash {Sorbus scopulina) p.214 Rough Fruit Berry {Rubus lasiococcus) p.210 Creeping Snowberry {Symphoricarpos mollis) p. 106 Orange Honeysuckle {Lonicera ciliosa) p. 104 Vine with Radial Flowers, 5 Petals, Leaves Toothed or Lobed Flowers pinkish-white, nodding 32 Swamp Currant {Ribes lacustre) p. 130 Gummy Gooseberry {Ribes lobbii) p. 132 Vine with Radial Flowers, 5 Petals, Leaves Entire Flowers bell-shaped, pinkish-white Flowers trumpet-shaped, orange 563 Snowberry (Symphoricarpos albus) p. 106 Creeping Snowberry {Symphoricarpos mollis) p. 106 Shrub with Radial Flowers, 5 Petals, Leaves Compound Plant thorny or prickly Leaves pinnately compound Leaves palmately compound Leaflets 3 Leaflets 5 Plant without thorns or prickles Leaves opposite Inflorescence flat-topped Inflorescence dome-shaped Leaves alternate Leaflets 9-13 Leaflets 3 562 Twinflower {Linnaea borealis) p. 102 Creeping Snowberry {Symphoricarpos mollis) p. 106 The Key System and How It Works RADIAL FLOWERS WITH 6 PETALS OR PETAL-LIKE PARTS 511 Radial Flowers, 6 Petals, No Apparent Leaves Lower lip of flower white with purple spots Lower lip of flower not clearly spotted Sepals and upper 2 petals clearly striped Sepals and upper 2 petals not clearly striped 622 Glacier Lily (Erythronium grandiflorum) p. 146 .....Jayne's Canyon Buckwheat (Eriogonum diclinum) p.180 Sulfur Flower (Eriogonum umbellatum) p. 182 Naked Buckwheat (Eriogonum nudum) p. 180 Western Tofieldia (Tofieldia occidentalis) p. 150 Pacific Onion (Allium validum) p. 142 White Marsh Marigold (Caltha leptosepala) p. 196 Bride's Bonnet (Clintonia uniflora) p. 144 Sheep Sorrel (Rumex acetosella) p. 184 Blue-eyed Grass (Sisyrinchium bellum) p.140 Radial Flowers, 6 Petals, Basal Leaves Only, Leaves Toothed or Lobed Flowers red Flowers white or yellow Leaves rounded, as wide as long.... Leaves dandelion-like, at least 3x longer than wide 632 Striped Coralroot (Corallorhiza striata) p. 164 Western Coralroot (Corallorhiza mertensiana) p. 164 Radial Flowers, 6 Petals, Basal Leaves Only, Leaves Entire Flowers yellow Basal leaves two, large, tulip-like Basal leaves many, small Flowering stem 11 cm or shorter Flowering stem 12 cm or taller Bracts whorled beneath umbel Bracts not present Flowers white, pink, purple, blue, or red Petals white or pale pink Inflorescence dense head-like or spike-like Stem sticky Stem not sticky Inflorescence solitary flower Stamens 10 or more, pistils 5 or more Stamens 6, pistil 1 Petals red, purple, or blue Flowers many, small, < 5 mm wide. Flowers few, large, >15 mm wide 623 Spotted Coralroot (Corallorhiza maculata) p. 162 Sheep Sorrel (Rumex acetosella) p. 184 White Marsh Marigold (Caltha leptosepala) p. 196 Naked-stem Hawksbeard (Crepis pleurocarpa) p.72 Radial Flowers, 6 Petals, Alternate Leaves, Leaves Entire Plant green and photosynthetic Flowers yellow or bright blue-purple Flowers yellow Petal tips shallowly lobed or notched Leaves sticky Leaves not sticky Petal tips entire Flowers more or less 5mm wide Flowers more or less 10 mm wide Flowers bright blue-purple Flowers white, green, or pale pink Stem stout; plant 60 cm or taller Leaves wide and pleated Flowers white Ovary glabrous; petals mostly entire Greene's Goldenbush (Ericameria greenei) p.72 Naked-stem Hawksbeard (Crepispleurocarpa) p.72 Naked Buckwheat (Eriogonum nudum) p. 180 Plantain-leaf Buttercup (Ranunculus alismifolius) p.200 Blue-eyed Grass (Sisyrinchium bellum) p. 140 California False Hellebore (Veratrum californicum) p. 152 33 The Key System and How It Works Ovary hairy; petals fringed Flowers green Leaves more or less narrow and not pleated Stem slender; plant less than 60 cm tall Petals long, 3 cm or longer Petals short, 15 mm or shorter Flowers bell-shaped, nodding Flowers fully open, erect Inflorescence dense, many-flowered Inflorescence loose, few-flowered Plant not green Lower lip of flower white with purple spots Lower lip of flower not clearly spotted Sepals and upper 2 petals clearly striped Sepals and upper 2 petals not clearly striped 633 Siskiyou False Hellebore (Veratrum insolitum) p. 152 Green False Hellebore (Veratrum viride) p. 152 Willow Dock (Rumex salicifolius) p. 186 Yellow-flowered Iris (Iris chrysophylla) p.140 Hooker's Fairybells (Disporum hookeri) p. 144 False Solomon's Seal (Smilacina racemosa) p. 148 Starry False Solomon's Seal (Smilacina stellata) p. 150 Spotted Coralroot (Corallorhiza maculata) p. 162 Striped Coralroot (Corallorhiza striata) p. 164 Western Coralroot (Corallorhiza mertensiana) p. 164 Radial Flowers, 6 Petals, Alternate Leaves, Leaves Toothed or Lobed Flowers greenish 634 Willow Dock (Rumex salicifolius) p. 186 Radial Flowers, 6 Petals, Alternate Leaves, Leaves Compound Inflorescence wide, flat-topped Inflorescence narrow, dome-shaped 642 Radial Flowers, 6 Petals, Opposite or Whorled Leaves, Leaves Entire Flowers large, >25 mm wide Petals recurved Petals yellow with purple or red dots Petals white or pink with magenta dots Petals straight, purplish-brown Flowers small, <15 mm wide Flowers white Flowers yellow 644 Drummond's Anemone (Anemone drummondii) p. 194 White Pasque Flower (Anemone occidentalis) p. 194 Lyall's Anemone (Anemone lyallii) p. 196 Golden Chinquapin (Chrysolepis chrysophyllus) p. 128 Bush Chinquapin (Chrysolepis sempervirens) p. 128 Sulfur Flower (Eriogonum umbellatum) p. 182 Shurb with Radial Flowers, 6 Petals, Leaves Toothed or Lobed Flowers yellow, leaves holly-like 34 Western Starflower (Trientalis latifolia) p. 190 Sulfur Flower (Eriogonum umbellatum) p. 182 Shrub with Radial Flowers, 6 Petals, Leaves Entire Flowers green Leaf tips pointed Leaf tips rounded Flowers yellow 653 Wiggin's Lily (Lilium pardalinum) p. 146 Washington Lily (Lilium washingtonianum) p. 148 Chocolate Lily (Fritillaria atropurpurea) p. 146 Radial Flowers, 6 Petals, Opposite or Whorled Leaves, Leaves Compound Fruiting heads erect Fruiting head size of large marble Fruiting head size of golf ball Fruiting heads nodding 652 Yarrow (Achillea millefolium) p.60 Baneberry (Acteae rubra) p. 192 Oregon Grape (Berberis nervosa) p.90 The Key System and How It Works RADIAL FLOWERS WITH 7 OR MORE PETALS OR PETAL-LIKE PARTS 711 Radial Flowers, 7 or More Petals, No Apparent Leaves Flowers yellow, dandelion-like 722 Radial Flowers, 7 or More Petals, Basal Leaves Only, Leaves Entire Petals white Petals orange or yellow Petals orange Phyllaries (sepal-like structures) glabrous Phyllaries (sepal-like structures) hairy Petals yellow Leaves glabrous Phyllaries (sepal-like structures) soft-hairy Phyllaries mottled with pink Phyllaries pale green Phyllaries (sepal-like structures) glabrous Leaves hairy Leaf hairs bristly Leaf hairs soft Leaf hairs spreading Leaf hairs tightly appressed, silky 723 Scouler's Hawkweed (Hieracium scouleri) p.80 White Marsh Marigold (Caltha leptosepala) p. 196 Orange Agoseris (Agoseris aurantiaca) p.62 Tall Agoseris (Agoseris elata) p.62 Large-flowered Agoseris (Agoseris grandiflora) p.64 Pale Agoseris (Agoseris glauca) p.66 Alpine Lake Prairie Dandelion (Nothocalais alpestris) p.84 Hairy Cat's Ear (Hypochaeris radicata) p.80 Scouler's Hawkweed (Hieracium scouleri) p.80 Silky Raillardella (.Raillardella argentea) p.86 Radial Flowers, 7 or More Petals, Basal Leaves Only, Leaves Toothed or Lobed Petals white White Marsh Marigold (Caltha leptosepala) Petals orange or yellow Petals orange Phyllaries (sepal-like structures) glabrous Orange Agoseris (Agoseris aurantiaca) Phyllaries (sepal-like structures) hairy Tall Agoseris (Agoseris elata) Petals yellow Stem simple, flowers solitary Phyllaries (sepal-like structures) hairy Phyllaries mottled with pink Large-flowered Agoseris (Agoseris grandiflora) Phyllaries pale green Pale Agoseris (Agoseris glauca) Phyllaries (sepal-like structures) glabrous Outer phyllaries straight, purple-dotted...Alpine Lake Prairie Dandelion (Nothocalais alpestris) Outer phyllaries relfexed Dandelion (Taraxacum officinale) Stem branched, more than one flower per plant Leaves glabrous Naked-Stem Hawksbeard (Crepis pleurocarpa) Leaves bristly-hairy Hairy Cat's Ear (Hypochaeris radicata) 732 p. 196 p.62 p.62 p.64 p.66 p.84 p.90 p.72 p.80 Radial Flowers, 7 or More Petals, Alternate Leaves, Leaves Entire Petals yellow Flowers daisy-like Plant of wet places Plant with black glands, very sticky Plant without glands, not sticky Stem stout, winged Stem slender, not winged Plant of dry places Bolander's Madia (Madia bolanderi) p.82 Tall Sneezeweed (Helenium bigelovii) p.78 Plantain-lead Buttercup (Ranunculus alismifolius) p.200 35 The Key System a n d How It Works Leaves woolly and more or less white Oregon Sunshine (Eriophyllum lanatum) p.76 Leaves glabrous, green Inflorescence more or less flat-topped Western Butterweed (Senecio integerrimus) p.86 Inflorescence more or less diamond-shaped Canada Goldenrod (Solidago canadensis) p.88 Flowers dandelion-like Leaves and phyllaries (sepal-like parts) hairy Flowers bright yellow Phyllaries densely hairy Scouler's Hawkweed (Hieracium scouleri) p.80 Phyllaries sparsely hairy Lower surface of outer petals brownish-red Hairy Cat's Ear (.Hypochaeris radicata) p.80 Lower surface of outer petals yellow Naked-stem Hawksbeard (Crepis pleurocarpa) p.72 Flowers pale yellow or cream White Hawkweed (Hieracium albiflorum) p.78 Leaves and phyllaries glabrous Phyllaries recurved, very sticky Idaho Gum Weed (Grindelia nana) p.76 Phyllaries straight Flowers many per plant (20+) Canada Goldenrod (Solidago canadensis) p.88 Flowers few per plant (1-3) Yellow Salsify (Tragopogon dubius) p.88 Petals white, cream, pink, purple, or blue Flower daisy-like Leaf margins wavy; phyllaries (sepal-like parts) not leafy Branched at mid-stem, 1-7 flowers Alice's Daisy (Erigeron alicea) p.72 Branched at top of stem, 1-4 flowers Subalpine Fleabane (Erigeronperegrinus) p.74 Leaf margins straight, phyllaries leafy Leafy Aster (Asterfoliaceus) p.68 Flower dandelion-like Flowers white or cream White Hawkweed (Hieracium albiflorum) p.78 Flowers purple American Sawwort (Saussurea americana) p.84 733 Radial Flowers, 7 or More Petals, Alternate Leaves, Leaves Toothed or Lobed Flowers yellow Stems and phyllaries (sepal-like parts) very sticky Phyllaries straight with black oil glands Phyllaries recurved with no obvious glands Stems and phyllaries not sticky Leaves and stem full of white woolly hairs Leaves and stem glabrous, or if hairy, not woolly hairs Leaves more or less triangular Leaves linear to lanceolate Flowers purple or blue Leaves and stems spiny Leaves and stems without spines Leaves deeply lobed Leaves finely toothed 734 Arrowhead Butterweed (Senecio triangularis) p.88 Western Butterweed (Senecio integerrimus) p.86 Bull Thistle (Cirsium vulgare) p.70 Chicory (Cichorium intybus) p.70 American Sawwort (Saussurea americana) p.84 Yarrow (Achillea millefolium) p.60 Baneberry (Acteae rubra) p. 192 Radial Flowers, 7 or More Petals, Opposite or Whorled Leaves, Leaves Entire Leaves whorled Leaves opposite Flowers yellow Plant hairy (glandular or not) 36 Oregon Sunshine (Eriophyllum lanatum) p.76 Radial Flowers, 7 or More Petals, Alternate Leaves, Leaves Compound Inflorescence wide, flat-topped Inflorescence narrow, dome-shaped 742 Bolander's Madia (Madia bolanderi) p.82 Idaho Gum Weed (Grindelia nana) p.76 Western Starflower (Trientalis latifolia) p. 190 The Key System and How It Works Plant sticky at top Plant not sticky, stem and leaves white with hairs Plant glabrous Flowers white 743 Radial Flowers, 7 or More Petals, Opposite or Whorled Leaves, Leaves Toothed or Lobed Plant hairy (glandular or not) Plant sticky at top Plant not sticky, stem and leaves white with hairs Plant glabrous 744 Bolander's Madia (Madia bolanderi) p.82 Oregon Sunshine (Eriophyllum lanatum) p.76 Heartleaf Arnica (Arnica cordifolia) p.66 Radial Flowers, 7 or More Petals, Opposite or Whorled Leaves, Leaves Compound Fruiting heads erect Fruiting head size of large marble Fruiting head size of golf ball Fruiting heads nodding, petal tips rounded 752 Bolander's Madia (Madia bolanderi) p.82 Oregon Sunshine (Eriophyllum lanatum) p.76 Heartleaf Arnica (Arnica cordifolia) p.66 Common Chickweed (Stellaria media) p.l 10 Drummond's Anemone (Anemone drummondii) p. 194 White Pasque Flower (Anemone occidentalis) p.194 Lyall's Anemone (Anemone lyallii) p. 196 Shrub, Radial Flowers, 7 or More Petals, Leaves Entire Leaf tips pointed Leaf tips rounded Golden Chinquapin (Chrysolepis chrysophyllus) p. 128 Bush Chinquapin (Chrysolepis sempervirens) p. 128 FLOWER PARTS INDISTINGUISHABLE 822 Flower Parts Indistinguishable, Basal Leaves Only, Leaves Entire Stamens exerted Flowers creamy white or pink with red anthers Flowers green or colorless with white anthers Stamens not exserted 823 Pussy Paws (Calyptridium umbellatum) p. 186 English Plantain (Plantago lanceolata) p. 174 Scabland Fleabane (Erigeron bloomeri) p.74 Flower Parts Indistinguishable, Basal Leaves Only, Leaves Toothed or Lobed Flowers yellow Flowers white 824 Flower Parts Indistinguishable, Basal Leaves Only, Leaves compound Flowers yellow 832 Cut-leaf Daisy (Erigeron compositus) p.74 Trail Plant (Adenocaulon bicolor) p.62 Cut-leaf Daisy (Erigeron compositus) p.74 Flower Parts Indistinguishable, Alternate Leaves, Leaves Entire Leaves glabrous Flowers brown, purple, red, or orange Flowering head solitary, subtended by whorl of bracts Western Coneflower (Rudbeckia occidentalis) Flowering heads several, not subtended by whorl of bracts Flowers purple, stem green American Sawwort (Saussurea americana) Flower red-orange, stem brown-red Giant Red Indian Paintbrush (Castilleja miniata) Flowers white, cream, pink, or yellow Plant prostrate Pussy Paws (Calyptridium umbellatum) Plant erect p.84 p.84 p.224 p. 186 37 The Key System and How It Works Flowers yellow Flowers pink and white Leaves hairy Leaves and stems white, gray, or frosted from dense hairs Flowers white, yellow, or pink Plant strongly scented Plant not scented or slightly scented Leaves erect to appressed Leaves widely spreading Flowers red or orange Leaves and stems green with sparse hairs Plant tall, at least 60 cm Flowers brown Flowers purple Plant short, 15 cm or shorter 833 Scabland Fleabane (Erigeron bloomeri) p.74 Mountain Owl's Clover (Orthocarpus imbricatus) p.232 Wright's Cudweed (Gnaphalium canescens) p.76 Rosy Pussytoes (Antennaria rosea) p.64 Pearly Everlasting (Anaphalis margariticea) p.64 Frosted Indian Paintbrush (Castillejapruinosa) p.224 ... Western Coneflower (Rudbeckia occidentalis) p. 84 American Sawwort (Saussurea americana) p.84 Cut-leaf Daisy (Erigeron compositus) p.74 Flower Parts Indistinguishable, Alternate Leaves, Leaves Toothed or Lobed Flowers yellow Plants aromatic Plant woody at base Mountain Sagebrush (Artemisia tridentata) p.68 Plant not woody Plant pineapple scented, flower cone-shaped Pineapple Weed (Chamomilla suaveolens) p.70 Plant not pineapple scented, flower not cone-shaped Mugwort (Artemisia douglasiana) p.66 Plants not aromatic Leaves lobed Cobwebby Paintbrush (Castilleja arachnoidea) p.224 Leaves toothed Leaves narrow, elliptic to linear Shasta Tansy Aster (Machaeranthera canescens) p.80 Leaves wide, ovate Whitney's Bristleweed (Hazardia whitneyi) p.78 Flowers red, orange, purple, pink, or white Leaves lobed Leaves and stems thorny Bull Thistle (Cirsium vulgare) p.70 Leaves and stems not thorny Flowers red-orange Frosted Indian Paintbrush (Castillejapruinosa) p.224 Flower white and/or pink Leaves wide, triangular Trail Plant (Adenocaulon bicolor) p.62 Leaves narrow, 3-lobed Siskiyou Mountains Owl's Clover (Orthocarpus cuspidatus) p.230 Leaves toothed Flowers brown Western Coneflower (Rudbeckia occidentalis) p.84 Flowers purple American Sawwort (Saussurea americana) p.84 834 Flower Parts Indistinguishable, Alternate Leaves, Leaves Compound Flowers yellow Plant aromatic (smells like pineapple) Plant not aromatic Flowers green 843 Flower Parts Indistinguishable, Opposite or Whorled Leaves, Leaves Toothed or Lobed Flowers small and green, plant with stinging hairs 38 Pineapple Weed (Chamomilla suaveolens) p.70 Cobwebby Paintbrush (Castilleja arachnoidea) p.224 Western Burnet (Sanguisorba occidentalis) p.214 Stinging Nettle (Urtica dioica) p.238 The Key System and How It Works 852 Shrubs with Flower Parts Indistinguishable, Leaves Entire Leaf tips pointed Leaf tips rounded 853 Golden Chinquapin (Chrysolepis chrysophyllus) p. 128 Bush Chinquapin (Chrysolepis sempervirens) p. 128 Shrubs with Flower Parts Indistinguishable, Leaves Toothed or Lobed Flowers fixzzv white Willow (Salix eastwoodiae) i, not fuzzy or wider or narrower p.218 Mountain Maple (Acer glabrum) p.52 Mountain Sagebrush (Artemisia tridentata) p.68 39 FART 3 Family Descriptions and Flower Descriptions "I will be the gladdest thing under the sun! I will touch a hundred flowers and not pick Edna St. Vincent Millay Afternoon on a Mill (1917) Family Descriptions FAMILY DESCRIPTIONS Maple Family (Aceraceae) Numbers 2 genera and 120 species Distribution mostly northern temperate areas and tropical mountains Plant Types shrubs and trees Leaves opposite, simple, generally palmately lobed Flowers small and radial, sepals 4 or 5, petals 1-6 but usually 5, stamens 5-12 but usually 8; pistil 1, ovary superior; some bisexual, some unisexual Fruits achenes. Mountain Maple, a member of the Maple Family Carrot or Parsley Family (Apiaceae) Numbers 300 genera, 3000 species Distribution worldwide, especially in temperate areas Leaves usually compound, often finely cut, sheathed at base Stems hollow and often ridged Inflorescence simple or compound umbels Flowers small, sepals 0 or 5, petals 5, stamens 5, pistil 2-parted Fruits schizocarp – splits into two parts Interesting family members include carrots, celery, parsley, anise, dill, and fennel. Birthwort Family (Aristolochiaceae) Etymology Greek; aristos means best, lochia means delivery; wort is old English word for plant; Plants in this family thought to be of use during child birth. Numbers 10 genera, 600 species Distribution worldwide, mostly tropical and warm temperate areas Leaves simple, usually reniform or cordate, alternate or basal, stem and/or leaves aromatic from oils Flowers reddish-brown, no petals, 3 showy sepals that are fused; often fetid odor to attract pollinators (flies, fungal gnats, beetles), compound pistil 6-chambered Fruits many-seeded capsules Wild Ginger, a member of the Birthwort Family 41 Family Descriptions Sunflower Family (Asteraceae) Numbers 1300 genera, 21000+ species, largest family in the world Distribution worldwide Inflorescence head – often mistaken for one large flower Flowers many small flowers on the head; sometimes of two forms – disk flowers (tube shaped) in the center of the head and ligulate or ray flowers around the edge of the head. Head can have only disk flowers, only ray flowers, or both. All flowers are attached to the receptacle. Individual flowers have 5 petals fused into a tube or a strap-shaped ray, 5 stamens fused into a tube around 1 pistil. Sepals are modified into hairs collectively called pappus. Many bracts subtend the head of flowers; these bracts are collectively referred to as the involucre. Fruits achene Interesting plants in this family include lettuce, artichoke, endive, sunflower, aster, chrysanthemums, coreopsis, marigolds, and zinnia. Disk Flower Ray Flower Receptacle Involucral Bracts Barberry Family (Berberidaceae) Numbers 16 genera, 670 species Distribution worldwide, mostly temperate and tropical Leaves usually alternate or basal, simple or compound Flowers 4-many sepals, 6-9 petals, many stamens, 1 simple pistil, superior ovary Fruits berry or capsule Borage Family (Boraginaceae) Numbers 100 genera 2000 species Distribution mostly tropical and temperate, especially in western North America and the Mediterranean Leaves simple, alternate, entire, typically rough- or stiff-hairy Stems rough- or stiff-hairy Inflorescence usually tight one-sided coiled cyme that uncoils as the flowers open Flowers radial, 5 sepals, 5 petals, 5 stamens, 1 pistil, superior ovary, petals have modified glands called corona Fruits nutlets, usually 4 Interesting plants in this family include forget-me-nots, borage, hound’s tongue, and comfrey. 42 Family Descriptions Mustard Family (Brassicaceae) This family was formerly known as Cruciferae because the four petals are in the shape of a Maltese cross. Numbers 376 genera, 3200 species; in U.S./Canada 94 genera, 634 species Distribution worldwide especially in cool regions Leaves usually alternate, if basal leaves are present they are often different from stem leaves, watery/acrid sap Flowers 4 sepals, 4 petals in cross shape, usually 6 stamens (4 tall, 2 short), 1 pistil 2parted; superior ovary Fruits siliques, silicles Interesting plants in this family include cabbage, turnips, broccoli, and brussel sprouts. Bellflower Family (Campanulaceae) Numbers 70 genera 2000 species Distribution worldwide Leaves generally simple and alternate Flowers radial or bilateral, sepals 5, petals 5, stamens 5, pistil 1, ovary inferior Fruits capsule Honeysuckle Family (Caprifoliaceae) Numbers 12 genera 450 species Distribution mostly in northern temperate areas Plant Types shrubs, vines, and small trees Leaves opposite, simple or compound Flowers radial or bilateral, 5 sepals fused, 5 petals fused at base, stamens 5, pistil 1, ovary inferior Fruits berry, drupe, or capsule Pink Family (Caryophyllaceae) This common family name is not for the color pink, but for the usually-notched petals, as in pinking shears. Numbers 85 genera, 2400 species Distribution worldwide, mostly in arctic, alpine, and temperate areas in the northern hemisphere Leaves usually simple, opposite, sometimes whorled Stems often swollen at nodes (think carnations, which are in this family) Flowers 5 sepals free or fused, 5 petals, 5-10 stamens, compound pistil with 2-5 parts Fruits capsule usually opening at the top with recurved teeth Paxistima sp., a member of the Staff-Tree Family Staff-Tree Family (Celastraceae) Numbers 50 genera, 800 species Distribution worldwide, especially in southeast Asia Plant Types shrubs and trees Leaves simple, alternate or opposite Flowers radial, small, sepals 4-5, petals generally 4-5, stamens 4-5, pistil 1, ovary superior Fruits capsule, achene, berry, drupe, or nutlet 43 Family Descriptions Stonecrop Family (Crassulaceae) Numbers 30 genera, 1500 species Distribution worldwide, especially dry temperate areas Leaves succulent or fleshy, opposite, alternate or whorled, usually simple Flowers radial, 4-30 sepals, petals in the same number as sepals, ovaries usually as many as sepals, superior ovaries Fruits 1-chambered follicles Heath or Wintergreen Family (Ericaceae) Numbers 100 genera, 3000 species Distribution worldwide, but not found in deserts Plant Types Herbaceous or woody-based perennials Leaves simple, alternate, opposite or whorled, vary from evergreen to reduced and lacking chlorophyll Flowers radial or slightly bilateral, usually 4-5 sepals, usually 4-5 petals, 8 or 10 stamens, 1 pistil, superior ovary Fruits 4-5 chambered capsule Legume or Pea Family (Fabaceae) Numbers 650 genera 18000 species; US/Canada 142 genera, 1521 species; 3rd largest family in the world after Sunflower Family (Asteraceae), and Orchid Family (Orchidaceae) Distribution worldwide Plant Types trees, vines, and herbaceous species Leaves compound, usually alternate, usually stipules, sometimes with tendrils or thorns Flowers usually bilateral, often tiny and clustered on heads, 5 sepals, 5 petals, 10+ stamens, 1 simple pistil Fruits legume This family is economically important providing food (peas, peanuts, lentils, beans, soybeans), forage and fodder (clover, alfalfa), dyes, gums, resins, oils, medicines and timbers. Oak Family (Fagaceae) Numbers 7 genera, 900 species Distribution mostly in northern hemisphere Plant Types shrubs and trees Leaves simple, alternate, entire or lobed, small stipules Flowers unisexual; staminate flowers arranged in catkin, 5-6 sepals, no petals, 4-12+ stamens; pistillate flowers situated above the catkin, 6 sepals, no petals, 1 pistil, inferior ovary Fruits acorn or nuts surrounded by spiny bracts Species in this family provide valuable wood, cork, and food products. 44 Geranium or Cranesbill Family (Geraniaceae) Etymology Geranos is Greek for crane Numbers 14 genera, 775 species; U.S./Canada 3 genera, 62 species Distribution worldwide Leaves deeply palmately lobed, usually with 5 lobes, alternate Flowers 5 sepals, 5 petals, 5-15 stamens, 5-parted pistil Fruits schizocarp – breaks into 5 parts Gooseberry Family (Grossulariaceae) Erodium cicutarium fruits that look Numbers 1 genus, 120 species like cranes’ bills. Distribution primarily in northern hemisphere and temperate south America Plant Types shrubs Leaves simple, alternate, generally lobed Flowers radial, sepals 5, petals5 , stamens5, 1 pistil, ovary inferior Fruits berry This family was formerly included in the Saxifrage family (Saxifragaceae). © University of California Press Family Descriptions Waterleaf Family (Hydrophyllaceae) Numbers 20 genera, 300 species Distribution mostly in western U.S. Leaves alternate, opposite or basal rosettes, entire or pinnately divided Inflorescence usually one-sided coiled cyme Flowers radial, 5 sepals partially fused, 5 petals partially fused into bell shape, 5 stamens, 1 pistil, superior ovary Fruits 1 chambered capsule St. John’s Wort Family (Hypericaceae) Numbers 10 genera, 400 species Distribution worldwide Leaves simple, opposite or whorled, often with black dots Flowers radial, sepals 5 often fused at base, petals 5, stamens many, pistil 1, ovary superior Fruits capsule Iris Family (Iridaceae) Numbers 80 genera, 1500 species Distribution worldwide, especially in Africa Leaves basal, overlapping in 2 ranks, linear to lanceolate, entire with parallel veins Flowers radial or bilateral, 6 petal-like parts in 2 series, all generally fused at base, 3 stamens, 3-parted pistil, inferior ovary Fruits 3 chambered capsule Familiar plants in this family include iris, gladiolus, crocus and freesia. 45 Family Descriptions Mint Family (Lamiaceae) Numbers 200 genera, 5500 species Distribution worldwide Stems square Leaves aromatic, opposite Flowers usually bilateral, 5 fused sepals, 5 petals fused into a two-lipped flower, 2 or 4 stamens, 4-parted pistil, ovary superior Fruits nutlets, usually 4 This family is an important source of aromatic oils (lavender, rosemary, mint) and herbs (basil, thyme, and savory). Lily Family (Liliaceae) Numbers 300 genera, 4600 species Distribution mostly in dry temperate and subtropical areas Leaves basal, alternate or whorled, entire with parallel veins Flowers radial; generally 3 sepals and 3 petals that look the same (tepals), 3 stamens, 1 pistil, superior ovary Fruits 3-chambered capsule or berry Familiar plants in this family include some edibles - onions, garlic, leeks – and some ornamentals - tulips, lilies, hyacinths, hostas. Flax Family (Linaceae) Numbers 13 genera, 300 species Distribution worldwide, especially temperate areas Leaves alternate, opposite or whorled, simple Flowers radial, sepals 4-5, petals 4-5, stamens 4-5, pistil 1 with 2-5 styles, ovary superior Fruits capsule Loasa Family (Loasaceae) Numbers 15 genera, 200 species Distribution mostly in the Americas, Africa, and the Pacific Leaves generally alternate and lobed Flowers radial, sepals 5, petals 5, stamens 5, pistil 1, inferior ovary Fruits capsule (utricle) Mentzelia sp., a member of the Loasa Family Mallow Family (Malvaceae) Numbers 100 genera 2000 species Distribution worldwide, especially in warm regions Leaves simple, alternate, usually with stipules Flowers radial, 5 sepals, 5 petals, many stamens, 1 pistil Fruits capsule or berry 46 Family Descriptions Evening Primrose Family (Onograceae) Numbers 15 genera, 650 species Distribution worldwide, mostly in western North America Leaves simple, alternate or opposite Flowers radial, 4 sepals, 4 petals, 4-8 stamens, 1 pistil, inferior ovary Fruits 4-chambered capsule Orchid Family (Orchidaceae) Numbers 800 genera, 18000 species Distribution worldwide except in deserts, mostly in tropical areas Leaves linear to round, entire, parallel veins Flowers bilateral, sepals 3 and petal-like, petals 3, stamen 1 fused with style and stigma into a column, ovary inferior Fruits Capsule Broom-rape Family (Orobanchaceae) Numbers 14 genera, 200 species Distribution mostly in northern temperate Leaves no true leaves but sometimes scaly bracts are present Flowers bilateral, sepals 0-5 fused into tube, petals generally 5 often fused into tube, stamens 4, pistil 1, ovary superior Oxalis Family (Oxalidaceae) Etymology Greek for sour Numbers 8 genera 575 species; U.S./Canada 1 genera, 30 species Distribution mostly temperate areas Leaves compound with 3 leaflets, usually clover-like, each leaflet has pleat or fold down the center, sour, watery juice Flowers 5 sepals free, 5 petals fused at base, 5-15 stamens, 5parted pistil Fruits capsule, explodes when disturbed Folded Oxalis leaf Poppy Family (Papaveraceae) Numbers 40 genera, 400 species Distribution mostly in north temperate and north tropical areas and south Africa Leaves deeply lobed, alternate, stem or leaf juice is usually colored (often red or yellow, sometimes orange) Flowers showy and large, usually 2 sepals that fall off early, 4-12 petals, many stamens, 1 4-parted pistil Fruits 4-parted capsule 47 Family Descriptions Plantain Family (Plantaginaceae) Numbers 3 genera, 270 species Distribution worldwide, especially temperate Leaves basal, simple with longitudal ribs Flowers unisexual or bisexual, radial, sepals 4 fused at base, petals 4 papery or colorless, stamens 2, pistil 1, ovary superior Psyllium, which is cultivated for a laxative, is in this family. Phlox Family (Polemoniaceae) Numbers 19 genera, 320 species Distribution Americas, north Europe, north Asia Leaves alternate, simple or compound, if simple often entire Flowers radial, 5 sepals fused into a tube, 5 petals fused at the base to form a trumpetshaped flower, 5 stamens, 1 pistil, superior ovary Fruits 3-chambered capsule There are a few ornamental species in this family. Buckwheat Family (Polygonaceae) Numbers 50 genera 1100 species Distribution worldwide, especially in northern temperate areas Leaves alternate, sheathing stipules Stems often with swollen nodes Flowers small, radial, numerous in terminal or axillary clusters, often unisexual, 6 petallike parts in 2 whorls of 3, stamens usually 6-9, 1 simple pistil, superior ovary Fruits achene – flat, angled, or winged Buckwheat and Rhubarb are in this family. Purslane Family (Portulacaceae) Numbers 19 genera, 400 species Distribution worldwide, mostly in warm, temperate regions of the Americas, Australia, and south Africa Leaves fleshy to succulent, usually opposite, entire, simple Flowers radial, 2 sepals, 4-many petals, 4-5+ stamens, compound pistil with 2-8 parts Fruits capsule Primrose Family (Primulaceae) Numbers 25 genera, 600 species Distribution mostly in the northern hemisphere Leaves simple, basal, alternate, opposite or whorled Flowers radial, sepals, petals and stamens in 4s or 5s, 1 pistil, ovary superior Fruits capsule 48 Family Descriptions Buttercup Family (Ranunculaceae) Etymology Latin for Place of the Frog because many species grow in wet places. Numbers 60 genera, 1700 species Distribution worldwide, especially in north temperate areas and tropical mountains Leaves often palmately divided or compound, usually alternate, leaves have sheathed base Flowers 5+ sepals, 0-many petals but usually 5, many stamens, many pistils Fruits achenes, berries, follicles Buckthorn Family (Rhamnaceae) Numbers 55 genera, 900 species Distribution mostly tropical and subtropical Plant Types shrubs, trees, and vines Leaves simple, generally alternate Flowers radial, sepals 4 or 5, petals 0,4 or 5, stamens 4 or 5, pistil 1, superior ovary Fruits capsule, drupe Rose Family (Rosaceae) Numbers 110 genera, 3000 species; U.S./Canada 62 genera, 870 species Distribution worldwide Plant Types trees, vines, and herbaceous species Leaves usually alternate, often compound, usually stipulate, stems often have thorns Flowers radial, 5 sepals (partially fused), usually 5 petals, many stamens, 1 to many simple pistils Fruits hips, pomes, drupes, aggregates, achenes, Familiar plants in this family includes strawberry, blackberry, raspberry, plums, cherries, peaches, apples, and pears. Madder Family (Rubiaceae) Numbers 500 genera, 6000 species Distribution worldwide, especially tropical Leaves entire, opposite or whorled Flowers small, radial, sepals 0 or 4, petals 4, pistil 1 or 2 fused Fruits berry, drupe, capsule, 2-4 nutlets Willow Family (Salicaceae) Numbers 2 genera, 340 species Distribution mostly in temperate areas Plant Types shrubs and trees Leaves simple, alternate, deciduous Flowers unisexual, radial, no sepals, no petals, staminate flowers have 1-many stamens, pistillate flowers have 1 pistil and superior ovary Fruits capsule 49 Family Descriptions Saxifrage Family (Saxifragaceae) Etymology Latin for “rock breaking;” over long periods of time, plants that grow in rocky areas help to break down the rocks Numbers 40 genera, 600 species Distribution mostly in north temperate, arctic, alpine areas Leaves usually alternate or basal forming rosettes, simple, sometimes lobed Flowers usually small and clustered, 5 sepals, 4-5 petals, 5 or 10 stamens, 1 pistil with 2 parts, superior ovary Fruits capsule, follicle, berry Figwort Family (Scrophulariaceae) Numbers 200 genera 3000 species Distribution worldwide Leaves alternate, simple, usually more or less entire Flowers bilateral, usually 2-lipped with 4-5 lobes, number of stamens vary – typically 4 with a 5th infertile stamen in Penstemon, 2 stamens in Veronica, 5 stamens in Verbascum, 1 pistil, ovary superior Fruits usually 2-chambered capsule Foxgloves (Digitalis) are in this family. Nettle Family (Urticaceae) Numbers 50 genera, 700 species Distribution worldwide Leaves alternate or opposite, glabrous or with stinging hairs Stems glabrous or with stinging hairs Flowers usually unisexual, radial, sepals 4-5 free or fused, petals 0, staminate flowers with 4-5 stamens, pistillate flowers with 1 pistil, superior ovary Fruits achene or drupe Valerian Family (Valerianaceae) Numbers 17 genera 300 species Distribution worldwide, except in Australia; mostly in temperate areas Leaves simple, pinnately lobed or compound, opposite Flower radial to bilateral, 5-15 sepals coiled inward becoming plumose in fruit, 5 petals usually partially fused often bilabial, 1-3 stamens, 1 pistil, ovary inferior Fruits achene Violet Family (Violaceae) Numbers 15 genera, 600 species; U.S./Canada 3 genera, 106 species Distribution worldwide, mostly in temperate areas Plant Types in U.S., herbaceous species; in tropics, tree and shrub species Leaves Flowers simple, usually cordate, alternate or basal, stipules at base 5 sepals, 5 petals, 5 stamens flattened against the 3-parted pistil, ovary superior; usually bilateral flower with spur; some species are cleistogamous Fruits capsule. Approximately 120 species in this family are cultivated as ornamentals, including pansies. 50 Plant Descriptions In this section, the flower descriptions are listed alphabetically by scientific name according to family, genus, and species. To use this section most effectively, use the key system that starts on page 17 to identify your plant, and then turn to the page(s) on which your species is described. 51 Plant Descriptions Maple Family (Aceraceae) Mountain Maple Acer glabrum Torrey var. torreyi (E. Greene) F.J. Smiley Native. Maple Family (Aceraceae). On Mount Ashland, Mountain Maple is a small shrub. Typical maple-shaped leaves grow opposite each other on slender, reddish twigs. The three primary lobes are irregularly toothed. Drooping inflorescences, with less than 10 greenish flowers each, appear before the leaves do. The flowers have either stamens, or pistils, not both, making these flowers imperfect. The staminate flowers have 5 tiny sepals, 5 tiny petals, and 8 stamens, while the pistillate flowers have 5 sepals, 5 petals, and 1 pistil. Because both male and female flowers grow on the same shrub, Mountain Maple is called dioecious. This short shrub grows in moist to dry slopes in forested areas along the Pacific Crest Trail. You have to be on the mountain early in the season to find it in flower. Acer is the Latin name for Maple. Glabrum is derived from the Latin word for smooth, referring to the absence of hairs. When a plant is not hairy, it is called glabrous. Dr. John Torrey (1796-1879) was a botanist of western North America. He reviewed and described the collections of many other noted botanists, including William Baldwin, and David Douglas. Carrot Family (Apiaceae) Angelica Angelica arguta Nutt. Also called Lyall’s Angelica, and Shining Angelica. Native. Carrot Family (Apiaceae). This stout plant with many white flowers arranged in an umbel of umbels is abundant on the south slopes in July. The dime-sized umbels look like little buttons, and each plant has 1-5 compound umbels made up of tiny radial flowers with 5 petals, 5 stamens, and 1 pistil; there are no sepals. The stem leaves are compound with very obvious sheathing bases, and the leaflets are coarsely toothed all along the margin. The basal leaves of Angelica are a perfect example of a ternate leaf – a compound leaf with 3 major divisions. This plant often grows up to 1 meter, and the stout stem is glabrous with a powdery white substance, called bloom, that rubs off when touched. Look for this tall perennial in wet areas on the south slopes of Mount Ashland. It is often found near Gray’s Lovage (Ligusticum grayi) and Western Columbine (Aquilegia formosa). Angelica is Latin for angelic, in reference to the medicinal properties of some of the plants in this genus. Arguta is Greek for shining or bright. Cow Parsnip Heracleum lanatum Michaux Also called Indian Rhubarb, Cow-cabbage, Masterwort, Hercules Parsnip, and Common Cow Parsnip. Native. Carrot Family (Apiaceae). Many small white flowers with 5 petals, 5 stamens and 1 pistil are borne in several umbels of umbels. Usually, one central umbel is flanked by two smaller umbels. Each flowering stem is subtended by an off-white, balloon-like sheath that tapers to a small, palmately lobed green leaf. The lower leaves are alternate and compound with leaflets that are toothed and hairy. The hollow stems are stout, and ridged, with coarse hairs. This perennial, which usually grows 60-150 cm tall, requires three years to mature from seed. Cow Parsnip is found in all the wet meadows on the south slopes with Big-leaf Lupine (Lupinus polyphyllus var. pallidipes) and Western Columbine (Aquilegia formosa). The sap can cause temporary skin irritations. There is contradictory information regarding the edibility of this plant for animals and humans. Some sources say Cow Parsnip is poisonous to cattle and humans, while other sources claim that wild and domestic animals forage for the leaves in spring, and parts of the plant can be consumed by humans if properly prepared. If you wish to forage plants for food, seek the advice and confirmation of a professional botanist or ethnobotanist. This huge, Herculean plant is named for the Greek hero, Heracles (more familiar by the Roman name of Hercules), who is purported to have found medicinal qualities in parts of the plant. Lanatum is Latin for wooly because the plant is so hairy. 52 Plant Descriptions Mountain Maple (Acer glabrum var. torreyi) Angelica (Angelica arguta) Cow Parsnip (Heracleum lanatum) 53 Plant Descriptions Gray’s Lovage Ligusticum grayii J. Coulter & Rose Also called Gray’s Licorice-root. Native. Carrot Family (Apiaceae). Tiny white radial flowers with 5 minute sepals, 5 petals, 5 stamens and 1 pistil are borne in an umbel of umbels, with each plant producing several compound umbels. Most plants have only one compound stem leaf with an enlarged base that clasps the stem. Many compound basal leaves surround the base of the stem. The leaves are twice pinnate, and the leaflets are coarsely and irregularly lobed. The glabrous flowering stem, which is often mottled reddish-brown, rises high above the basal leaves reaching heights of 20-80 cm. This perennial requires the moist soil near wet meadows and in partly shaded areas. Nearby, you may find Wiggin’s Lily (Lilium pardalinum ssp. wigginsii) or Angelica (Angelica arguta). Dr. Asa Gray (1810-1888) is honored in both the common name and the specific epithet for this plant. Dr. Gray and his teacher, Dr. John Torrey, published A Flora of North America in 1843. Dr. Gray specialized in composites, or the Sunflower Family. Ligusticum is a Latin derivative of the town of Liguria in Italy, which is home of the related Levisticum. California Lomatium Lomatium californicum (Torrey & A. Gray) Mathias & Constance Native. Carrot Family (Apiaceae). Compound umbels display tiny radial yellow flowers that have 5 petals, 5 stamens, and 1 pistil. The stems that bear the secondary umbels, are numerous and long. The stems and leaves of this perennial are glabrous and glaucous (covered with a white, powdery substance that is easily rubbed off). The plant has many pinnately compound basal leaves with leaflets that are wedge-shaped, and lobed. The lobes are again toothed or lobed. Stem leaves are few and similar to basal leaves. California Lomatium can grow to 120 cm, and is usually found in wooded areas or on brushy slopes at lower elevations. Local Native Americans call(ed) this plant Icknish. They would collect the green sprouts while they were soft and tender, and eat it like celery. A favorite modern dish is called Icknish and Hotdogs. Try crushing the foliage on this plant – does it smell like celery? Lomatium is derived from the Greek word for bordered, referring to the fruits that have prominent marginal wings. Californicum is the Latinized version of California referring to the general range of this species. Piper’s Desert Parsley Lomatium piperi J. Coulter & Rose Also called Indian Biscuit-root. Native. Carrot Family (Apiaceae). This low-lying perennial bears small radial white flowers in compound umbels. The flowers are made up of 5 petals, 5 stamens with purple anthers, and 1 pistil. Several basal leaves surround the flowering stem. The basal leaves are pinnately or ternately compound with linear leaflets. Stem leaves are quite different from the basal leaves; they are simple, linear, and entire. The underground parts of this plant are important in determining Piper’s Desert Parsley (Lomatium piperi) from California Indian Potato (Orogenia fusiformis). Piper’s Desert Parsely (L. piperi) has a slender root that ends in a spherical tuber. California Indian Potato (O. fusiformis), on the other hand, has a slender, carrot-like taproot. If you decide to dig up these plants to look at the underground parts, please re-plant them. Lomatium is derived from the Greek word for bordered, referring to the fruits that have prominent marginal wings. Piperi is the latinized version of the name Piper, after C.V. Piper (1867–1926), an American botanist who authored more than five major floras, including the first Flora of the State of Washington. 54 Plant Descriptions © University of California Press Gray’s Lovage (Ligusticum grayi) © Stanford University Press California Lomatium (Lomatium californicum) Piper’s Desert Parsley (Lomatium piperi) 55 Plant Descriptions California Indian Potato Orogenia fusiformis S. Watson Native. Carrot Family (Apiaceae). As a member of the carrot family, this flower has the distinctive inflorescence – an umbel of umbels. The tiny white radial flowers have 5 petals, 5 stamens, and 1 pistil. The compound basal leaves have entire, linear leaflets. The plant grows very close to the ground, reaching heights of only 4-8 cm. Look for this low-lying perennial in May and early June at the edge of snow melt areas on the south slopes in loose, gravelly soil. Because it appears so early in the season in loose soil, there will probably not be any other flowers blooming near California Indian Potato. Orogenia comes from Greek, and means mountain race, referring to the montane habitats of this genus. Fusiformis is derived from Latin and means a spindly form or shape, referring perhaps to the linear leaflets. Western Sweet Cicely Osmorhiza chilensis Hook. & Arn. Native. Carrot Family (Apiaceae). The small, radial creamy white flowers are inconspicuous. The inflorescence is the typical form for members of the carrot family – an umbel. Each plant can have from one to several loose umbels that are short stalked when in flower and long stalked when in fruit. Western Sweet Cicely has basal leaves and alternate stem leaves that all look the same: compound leaves that are twice divided into 3s. The leaflets are coarsely toothed. When in flower, Western Sweet Cicely is usually 10-17 cm tall, but when the fruits mature the plant usually stands 20-30 cm tall. You’ll find this perennial in shaded, forested areas along the Pacific Crest Trail blooming in late June, and the bristly-hairy fruits are obvious through July. Growing nearby, you may see Large-leaf Sandwort (Moehringia macrophylla) or Jessica’s Stickseed (Hackelia micrantha). Osmorhiza is derived from the Greek word for sweet root, referring to the wonderful anise aroma of the roots of some plants in this genus. Chilensis is New Latin, meaning of Chile because it grows in South America, as well as North America. Western Sweet Root Osmorhiza occidentalis (Nutt.) Torrey Also called Mountain Sweet Cicely. Native. Carrot Family (Apiaceae). If you see a dainty umbel of umbels with yellowish-green flowers, scratch a stem or leaf and sniff. If you smell anise or licorice, you’ve got Western Sweet Root. This perennial has very small, regular, yellow flowers – you will need a hand lens to see the 5 petals, 5 stamens, and 1 pistil. A mature plant will have as many as five compound umbels. The twice pinnately compound leaves are mostly basal, with 5 to 7 leaflets that are also compound. The leaflets are irregularly and coarsely toothed. Each inflorescence is subtended by smaller compound leaves with three leaflets. The stem and alternate leaves have short hairs that can only be seen with a hand lens; however, there are conspicuous cob-webby hairs where the pedicel branches off the main stem. The fruits are long, linear and ribbed with no hairs. If scratched, these also smell of anise. Western Sweet Root usually grows 30-60 cm tall. Look for this flower in May and early June growing in partly shaded areas on dry hillsides. It is often found near Meadow Larkspur (Delphinium nuttallianum) and Greenleaf Manzanita (Arctostaphylos patula). Osmorhiza is derived from the Greek word for sweet root, referring to the wonderful anise aroma of the roots of this species. I don’t want you to dig it up, but scratch and sniff the stem and you’ll get an idea of how the root smells. Occidentalis is Latin for of the west, referring to the range of this species. 56 Plant Descriptions Photo by Br. Alfred Brousseau, St. Mary’s College California Indian Potato (Orogenia fusiformis) Western Sweet Cicely (Osmorhiza chilensis) Western Sweet Root (Osmorhiza occidentalis) 57 Plant Descriptions Yampah Perideridia oregana (S. Watson) Mathias Also called Squaw Potato, and Squawroot. Native. Carrot Family (Apiaceae). This delicate plant could be easily overlooked because the stem is so slender and the flowers so dainty. It could also be mistaken for a weak-looking Gray’s Lovage (Ligusticum grayi). But if you look closely, you will see that the leaves are quite different from Gray’s Lovage. The small radial white flowers of Yampah (P. oregana) have 5 sepals, 5 petals, 5 stamens, and 1 pistil, and are borne in an umbel of umbels. The slender, erect stem is glabrous and sometimes covered with a white bloom (pronounced blome), a powdery substance that easily rubs off. The plant has basal leaves and stem leaves, but the basal leaves may be difficult to find. They are pinnately compound and the leaflets are linear and entire. The one or two stem leaves are similar to, but smaller than the basal leaves. This perennial usually grows between 20-50 cm tall, and blooms in August in moist, open meadows. Growing nearby, you may find Swamp White Heads (Sphenosciadium capitellatum) or Western Coneflower (Rudbeckia occidentalis). Perideridia is derived from the Greek words meaning around the neck, referring to the involucre the ring of bracts at the base of the primary umbel. Oregana is the latinized version of Oregon, referring to the general range of this plant. Northern Sanicle Sanicula graveolens DC. Also called Sierra Snakeroot. Native. Carrot Family (Apiaceae). Take a long sniff of this plant. Does it smell like a familiar culinary herb? I think it smells like cilantro! This low-growing to erect plant has large obvious leaves and tiny yellowish-green flowers. The flowers are radial with 5 sepals, 5 petals, 5 stamens, and 1 pistil, and are arranged in rounded compound umbels. The pinnately compound leaves are alternate and the lowermost leaves are often attached below the ground level, and seem to arise separately from the stem. The leaflets are toothed or lobed. This perennial usually only grows 5-20 cm. Northern Sanicle is an early bloomer on Mount Ashland, showing up in early June in dry areas or on rocky slopes. You may find Meadow Larkspur (Delphinium nuttallianum) or Chocolate Lily (Fritillaria atropurpurea) growing nearby. Sanicula comes from the Latin word meaning to heal, referring to the medicinal properties of some of the species in this genus. Graveolens is derived from two Latin words meaning to emit a heavy smell, referring to the strong cilantro scent. Swamp White Heads Sphenosciadium capitellatum A. Gray Also called Woolly Head Parsnip, and Ranger’s Buttons. Native. Carrot Family (Apiaceae). Tiny radial flowers are arranged in compound umbels, and the secondary umbels are so tightly packed with the tiny white flowers that they look like little round buttons. The tall, stout stems are rough to the touch, owing to short, stiff hairs. The large basal leaves are pinnately compound and the large leaflets are coarsely toothed. Stem leaves are alternate and small, also pinnately compound, with a large conspicuous sheath at the base. This large perennial usually reaches heights of 90-180 cm, and grows in open, wet meadows. Swamp White Heads bloom in July and if you are lucky, you will find Richardson’s Geranium (Geranium richardsonii) and Wiggin’s Lily (Lilium pardalinum ssp. wigginsii) blooming nearby. Sphenosciadium is derived from Greek words meaning wedge umbrella, referring to the umbel. Capitellatum is derived from several Latin words meaning small head, referring to the tight rounded umbels of flowers. 58 Plant Descriptions Photo by Bob Vos Yampah (Perideridea oregana) Northern Sanicle (Sanicula graveolens) Swamp White Heads (Sphenosciadium capitellatum) 59 Plant Descriptions Birthwort Family (Aristolochiaceae) Wild Ginger Asarum caudatum Lindley Also called Ginger Root, Long-Tailed Ginger, British Columbia Wild Ginger, and Western Wild Ginger. Native. Birthwort Family (Aristolochiaceae). Because the blooms lie on the ground beneath the large “pillowy” leaves, flower will be difficult to find unless you can identify the leaves, which rise in pairs above the flower. The uniformly green basal leaves are heart-shaped (cordate) to kidney-shaped (reniform) with entire margins. The maroon flower is worth searching for. Gently push the leaves aside until you find the three long, tapering “petals” that are actually modified sepals. The tube that encloses the 12 stamens and 1 pistil is made up of fused sepal bases. The tube is white on the inside with one maroon stripe. This perennial prefers moist, shaded areas, so search for it along the forested portions of the Pacific Crest Trail from late June to mid July. Look for Heart-leaf Spring Beauty (Claytonia cordifolia) and Baneberry (Acteae rubra) growing nearby. The edible rhizome has a ginger scent when crushed, but the ginger in your spice rack is from a tropical plant, Zingiber spp. Asarum is Greek, but the derivation is unknown. It could mean “without broom,” but its reference is uncertain. Caudatum is Latin for tail, or trunk of a tree. This could be in reference to the tapering sepals that appear to have tails, or it could indicate the fact that these flowers are only found in forested areas, and therefore, at the base of tree trunks. Sunflower Family (Asteraceae) Yarrow Achillea millefolium L. Also known as Soldier’s Wound-wort, Devil’s Plaything, Milfoil, Tansy, Thousand-leaf, and Nosebleed. Native. Sunflower Family (Asteraceae). This perennial has alternate leaves that are three times pinnately compound, giving them a finely dissected, fern-like appearance. Usually, there are many basal leaves surrounding the flowering stem; all leaves are hairy. The inflorescence is a flat-topped corymb, and what appears to be one regular flower is actually a composite of three to eight tiny whitish ray flowers (that look like petals) and many disk flowers (the yellow, white, or purple center of the “flower”). This erect plant stands from 20-40 cm tall, and is found in dry, open areas. Look for Yarrow from late June through July around Rabbit Ears, Grouse Gap, and in the picnic area. Nearby, you may also find Siskiyou Mountain Owl’s Clover (Orthocarpus cuspidatus var. cuspidatus), or Common Blue Flax (Linum lewisii). Yarrow has a long and honorable history of medicinal use. Native peoples across America and in ancient Europe used Yarrow to make a tea to induce sweating and thereby reduce fevers, or to improve indigestion. Leaves were chewed and the poultice was placed on wounds to stop bleeding and assist the healing process. Yarrow was also used as an insect repellant: the stems and leaves were placed on hot coals to repel mosquitoes. In addition to these and many other medicinal applications, Yarrow was used in ancient Europe and China for divination, hence the name Devil’s Plaything. The genus name, Achillea, comes from the legend that the plant was used by Achilles of ancient Greek mythology to heal the wounds of his soldiers. Homer’s Iliad contains the legend that Chiron, wisest of the centaurs and skilled in medicine, advised Achilles to make an ointment from yarrow to heal his soldiers’ wounds after the siege of Troy. Yarrow was the Greek emblem for war, perhaps because of its use in healing wounds. Millefolium is Latin for thousand leaves, referring to the highly dissected leaves. 60 Plant Descriptions Wild Ginger (Asarum caudatum) Yarrow (Achillea millefolium) 61 Plant Descriptions Trail Plant Adenocaulon bicolor Hook. Native. Sunflower Family (Asteraceae). Trail Plant is a composite, therefore it is made up of many small flowers densely packed in one head. The tiny heads are arranged in an open panicle-like cyme. The outer flowers on each head are pistillate (contain only the female reproductive parts) and the inner flowers on each head are staminate (contain only the male reproductive parts). The staminate flowers look like a radial flower with five tiny white petals, but the pistillate flowers appear to have four or five small white petals and many glandular hairs, making these flowers sticky. A basal rosette of triangular leaves surround the single leafless flowering stem. Each shallowly lobed leaf has a thin, white cottony covering on the lower surface. The stem has glandular hairs near the top. Look for this common perennial in shaded areas and in the forest at lower elevations. Other flowers that may be blooming nearby include White-Veined Wintergreen (Pyrola picta), and White Hawkweed (Hieracium albiflorum). This flower is called Trail Plant because when a human or an animal touches it, the underside of the leaf faces upward. Because the undersides of the leaves are whitish in color, they act as a marker for the trail. Adenocaulon is a combination of Greek and Latin roots that mean glandular stem. Feel the upper part of the stem – it is sticky because these glands contain sticky oils. Bicolor is Latin for two colors, referring to the leaves which are green on the upper surface and white on the lower surface. Orange Agoseris Agoseris aurantiaca (Hook.) E. Greene Also called Slender Agoseris. Native. Sunflower Family (Asteraceae). When I first noticed this plant, I though it looked like a tall, orange dandelion. This perennial has only basal leaves that are oblanceolate, and generally entire, but may have irregular, shallow teeth along the lower 2/3 of the leaf. The erect stem has sparse white woolly hairs that grow more dense toward the top. What appears to be one regular flower is actually a composite head of many small ligulate flowers. Each flower will produce a seed with a hairy parachute, similar to dandelions. This flower grows 25-35 cm, and is found in meadows. Look for Orange Agoseris from late June through July around Rabbit Ears and around the picnic area. You may also see Sitka Valerian (Valeriana sitchensis) or Western Sweet Root (Osmorhiza occidentalis) growing in the shade nearby. Agoseris is Greek for goat chicory; aurantiaca is New Latin for orange-colored, obviously referring to the color of the many tiny flowers on the head. Tall Agoseris Agoseris elata (Nutt.) E. Greene Native. Sunflower Family (Asteraceae). Tall Agoseris has a fairly stout stem with long white cobwebby hairs that vary from sparse to dense, and the stem is often mottled with pink. The basal leaves are at least 1 cm wide (usually wider), and vary from entire to toothed or lobed, and from glabrous to sparsely hairy. The leaves and stem exude a milky sap when broken. The phyllaries, which may be mistaken for sepals, overlap in 3 or 4 series. They are short, wide, and soft-hairy. This plant looks similar to Orange Agoseris (A. aurantiaca), but there are several features that distinguish the two. The leaves of Tall Agoseris are wider than those of Orange Agoseris. The phyllaries of Tall Agoseris are shorter and wider than those of Orange Agoseris, and they are hairy, while those of Orange Agoseris are glabrous. Most textbooks claim that the flowering head of Tall Agoseris is yellow, drying pinkish. However, the flowers on Mount Ashland are orange, just like the flowering heads of Orange Agoseris. This perennial often grows 30-65 cm, and is found in open meadows, usually on dry soil. Nearby, you may find Large-flowered Agoseris (A. grandiflora) and Orange Agoseris (A. aurantiaca). Agoseris is Greek for goat chicory; elata is Latin for high, or lofty, referring to the plant’s height. Tall Agoseris towers over many other plants. 62 Photo by Bob Vos Plant Descriptions © Stanford University Press Trail Plant (Adenocaulon bicolor) Orange Agoseris (Agoseris aurantiaca) Tall Agoseris (Agoseris elata) 63 Plant Descriptions Large-flowered Agoseris Agoseris grandiflora (Nutt.) E. Greene Also called Grand Mountain Dandelion. Native. Sunflower Family (Asteraceae). That is not one yellow flower at the top of the stem, rather it is a head of many tiny yellow flowers. What appears to be one petal is one ligulate flower. Looking closely at the tip of the ligule, you will notice that there are 5 lobes indicating that one ligule is made up of five petals that are fused together. What, at first glance, appears to be the sepals are bracts that are called phyllaries. The narrow, lanceolate phyllaries appear glabrous without magnification, and are usually purple-tipped and mottled with pink. The basal leaves are either entire and linear, or lobed along the upper half of the leaf margin. The upward-pointing lobes and the blade along the midrib are narrow. The leafless stem is sparsely hairy with long, white hairs. The hairs are especially dense just below the flowering head. Large-flowered Agoseris is a perennial that grows in dry, open areas on the south slopes. Nearby, you might also find Orange Agoseris (A. aurantiaca) and Tall Agoseris (A. elata). Agoseris is Greek for goat chicory; grandiflora is Latin for large flower. Pearly Everlasting Anaphalis margaritacea (L.) Benth. & Hook. Also called Western Pearly Everlasting. Native. Sunflower Family (Asteraceae). Prepare yourself for a tactile treat, for the velvety white leaves and stem make this plant a delight. As a species in the Sunflower Family, what appears to be one flower is actually a head of many flowers, but the white and yellow flowers are so small that the individual parts often cannot be distinguished without magnification. When the plant first blooms, the yellow flowers in the center are not visible. But as this perennial matures, the white bracts spread open revealing the yellow flowers. The leaves are alternate, entire and sensuously fuzzy. Pearly Everlasting can be found in dry, open areas at the base of the ski slopes, and along the Pacific Crest Trail from USFS Road 20 to Grouse Gap. Look for Wright’s Cudweed (Gnaphalium canescens ssp. thermale) or Pussy Paws (Calyptridium umbellatum) growing nearby. Anaphalis is Greek and means male parts throughout. This must be in reference to the fact that on many of the plants in this genus, the male (staminate) and female (pistillate) flowers are on separate plants. Therefore, the plants with only staminate flowers would have “male parts throughout.” Margariticea is Greek for a pearl, which could be in reference to the overall color of the plant, or to the small, white, rounded heads of flowers that may have reminded someone of pearls. Rosy Pussytoes Antennaria rosea E. Greene Also called Pussy Toes, Pink Pussytoes, Cat’s Paws, Everlasting, Ladies Tobacco, and Rosy Everlasting. Native. Sunflower Family (Asteraceae). In the early stages of blooming, these pink and white flower heads nod, eventually standing erect as the plant matures. What appear to be pink or rose-colored petals are in fact showy bracts. The white portion in the center is surprisingly made up of many tiny flowers. The individual flowers are too small to be distinguished without magnification. The oblong to linear stem leaves are entire and alternate, and the basal leaves are oblanceolate. The fuzzy hairs on the stem and leaves are a delight to touch. This perennial grows 10-25 cm tall. Look for Rosy Pussytoes in open, dry areas on the south slopes from mid-June through mid-July. Nearby, you should see Mt. Ashland Lupine (Lupinus aridus var. ashlandensis) and Dusty Maidens (Chaenactis douglasii). This plant was discovered in 1881, and has had its name changed several times. Some authorities consider it to be synonymous in botanical nomenclature to two other species. Antennaria is Latin for antenna, and comes from the supposed resemblance of this plant to the antennae of insects. Rosea is Latin for rosy or rose colored, referring to the color of the bracts that surround the white flowers. 64 Plant Descriptions Photo by Bob Vos Large-flowered Agoseris (Agoseris grandiflora) Pearly Everlasting (Anaphalis margariticea) Rosy Pussytoes (Antennaria rosea) 65 Plant Descriptions Pale Agoseris Agoseris glauca (Pursh) Raf. Also called Short-beaked Agoseris. Native. Sunflower Family (Asteraceae). First of all, you need to know that the “flower” at the top of the stem is not one yellow flower, but a head of many tiny yellow flowers. What appears to be one petal is one ligulate flower. The tip of each ligule has five tiny lobes indicating that one ligule is made up of 5 fused petals. The phyllaries, which could be mistaken for sepals, are long-tapered and can vary from glabrous to densely soft-hairy. They overlap in several series. Basal leaves that vary from entire to toothed to lobed surround the glabrous stem. When torn, the leaves and stem exude a milky sap. This perennial usually grows 3-30 cm, and is found in coniferous forests or moist meadows. You may confuse Pale Agoseris with Large-flowered Agoseris (A. grandiflora) or Dandelion (Taraxacum officinale). To tell them apart, look first at the phyllaries. Those of Large-flowered Agoseris are narrow and lanceolate, appear glabrous without magnification, and are usually purple-tipped and mottled with pink. The phyllaries of Dandelion are glabrous and the outer series is reflexed. Then look at the stem. The stem of Large-flowered Agoseris is sparsely long-hairy. The other common name for this plant, Short-beaked Agoseris refers to the fruits, which consist of a seed attached to a fluffy, white parachute made up of many fine bristles. The seed is attached to the parachute by a short structure called a beak. Agoseris is Greek for goat chicory. Glauca is derived from the Latin word for gray, referring to the stem, which is grayish as a result of hairs or bloom (pronounced blome), a white substance that easily rubs off when the plant is touched. Heartleaf Arnica Arnica cordifolia Hook. Also called Leopard’s Bane. Native. Sunflower Family (Asteraceae). Here is another bright member of the Sunflower Family. What appears to be one yellow flower is actually a head of many flowers. The “petals” are ray flowers and the center is filled with disk flowers. The “sepals” are bracts that are collectively referred to as the involucre. These are pointed at the tip and have long, white hairs. The two to four opposite leaves are coarsely toothed and the base is often, but not always, heart-shaped (cordate). Many basal leaves are present and often persist after the flower dies back. The stem and leaves are slightly hairy, and sometimes a bit sticky. Heartlead Arnica grows 10-25 cm tall. This perennial is usually found in partly shades areas on the north slopes. It flowers in late June to early July with Sitka Valerian (Valeriana sitchensis) and Spotted Coralroot (Corallorhiza maculata). In the past, Heartleaf Arnica was popular for several medicinal treatments. The flowers can be steeped in water and applied to cuts and wounds, and it is also good for chapped lips. It is uncertain whether Arnica is an ancient Latin or Greek name. Cordifolia means heart-shaped leaf. Mugwort Artemisia douglasiana Besser Also called Douglas’ Sagewort. Native. Sunflower Family (Asteraceae). This highly aromatic perennial smells similar to Mountain Sagebrush (Artemisia tridentata var. vaseyana). The small, rather inconspicuous yellow flowers are actually heads of several tiny flowers, but you’ll need some form of magnification to distinguish each individual flower. The heads are generally bell-shaped and nodding. Alternate grayish-green leaves are evenly spaced along the tall stem. The lower surface of the leaves has many tiny, white hairs, while the upper surface has fewer white hairs. The leaves are irregularly lobed, and the tips of the lobes are pointed. Some of the upper leaves may be entire. Stems are fairly stout and can grow up to 120 cm. Mugwort can be found along County Road 1151 between miles 7 and 8, and between Road 300 and Grouse Gap. There is also one clump of it at the base of the Sonnet chair lift. The genus name, Artemisia, is for the Greek goddess of the hunt, Artemis. Douglasiana honors David Douglas (1798-1834), the celebrated Scottish botanist who explored the western coast of North America with John Scouler from 1825-1827, and again from 1830-1832. Douglas died while exploring the Hawaiian islands, and is buried there. 66 Photo by Charles Webber, California Academy of Sciences Plant Descriptions Pale Agoseris (Agoseris glauca) Heartleaf Arnica (Arnica cordifolia) Mugwort (Artemisia douglasiana) 67 Plant Descriptions Mountain Sagebrush Artemisia tridentata Nutt. var. vaseyana (Rydb.) Beetle Also called Sage, Sagebrush, and Mountain Big Sagebrush. Native. Sunflower Family (Asteraceae). Mountain Sagebrush, which is very popular for its soothing aroma, has small yellow flowers that are quite inconspicuous. This is a member of the Sunflower Family, which means that what appears to be one flower is actually a head of several tiny flowers. A hand lens or magnifying glass will make it easier to distinguish the individual flowers. The flowering stalk appears in early August, and by mid-August, the flowers are in bloom. The alternate leaves are grayish-green because of the dense white hairs that cover them, and usually have three lobes at the tip. Occasionally, some leaves will have four or five lobes. This shrub usually grows to about 60-90 cm tall, and is found on the dry, open south slopes. Growing nearby will be Sulfur Flower (Eriogonum umbellatum) and Leafy Aster (Aster foliaceus). The genus name, Artemisia, is for the Greek goddess of the hunt, Artemis. Tridentata is Latin for three-toothed, referring to the three lobes at the apex of most leaves. Leafy Aster Aster foliaceus Lindley var. lyallii (A. Gray) Cronq. Also called Leafy-headed Aster, and Lyall’s Aster. Native. Sunflower Family (Asteraceae). Because this flower is a composite, the blue “petals” on these plants are actually tiny flowers, called ray flowers. The yellow center is made up of many tiny yellow disk flowers. What appear to be leafy sepals are phyllaries that are collectively referred to as the involucre. The alternate leaves are entire and glabrous, and the base of the leaves clasps the stem. The stem carries minute hairs and is often, but not always, branched, and produces one to several heads. There are usually 4-12 heads per plant. Leafy Aster can grow up to 30 cm tall, and is often found in moist areas on the south slopes. Growing nearby, you will find Tall Agoseris (Agoseris elata), and Tall Annual Willowherb (Epilobium brachycarpum). Leafy Aster and Leafy-headed Aster are appropriate names because of the many, large phyllaries that are leaf-like. In Greek, aster means star. Foliaceus is Latin for leafy. Dusty Maidens Chaenactis douglasii (Hook.) Hook. & Arn. Also called Hoary Chaenactis, Pincushion, and Douglas’ Dusty Maiden. Native. Sunflower Family (Asteraceae). The creamy-white radial flowers in the terminal heads have 5 petals and exserted stamens. The branching stem usually has three to many flower heads, but it can have only one or two flower heads. The 15-25 cm erect stems are reddish with white cobwebby hairs. Leaves are compound and alternate with white cobwebby hairs, and often subtend branches on the upper portion of the stem. The larger leaves toward the bottom of the stem are usually twice pinnately compound. There is also a basal rosette of leaves that are twice pinnately compound. This is a biennial that sometimes flowers in its first year. These flowers are not abundant, but they grow around Rabbit Ears and in other dry, open areas in late June and early July. You may find Spear-leaf Stonecrop (Sedum lanceolatum) or Siskiyou Mountains Owl’s Clover (Orthocarpus cuspidatus) blooming nearby. In Greek, chaenactis means gaping ray, referring to the enlarged outer corollas of some of the species in this genus. Douglasii is derived from the name of David Douglas (1798-1834). Douglas arrived at the mouth of the Columbia River in April 1825 on a Hudson Bay Company ship. The Scottish botanist explored and collected plants in what is now Oregon, Washington, and Idaho from 1825-1827. 68 Plant Descriptions © Stanford University Press Mountain Sagebrush (Artemisia tridentata var. vaseyana) Leafy Aster (Aster foliaceus var. lyallii) Dusty Maidens (Chaenactis douglasii) 69 Plant Descriptions Pineapple Weed Chamomilla suaveolens (Pursh) Rydb. Also called Rayless Chamomile. Non-native from western Canada and northeast Asia. Sunflower Family (Asteraceae). You can confirm your identification of this plant by rubbing your fingers over the leaves or the flowers and taking a long, leisurely sniff. Mmmm…pineapple! The many tiny yellow flowers are held on a button-like head, and each plant produces several flowering heads. The alternate leaves are irregularly pinnately lobed. The plant usually is branched, and low-growing, only reaching heights of 510 cm. Pineapple Weed usually grows in disturbed soils, so look for it around the ski lodge and along County Road 1151 up to the ski lodge parking area from mid-July through mid-August. You may find Mountain Tarweed (Madia glomerata) or Sticky Cinquefoil (Potentilla glandulosa) blooming nearby. According to Webster’s New World Dictionary, chamomilla is derived from several Greek words that mean earth apple. Derived from Latin, suaveolens means sweet-scented, of course! Chicory Cichorium intybus L. Also called Blue Sailor, Ragged Sailors, Blue Daisy, Blue Dandelion, Coffee-Weed, and Wild Succory. Non-native from Europe. Sunflower Family (Asteraceae). There are not many of these sky-blue flowers growing on Mount Ashland. Because many people consider this non-native to be a weedy plant, they are happy that there are very few on Mount Ashland. The many blue or lavender petals are actually the ligule of individual flowers that will each produce its own fruit (a dry achene). In the center of the head, pistils of each of these minute flowers are obvious. The pistils arise out of a tube that is white at the base and blue at the top. Inside each tube are 5 stamens, making each tube with its corresponding ligule a complete flower. If you look closely at the tip of the ligule, you will see five lobes. These lobes represent five petals that are fused together to form the ligule. The alternate leaves of Chicory are coarsely toothed and/or pinnately lobed. This plant grows 30-120 cm. Look for it along County Road 1151 at lower elevations, next to Idaho Gum Weed (Grindelia nana) or Bull Thistle (Cirsium vulgare). Chicory is well known as a coffee substitute. Colonists from Europe and later, the Native Americans, and still later, southerners in the United States, roasted and ground the roots to use as coffee or to add to their coffee. Chicory also has many medicinal uses – almost as many as Dandelion (Taraxacum officinale). Cichorium is an old Arabic name. The only known meaning of intybus is endive or chicory. Bull Thistle Cirsium vulgare (Savi) Ten. Also called Common Thistle. Non-native from Europe. Sunflower Family (Asteraceae). Bull Thistle is a biennial, which means it completes its life cycle in two years. During the first year, only the spiny basal leaves will be obvious growing in a rosette on the ground. The second year, the plant sends up a flowering stalk that produces several purple heads of flowers that are surrounded by spiny bracts. The alternate leaves on the stem are lobed and each lobe is tipped with a spine. The stem, itself, is winged – it has green tissue similar to a leaf blade, and this tissue is also lobed and has many spines. There are very few of these weedy plants on Mount Ashland above 6200 feet, but look for them along County Road 1151 at lower elevations. You may find them growing next to Mugwort (Artemisia douglasiana) or Dalmatian Toadflax (Linaria genistifolia var. dalmatica). Cirsium is the Greek word for thistle. Vulgare is Latin for common-place, referring to the fact that this plant is very common, indeed too common for some people’s liking! 70 Plant Descriptions Pineapple Weed (Chamomilla suaveolens) Chicory (Cichorium intybus) Bull Thistle (Cirsium vulgare) 71 Plant Descriptions Naked-stem Hawksbeard Crepis pleurocarpa A. Gray Native. Sunflower Family (Asteraceae). The bright yellow five to eight “petals” on these “flowers” are actually the ligules of five to eight individual little flowers that are clustered together on a head. The tip of each ligule has 5 shallow lobes. Each ligule has a corresponding slender tube in the center of the “flower.” Inside this tube, there are five miniscule stamens, and the slender pistil that ends with two curled lobes at the tip. The sepal-like structures are called phyllaries, and they are elongated, narrow and linear. Naked-stem Hawksbeard has mostly basal leaves that are dandelion-like, with downward pointing lobes. In spite of the name, this plant may have a few stem leaves. If so, they will be short, grass-like, and entire. The stems are usually branched, and each plant may yield 7 to 40 heads. Break open a leaf or one of the small branches to see the white, milky sap. The stems can vary from glabrous to slightly short-hairy. This perennial grows 1545 cm on dry, open slopes. It flowers in June. Crepis is derived from the Greek word for boot or sandal. The reason for this genus name is unknown. Pleurocarpa is derived from two Greek words meaning ribbed fruit. If you use your hand lens, you will find 10 prominent ribs on the achenes. Greene’s Goldenbush Ericameria greenei (A. Gray) G. Nesom Native. Sunflower Family (Asteraceae). The number of “petals” or ray flowers on these flower heads varies from 2 to 6. The number of disk flowers varies from 7 to 20. Usually, several flowering heads are clustered at the end of a stem. The tightly packed alternate, sticky leaves are entire and more or less linear. The stems are woody at the base, and many grow together in large clumps. In fact, these plants are listed in botanical keys as shrubs. These flowers bloom late in August through September, but the sticky green leaves become obvious in early July. Look for this perennial on the dry, open slopes between the split in the road for the summit, and Grouse Gap Shelter. Nearby, Sulfur Flower (Eriogonum umbellatum), Coyote Mint (Monardella odoratissima), and many more flowers bloom. The genus name, Ericameria, is constructed from two Greek words: eric, which means heath, and meria, which means “a part.” Edward L Greene (1843-1915), an Episcopal priest turned botanist, collected and described many western species. Alice’s Daisy Erigeron aliceae Howell Also called Alice Eastwood’s Fleabane. Native. Sunflower Family (Asteraceae). These prolific plants are a common sight on the south slopes of Mount Ashland. The “petals” or ray flowers vary in color from lavender to pink to white, and the disk flowers in the center are yellow. The “sepals” or phyllaries have long, white hairs, some of which may be glandular. The alternate leaves are entire, often have wavy margins, are distinctly hairy, and clasp the stem at the base. These perennials grow 20-45 cm tall, and are found blooming in any open, dry areas in late June through July. Nearby, you may find Nettle-leaf Giant Hyssop (Agastache urticifolia) or Siskiyou Mountains Owl’s Clover (Orthocarpus cuspidatus). Alice’s Daisy could be mistaken for Subalpine Fleabane (Erigeron peregrinus). The best way to tell the difference is by looking at the leaves. If the leaves are distinctly hairy, the plant is Alice’s Daisy. If the leaves are glabrous or only sparsely hairy, the plant is Subalpine Fleabane. Erigeron is Greek and means either “a kind of plant” or “early old age.” It is uncertain which meaning is applicable or why. Aliceae is a Latin derivative for the name Alice, in honor of Alice Eastwood (1859-1953) an American botanist raised in the western United States. Eastwood authored three books of California flora, and was the Curator at the California Academy of Sciences from 18941949. She was listed in American Men of Science in 1906. 72 © Stanford University Press Plant Descriptions Naked-stem Hawksbeard (Crepis pleurocarpa) Greene’s Goldenbush (Ericameria greenei) Alice’s Daisy (Erigeron alicea) 73 Plant Descriptions Scabland Fleabane Erigeron bloomeri A. Gray Native. Sunflower Family (Asteraceae). This low-growing perennial with yellow button-like flowers looks exactly like Cut-leaf Daisy, except for the leaves. Scabland Fleabane has only basal leaves that are linear and entire, similar to small grass blades. However, the basal leaves of Cut-leaf Daisy are pinnately lobed or compound. For more details, refer to the description of Cut-leaf Daisy below. Look for Scabland Fleabane on dry, open slopes. Erigeron is Greek and could mean a kind of plant or early old age. Mr. Hiram G. Bloomer (18191874) is honored in this specific epithet. Mr. Bloomer arrived in San Francisco in 1852 from his native state, New York. As his obituaries claim, he was regarded as the pioneer resident botanist of the Pacific Coast. He worked as the Curator of the Academy of Natural Sciences in San Francisco for many years. Cut-Leaf Daisy Erigeron compositus Pursh Also called Dwarf Mountain Erigeron. Native. Sunflower Family (Asteraceae). The flowers of this little plant look like little yellow buttons. They are difficult to identify because the structures are so tiny. Believe it or not, what appears to be one flower is a head of many miniscule flowers. The corolla of each individual flower has 5 tiny lobes, and in the center of these lobes are five stamens and one pistil. The sepal-like structures are phyllaries. They are green with red tips and covered with long white hairs. Each stem produces one terminal flowering head. Stem leaves are few, alternate, and entire, while the many basal leaves are pinnately compound. The leaves and stems also have long white hairs. This perennial grows no taller than 12 cm. It blooms in June and early July in rocky crevices and on open, dry, sandy soil. Look for Cut-leaf Daisy around Rabbit Ears and up to the summit on the south slope. Davidson’s Penstemon (Penstemon davidsonii) and Cream Stonecrop (Sedum oregonense) may be blooming nearby. Scabland Fleabane (E. bloomeri) looks almost identical to Cut-Leaf Daisy. To tell them apart, look at the basal leaves. If they are pinnately compound, your specimen is Cut-Leaf Daisy (E. compositus). If the basal leaves are simple and entire, you have Scabland Fleabane (E. bloomeri). Erigeron is Greek and could mean a kind of plant or early old age. It is uncertain which meaning is applicable or why. Compositus is Latin meaning placed together, referring to the tiny flowers which grow close together in a tight head. Subalpine Fleabane Erigeron peregrinus (Pursh) E. Greene Also called Foreign Erigeron, Wandering Daisy, Wandering Fleabane. Native. Sunflower Family (Asteraceae). These abundant perennials are a common sight on the south slopes of Mount Ashland. On this plant, as on all plants of the Sunflower Family, the structures that appear to be petals are actually small flowers, called ray flowers. The ray flowers on this plant vary in color from lavender to pink to white, and the small disk flowers that make up the center part of the flowering head are yellow. The “sepals” or phyllaries have dense hairs that bear an oil gland at the tip. These are known as glandular hairs, and they usually make the surface sticky to the touch. The alternate leaves are entire, often have wavy margins, are glabrous to sparsely-hairy, and clasp the stem at the base. These perennials grow 20-45 cm tall, and are found blooming in any open, dry area in late June through July. Nearby, you may find Oregon Sunshine (Eriophyllum lanatum) or Scouler’s Hawkweed (Hieracium scouleri). The best way to tell the difference between Subalpine Fleabane and Alice’s Daisy (E. alicea) is by looking at the leaves. If the leaves are distinctly hairy, the plant is Alice’s Daisy. If the leaves are glabrous or only sparsely hairy, the plant is Subalpine Fleabane. Erigeron is Greek and could mean a kind of plant or early old age. It is uncertain which meaning is applicable or why. Peregrinus is derived from the Latin word meaning to wander or travel abroad, perhaps referring to the seeds which are similar to those of Dandelion and are, therefore, wind dispersed. 74 Photo by Bob Vos © Stanford University Press Plant Descriptions Scabland Fleabane (Erigeron bloomeri) Photo by Bob Vos Cut-leaf Daisy (Erigeron compositus) Subalpine Fleabane (Erigeron peregrinus) 75 Plant Descriptions Oregon Sunshine Eriophyllum lanatum (Pursh) James Forbes var. grandiflorum (A. Gray) Jepson Also called Woolly Sunflower, Common Woolly Sunflower, and Golden Yarrow. Native. Sunflower Family (Asteraceae) This yellow daisy-like flower is actually a head of many small flowers. What appears to be the petals is actually a row of 8-13 ray flowers, and the center of the head consists of small disk flowers, from 20 to 300. If you look closely with your hand lens, you will notice that each of the disk flowers has a short 5-lobed corolla. Although Oregon Sunshine tends to grow in clumps, there is only one flower head at the end of each stem. Leaves are alternate and entire with wavy margins and have many fine, white hairs that make the plant feel woolly. Oregon Sunshine blooms from June through September on dry, open hillsides. Look for it in open areas along USFS Road 20, the Pacific Crest Trail, and around Rabbit Ears. Coyote Mint (Monardella odoratissima) or Ballhead Sandwort (Arenaria congesta) may be blooming nearby. In Greek, eriophyllum means woolly leaf, and lanatum is Latin for wool. Both words describe the white, woolly hairs that grow on the leaves and stem. Wright’s Cudweed Gnaphalium canescens DC. ssp. thermale (E.Nelson) Stebb. & Keil Native. Sunflower Family (Asteraceae). A biennial or short-lived perennial, Cudweed is usually branched at the top bearing many heads of flowers in a panicle-like arrangement. Each tiny head of flowers contains pistillate flowers around the margin and staminate flowers in the center. All flowers are yellow. The phyllaries (sepal-like structures) are graded in several series and make the head more or less bell shaped. Leaves are alternate, entire, and linear. Stem, leaves, and phyllaries are covered with many white, woolly hairs giving the plant a velvety feel, and a whitish-gray color. The entire plant is strongly scented. Cudweed grows usually 25-45 cm tall, and is found roadside on Mount Ashland. Growing nearby, you may see a similar plant, Pearly Everlasting (Anaphalis margariticea), or Siskiyou Penstemon (Penstemon anguineus). Superficially, this plant resembles Pearly Everlasting (A. margariticea) because of the velvety, whitish-gray stems and leaves. But Cudweed is a more delicate plant than Pearly Everlasting: the stems are not as stout , the leaves are much smaller, and the flowering heads are much smaller. Gnaphalium is Greek for lock of wool, referring to the many white, woolly hairs on the stem and leaves. Canescens is Latin for becoming gray, again referring to the hairs which make the stem and leaves appear to be grayish in color. Thermale means of warm springs, referring to the general habitat of this subspecies. Idaho Gum Weed Grindelia nana Nutt. Also called Idaho Resin Weed, and Gum Plant. Native. Sunflower Family (Asteraceae). These large, bright yellow flowers appear to have many petals and a yellow center. Because this flower is a composite, the petal-like structures are complete flowers, called ray flowers, and the yellow center is made up of many tiny flowers called disk flowers. The many sepal-like structures, called phyllaries, are curled back and highly resinous. Alternate leaves are coarsely toothed and the upper-most leaves are reflexed. The stems are reddish-purple, glabrous, and usually branched. Gum Plant grows to 20-45 cm along USFS Road 20 at lower elevations. It blooms late July through mid-August. Next to it, you may find White Clover (Trifolium repens) or Woolly Mullein (Verbascum thapsus). The plants in this genus are named after D.H. Grindel (1776-1836), a Latvian botanist. Nana is a Greek word meaning dwarf; it is uncertain to what this refers. 76 Plant Descriptions © University of California Press Oregon Sunshine (Eriophyllum lanatum var. grandiflorum) Wright’s Cudweed (Gnaphalium canescens ssp. thermale) Idaho Gum Weed (Grindelia nana) 77 Plant Descriptions Whitney’s Bristleweed Hazardia whitneyi (A. Gray) E. Greene var. discoidea (J. Howell) W. Clark Native. Sunflower Family (Asteraceae). One terminal head of yellow flowers rises above the alternate, coarsely toothed leaves. The tough, leathery leaves superficially resemble holly leaves. The phyllaries are long, narrow and pointed, and are graduated in several series. The stem, which is often woody at the base, is green above and brownish below, and bears white hairs. These perennials grow in dense clumps, and are fairly low to the ground, growing only 15-20 cm. One small population of Whitney’s Hazardia grows 300 feet south of the top of the Sonnet chairlift on small rock outcrop, and more can be found on the Knoll at mile 8 on County Road 1151. Looking around, you may see Mugwort (Artemisia douglasiana) or Oregon Grape (Berberis nervosa). Barclay Hazard, a 19th century California botanist, is commemorated with this genus name. Mr. Whitney is honored in the specific epithet. Discoidea is a Greek derivative for a round plate, and is a descriptive word for the flowers, which are often called discoid. Tall Sneezeweed Helenium bigelovii A. Gray Also called Bigelow’s Sneezeweed. Native. Sunflower Family (Asteraceae). These large, yellow composite flowers seem to say, “Summer is here!” Many yellow ray flowers surround and subtend even more yellow disk flowers. The ray flowers are strongly three-lobed, but if you look closely at the disk flowers with your hand lens, you will see that these flowers have 4 or 5 lobes. The disk flowers are held on a rounded receptacle. One large terminal head of flowers rises high on a stem with alternate, entire leaves. The entire stem of this perennial and the mid-vein of the leaves carry curly white hairs, and the lower portion of the stem is winged. Because Tall Sneezeweed prefers damp or wet places, you will find it in all the wet meadows on the south slopes, along with Oregon Checkerbloom (Sidalcea oregana) and perhaps Western Coneflower (Rudbeckia occidentalis). This plant is poisonous to livestock. If eaten by an animal, a poisonous glycoside accumulates in the animal’s tissues and causes “spewing sickness.” This showy yellow plant owes its genus name to Helen of Troy, while the specific epithet honors John Milton Biglow (1804-1878). Mr. Bigelow was an American surgeon and botanist on the Whipple Expedition. White Hawkweed Hieracium albiflorum Hook. Also called White Flowered Hawkweed. Native. Sunflower Family (Asteraceae). Another composite, a panicle of small heads bears ligulate flowers. The ligules (petal-like structures) are white and have 5 small lobes at the tip. Each ligule is accompanied by a yellow pistil in the center of the head. The phyllaries (sepal-like structures) are long, narrow and pointed with sparse, long hairs. Each head-bearing stem has a scale-like bract. The erect, green, stem is densely hairy along the lower portion. There may be a few alternate leaves along the lower part of the stem, but most of the leaves are basal. The leaves are mostly entire, but some may be toothed, and all are long-hairy. This perennial with milky sap is found in shaded areas. Look for it in July in wooded areas along the Pacific Crest Trail. You may find Siskiyou Penstemon (Penstemon anguineus) or Leafy Lousewort (Pedicularis racemosa) nearby. It is possible that some Native Americans used this plant and its coagulated juice as chewing gum. Hieracium is a Greek derivative of the word for hawk, and comes from the ancient legend that hawks ate the sap of these plants to sharpen their eyesight. The common name, Hawkweed, also arises from this legend. Albiflorum means white flower in Latin. 78 © Stanford University Press Plant Descriptions © Stanford University Press Whitney’s Bristleweed (Hazardia whitneyi var. discoidea) Tall Sneezeweed (Helenium bigelovii) White Hawkweed (Hieracium albiflorum) 79 Plant Descriptions Scouler’s Hawkweed Hieracium scouleri Hook. Also called Scouler’s Woolly Weed, and Hound’s Tongue Hawkweed. Native. Sunflower family (Asteraceae). You will find the hairy basal leaves of this plant as early as June 1st, long before the gold-yellow flowers appear in early July. The basal leaves are simple and entire, and exude a milky sap when broken. The dandelion-type “flowers” grow on branching stalks, with 2 to 5 flowering heads per stalk. If leaves are present on the stem, they are small, simple, and entire, and generally subtend a branch from the main stalk. The stems are covered with minute, white hairs that are not nearly as noticeable as the long hairs on the basal leaves and “sepals.” As this plant is in the Sunflower Family, what appear to be sepals are actually phyllaries, and collectively referred to as the involucre. These erect perennials can grow 10-25 cm tall. You can find Scouler’s Hawkweed around Rabbit Ears in early July, growing near Scarlet Gilia (Ipomopsis aggregata) and Rosy Pussytoes (Antennaria rosea). Hieracium is a Greek derivative of the word for hawk, referring to the ancient legend that hawks ate the sap of these plants to sharpen their eyesight. The common name, Hawkweed, also arises from this legend. Scouleri honors Dr. John Scouler (1804-1871), a surgeon, geologist, zoologist, and botanist from Glasgow, Scotland. He explored the coast flora of the Pacific Northwest for seven months in 1824-1825. Hairy Cat’s Ear Hypochaeris radicata L. Also called Rough Cat’s Ear. Non-native from Europe. Sunflower Family (Asteraceae). These tall, spindly, dandelion-like “flowers” are heads made up of many small, yellow, ray flowers, or ligulate flowers. Many of the outer ray flowers have a red-brown stripe on back. The sepallike structures, called phyllaries, are deep, dark green. Most of the leaves are basal, and they vary from entire to toothed, and all have stiff hairs. The lower portion of the stem may bear one or two stem leaves that would be entire and hairy. The upper portion of the stem may have very small, scale-like bracts that have a pointed, brown tip. The flowering stems are 15-60 cm tall and usually branch to bear more than one flowering head. Hairy Cat’s Ear blooms from mid-July through August along County Road 1151, often growing near Shasta Tansy Aster (Machaeranthera canescens var. shastensis) and Scouler’s Hawkweed (Hieracium scouleri). Hypochaeris is the ancient Greek name given by Theophrastus (370-285 B.C.), a disciple of Aristotle who established the vocabulary for plant morphology. His concepts remained basically unchanged until the advent of microscopes. Radicata is a combination of Greek and Latin words that means downward spoke or downward ray. Shasta Tansy Aster Machaeranthera canescens (Pursh) A. Gray var. shastensis (A. Gray) B. Turner Also called Shasta Hoary Aster. Native. Sunflower Family (Asteraceae). It appears that the yellow “flowers” of this perennial have no petals. That is because what appears to be one flower is not a flower at all. Rather, it is a head of many tiny flowers! Using your hand lens, you should be able to distinguish each tiny flower: a radiate corolla with 5 lobes and a long pistil extending from the center. The head of flowers is subtended by many green bracts, which are collectively referred to as the involucre. The involucre, which may be mistaken for sepals, has glandular hairs making them sticky to the touch. The severral flowering heads are borne at the tips of the many branches which are covered with dense, fine, grayish-white hairs. The alternate leaves have shallow teeth, especially toward the base of leaf. The uppermost leaves are glandular. Shasta Tansy Aster blooms from mid-July through mid-August along County Road 1151, and in dry, open areas along the Pacific Crest Trail. It is usually found below 6500 feet in elevation, and often grows next to Scarlet Gilia (Ipomopsis aggregata) and Hairy Cat’s Ear (Hypochaeris radicata). Derived from several Greek words, machaeranthera means sword-like anthers, obviously referring to the shape of the anthers. Canescens is Latin for becoming gray, referring to the grayish-white hairs on the stems. Shastensis is a Latinized version of Shasta, indicating the general range of this subspecies. 80 Plant Descriptions Scouler’s Hawkweed (Hieracium scouleri) Hairy Cat’s Ear (Hypochaeris radicata) Shasta Tansy Aster (Machaeranthera canescens var. shastensis) 81 Plant Descriptions Bolander’s Madia Madia bolanderi (A. Gray) A. Gray Also called Bolander’s Tarweed. Native. Sunflower Family (Asteraceae). 8-12 showy, deep yellow ray flowers surround many yellow disk flowers that have black anthers. This bold “flower” appears to have yellow petals and a center that is yellow and black-dotted. Below all the flowers are many green bracts that might be mistaken for sepals. These bracts are called phyllaries, and they are covered with glands that contain a sticky oil. The large, heavy flowering heads are borne at the terminus of several branches. The fairly stout stems are covered with hairs that are tipped with black glands that also contain the same sticky oil. Go ahead – touch the stem. Pretty sticky, eh? The leaves are opposite on the lower half of the stem and alternate above, and they vary from entire to shallowly toothed. Bolander’s Madia blooms from mid-July through mid-September on the south slopes in wet meadows and along streams. Nearby, you will find Oregon Saxifrage (Saxifraga oregana) and a little belly flower called Primrose Monkey Flower (Mimulus primuloides). Madia is a Chilean name because many of the plants in this genus grow in southwest South America. Bolanderi recognizes Henry Nicholas Bolander (1832-1897), a German-born American botanist who was known in his time as the “State Botanist for California.” For many years, he collected plants in the San Francisco area. Mountain Tarweed Madia glomerata Hook. Also called Stinking Tarweed. Native. Sunflower Family (Asteraceae). To experience everything this little flower has to offer, first you should touch the top of the stem around the base of the flowering head. You’ll notice that it is very sticky – the result of glands that contain sticky oils. Now smell your fingers. Yuck! Indeed, Stinking Tarweed is an appropriate name! The slender stem of this stinky little plant bears one to several flowering heads. The lower portion of the stem is soft-hairy to bristly and the upper portion, as you already experienced, is glandular. Often the stem is simple with no branches; however, on some plants the stems branch once or twice and each branch bears a flowering head. There are usually only two to four small yellow ray flowers that surround many tiny yellow disk flowers that have black anthers. So, the center of the flower appears to be black-dotted. The narrow, lanceolate leaves tend to be opposite on the lower half of the stem and alternate above. They can vary from entire to shallowly toothed. Mountain Tarweed is found on dry, gravelly soil on the south slopes. It is abundant along USFS Road 20 from the picnic area to Grouse Gap shelter. It blooms from early- to mid-July through midAugust, keeping company with Leafy Aster (Aster foliaceus var. lyallii) and Large-flowered Collomia (Collomia grandiflora). Madia is a Chilean name because many of the plants in this genus grow in southwest South America. Derived from a Latin word, glomerata means a ball, referring to the spherical flowering heads. Slender Tarweed Madia gracilis (Smith) Keck Native. Sunflower Family (Asteraceae). The slender stem of this graceful plant is highly branched bearing many flowering heads in the leaf axils. There are usually only two to four small yellow ray flowers that surround many tiny yellow disk flowers that have black anthers. Therefore, the center of the flower appears to be black-dotted. The calyxes are densely hairy with black glandular tips. When the fruits mature, the papery, cream-colored chaff scales remain attached to the receptable. The alternate, entire, narrow leaves vary in shape from linear, to elliptic, to oblanceolate and are white hairy. The lower portion of the stems are bristly to long soft-hairy and the upper portion is covered with hairs that have a black, glandular tip. Slender Tarweed is found in many different habitats. Look for it along USFS Road 20 from the picnic area to Grouse Gap shelter. It blooms in September, next to Changeable Phacelia (Phacelia mutabilis) and Leafy Aster (Aster foliaceus var. lyallii). Many of the plants in this genus grow in the area of Chile in South America, therefore Madia is a Chilean name. Gracilis is derived from the Latin word for slender, referring to the stems. 82 © Stanford University Press Plant Descriptions Bolander’s Madia (Madia bolanderi) © Stanford University Press Mountain Tarweed (Madia glomerata) Slender Tarweed (Madia gracilis) 83 Plant Descriptions Alpine Lake Prairie Dandelion Nothocalais alpestris (A. Gray) Chambers Native. Sunflower Family (Asteraceae). As the name suggusts, this flower superficially looks like a scrawny Dandelion (Taraxacum officinale). Bright yellow flowering heads, made up of 13 or more ligulate flowers, sit atop leafless stems. The phyllaries, green structures below the flowers that could be mistaken for sepals, are glabrous, and evenly and finely purple-dotted. The many basal leaves vary in shape from linear to widely oblanceolate (widest toward the tip), and also vary from entire to toothed to pinnately lobed. Alpine Lake Prairie Dandelion blooms early in the subalpine season, and are short-lived. They can be found usually from mid-June through late-June, on the dry, gravelly south slopes, perhaps blooming next to California Indian Potato (Orogenia fusiformis) or Spreading Phlox (Phlox diffusa). Nothocalais is derived from Greek and means false Calais; the derivation and meaning of Calais is unknown. Alpestris is New Latin for mountains, referring to the habitat of this flower. Western Coneflower Rudbeckia occidentalis Nutt. var. occidentalis Native. Sunflower Family (Asteraceae). Wow! A native coneflower on Mount Ashland! The stout stem is usually simple, but may have one or two branches near the top. Each branch bears one terminal large flowering head that appears to be missing something. This plant has no petal-like structures, that is there are no ray flowers. Only dark purple-brownish disk flowers are held on a cone-shaped receptacle. Beneath the disk flowers, the long, narrow phyllaries are reflexed. Don’t be fooled – these are not sepals. This perennial has basal leaves and alternate stem leaves that vary from entire to shallowly toothed, and also vary from glabrous to hairy. The leaves can be up to 30 cm long, while the entire plant usually grows from 120 – 150 cm. I found Western Coneflower only in one wet meadow along the Pacific Crest Trail. If you locate the trailhead of the PCT on USFS Road 20, just beyond the Bull Gap trailhead, and hike south along the trail, after a short hike through the woods, you’ll come to an open wet meadow. Western Coneflower grows right next to the trail through this wet meadow, along with Red Elderberry (Sambucus racemosa) and American Sawwort (Saussurea americana). O.J. Rudbeck (1630-1702) and O.O. Rudbeck (1660-1740) were professors of botany at Upsala Sweden. They are both honored in this genus name. Occidentalis is derived from the Latin word meaning western, referring to the western range of this flower. American Sawwort Saussurea americana D. Eaton Native. Sunflower Family (Asteraceae). One fairly stout stem bears 3 to 12 small flowering heads with purple disk flowers. The terminal flowering heads have no petal-like structures, or ray flowers. The phyllaries are pale green with dark tips and have sparse woolly hairs. The stem also has sparse woolly hairs, and is very leafy, with alternate leaves from 5-15 cm long and coarsely toothed. This perennial reaches heights of 90-150 cm. You will find American Sawwort in the wet meadows on the south slopes of Mount Ashland. It blooms in August in the company of Mountain Larkspur (Delphinium glaucum) and California False Hellebore (Veratrum californicum). Theodore and Horace Saussure, 18th century Swiss naturalists, are remembered in this genus name. Americana is derived from Latin and means of or from America. 84 Plant Descriptions Alpine Lake Prairie Dandelion (Nothocalais alpestris) Western Coneflower (Rudbeckia occidentalis) American Sawwort (Saussurea americana) 85 Plant Descriptions Silky Raillardella Raillardella argentea (A. Gray) A. Gray Native. Sunflower Family (Asteraceae). Silky Raillardella is an inconspicuous, unimpressive plant (as far as appearances go, any way). The yellow flowering heads are made up of disk flowers only, the “flower,” which is actually a head of many tiny flowers, appears to have no “petals.” The leafless flowering stem rises only an inch or two above the mat of basal leaves. The leaves are entire and completely covered with long, fine, soft white hairs. This low-growing, mat-forming perennial grows only on the summit of Mount Ashland around the top of the bowl between the weather station buildings. You’ll find it blooming from mid-July through mid-August near Jayne’s Canyon Buckwheat (Eriogonum diclinum) and fluffy fruiting heads of Drummond’s Anemone (Anemone drummondii). When you consider the environmental implications of this flower’s habitat, the plant no longer seems unimpressive. Rather, it is absolutely remarkable that any plant can survive under these conditions. In the late fall and early winter, before the snows begin, this section of Mount Ashland is subject to high velocity winds. These winds not only provide a mechanical challenge for the plants to overcome, but it also dries out the soil. Later in the winter, the area is covered by a large amount of snow, and on top of the snow, skiers line up for the exciting rush down “the bowl.” When the snows melt and spring begins, this is the first area to dry out as the winds kick up again. Then the brutal sun of summer adds not only to the evaporative effect, but showers the plants with damaging ultra-violet light. Silky Raillardella is, in fact, quite an impressive little flower. Riallardella is of Latin derivation and means little raillardia. The derivation and meaning of raillardia is uncertain. Argentea is derived from the Latin word meaning silver or silvery, referring to the leaves that are covered with long, white hairs, making the leaves look silvery. Western Butterweed Senecio integerrimus Nutt. var. major (A. Gray) Cronq. Also called Columbia Ragwort, Tower Butterweed, and Tall Western Senecio. Native. Sunflower Family (Asteraceae). Small yellow radiate flowers consist of 8-13 ray flowers, which superficially resemble petals, surrounding 20-50 tiny disk flowers in the center. The ray flowers vary from bright yellow to off-white, while the disk flowers are always yellow. Phyllaries, structures that superficially resemble sepals, are green with a minute black tip. (The black-tipped phyllaries are the best characteristic for determining whether a composite plant is in the genus Senecio.) The erect unbranching stem yields 6-20 of these flowering heads in a corymb. The leaves are generally lanceolate, and the margins of the alternate leaves vary from entire to wavy to shallowly and coarsely toothed. The leaf stem is small, if it is there at all. Stem and leaves are soft hairy. Western Butterweed is a biennial or perennial and grows 20-50 cm. You will find Western Butterweed in dry soil usually at forest edges, but sometimes in open meadows. It blooms through June and early July, along with Meadow Larkspur (Delphinium nuttallianum) and Ballhead Sandwort (Arenaria congesta var. congesta). Senecio is a Latin derivative meaning old man, referring to the white pappus on each tiny flower that can only be viewed with a hand lens or magnifying glass after plucking an individual flower from the head. Integerrimus is based on the Latin word for whole, complete, or entire. This refers to the leaf margins which are usually entire. 86 Plant Descriptions Silky Raillardella (Raillardella argentea) Western Butterweed (Senecio integerrimus var. major) 87 Plant Descriptions Arrowhead Butterweed Senecio triangularis Hook. Also called Arrow-leaf Ragwort, Arrowleaf Groundsel, and Spear-head Senecio. Native. Sunflower Family (Asteraceae). The yellow flowering heads of Arrowhead Butterweed look just like the flowering heads of Western Butterweed (S. integerrimus). The leaf shape and habitat are the primary differences between the two plants. Western Butterweed has lanceolate leaves and is found in dry soil, while Arrowhead Butterweed has distinctly triangular-shaped leaves (hence, the name Arrowhead) and is found in wet soil. The leaf margins of Arrowhead Butterweed are usually coarsely toothed, but on some plants the leaves could be entire. The erect stem is glabrous, very leafy, and reaches heights of 30-85 cm. Senecio is a Latin derivative meaning old man, referring to the white pappus on each tiny flower that can only be viewed with a hand lens or magnifying glass after plucking an individual flower from the head. Triangularis is derived from Latin and means three corners, obviously referring to the shape of the leaves. Canada Goldenrod Solidago canadensis L. ssp. elongata (Nutt.) Keck Also called Graceful Goldenrod, and Meadow Goldenrod. Native. Sunflower Family (Asteraceae). Many small yellow flowering heads are arranged in a long, diamond shaped panicle: the panicle is widest in the center and tapered at the top and bottom. The flowering heads are only 3-5 mm wide, and are displayed at the top of a tall, leafy stem. Alternate leaves are irregularly toothed and prominently three-veined. The erect stem stands 30-150 cm tall. I saw only one or two Canada Goldenrod plants blooming in the open meadow close to the picnic area and above USFS Road 20 from mid-July through mid-August. Seep Monkey Flower (Mimulus guttatus) may be found in the nearby wet area, while Spreading Ground Smoke (Gayophytum diffusum) may be found in the surrounding dry soil. Goldenrods are often blamed for hay fever, but this common summer ailment is more likely the result of pollen from the inconspicuous flowers of Ragweed (Ambrosia sp.). Old herbalists believed Goldenrod to have curative properties. Although this has since been disproved, the Latin name remained the same: solidus which means whole, ago which means to make, and so…to cure. Several species of solidago are used to make excellent yellow dyes. From Latin, solidago means to cure, referring to the healing properties of some of the plants in this genus. Canadensis is New Latin meaning of Canada, referring either to the location of the first plant of this species that was identified, or referring to the preferred range of this species. Elongata means elongated and probably refers to the long inflorescence. Yellow Salsify Tragopogon dubius Scop. Also called Yellow Goatsbeard. Non-native from Europe. Sunflower Family (Asteraceae). A solitary yellow, radiate flowering head is snuggled inside the narrow, green, lanceolate phyllaries that are longer than the yellow ligules. These ligulate flowering heads are made up of only ray flowers - there are no disk flowers – with black anthers. The alternate leaves are grasslike: long, linear, and entire. The base of each leaf wraps half way around the unbranching stem. The stem, leaves, and phyllaries are glabrous and glaucous, except at the stem nodes where a patch of woolly hairs marks the spot. The erect stem widens at the top, just below the flowering head. When broken, the stem and leaves exude a milky sap. This biennial or perennial blooms mid-July through late-August in dry, open meadows on the south slopes. If you look around, you may also see Large-flowered Agoseris (Agoseris grandiflora) or Leafy Aster (Aster foliaceus var. lyallii). Tragopogon comes from Greek for goat’s beard, possibly referring to the thin, tapering, tufted, grass-like leaves. It is uncertain to what the name, dubius, refers but it is Latin for doubtful. 88 Plant Descriptions Arrowhead Butterweed (Senecio triangularis) Canada Goldenrod (Solidago canadensis) Yellow Salsify (Tragopogon dubius) 89 Plant Descriptions Dandelion Taraxacum officinale Wigg. Also called Lion’s Tooth, Blowball, Priest’s Crown, Swine’s Snout, Fortuneteller, Doonhead Clock, and Wet-a-Bed. (Asteraceae). Dandelions are so common and familiar that they hardly need description. Suffice it to say they are bright yellow flowers with lobed basal leaves that resemble lion’s teeth. Dandelions are first mentioned in 1513 while Henry VIII was on the throne in England. King Henry’s gardener maintained a perfectly manicured lawn and bowling green at Hampton Court Palace (Ward-Harris, 1983). The gardener hand cut the greens with a scythe, and pulled out dandelions by hand. Dandelions have been eaten and used for medicines for centuries. Dandelion is used for so many medical treatments that many herbalists refer to this plant as “nature’s medicine cabinet.” During World War II, dandelion roots were ground up and roasted and used as a substitute for coffee. The plant is still cultivated in France as a salad vegetable. Dandelion leaves contain iron, phosphorus, sodium, and large amounts of calcium and potassium. They are rich in vitamins B and C, and have more vitamin A than any garden plant. The flowers are used to make a delicious sweet wine (I’ve tried it!) and beer. Gypsies used the roots for a magenta dye, and Scotsmen made the same dye for their tartans using the whole plant, or just the roots. Experiments among weavers in North America have shown that dandelions on this side of the Atlantic yield only a yellow dye. Taraxacum appears to stem from the Greek word, tarassein, which means to stir up, possibly relating to the plant’s medicinal purposes. The specific name, officinale, is equivalent to early English “medicinal” since dandelion leaves make a tea for coughs, fevers, and internal inflammations. The root is diuretic, hence one of the French names, Pissenlit – Wet-a-Bed. Barberry Family (Berberidaceae) Oregon Grape Mahonia nervosa (Pursh) Nutt., or Berberis nervosa Pursh Also called Dwarf Oregon Grape, Long-leaved Oregon Grape, Little Oregon Grape, Mahonia, Holly Grape, Barberry, and Cascade Barberry. Native. Barberry Family (Berberidaceae). A shrub with holly-like leaves, Oregon Grape is one of a kind on Mount Ashland. The yellow, radial flowers have 6 petals in two whorls of three and 6 stamens. Beneath the petals are 9 green sepals in three whorls of three. The alternate, bright green, waxy leaves are compound with 7 to 23 spine-toothed leaflets. The bright, yellow flowers appear in late June, and the blue-purple berries ripen in August. This erect shrub is found mostly on the north slopes at the forest edge. Nearby, look for Golden Chinquapin (Chrysolepis chrysophyllus) and Greenleaf Manzanita (Arctostaphylos patula). The berries of this plant are edible and make good jam. A medicinal tea can be made from the root bark, and a yellow dye can be made from the entire root. The root bark tea can be mixed with a tea made from Prince’s Pine (Chimophila umbellata) and used as a remedy for kidney problems. The roots of some of the plants in this genus are often toxic, with spines that may inject fungal spores into the skin. The meaning and derivation of the genus name, Mahonia, is uncertain. Nervosa is Latin for vein or nerve. This could refer to the obvious leaf veins, or this specific epithet could imply that the toxic substances in the roots affect the nervous system. Berberis is the Latin derivation of the ancient Arabic name for Barberry. 90 Plant Descriptions Photo by Br. Alfred Brousseau, St. Mary’s College Photo by Charles Webber, California Academy of Sciences Photo by Br. Alfred Brousseau, St. Mary’s College Dandelion (Taraxacum offinale) Oregon Grape (Berberis nervosa) 91 Plant Descriptions Borage Family (Boraginaceae) Weak-stem Cryptantha Cryptantha flaccida (Lehm.) E. Greene Native. Borage Family (Boraginaceae). The coiled cymes are made up of many small, white flowers. Each radial flower has 5 sepals that are strongly bristly-hairy, 5 petals, 5 stamens, and 1 pistil. There is only fruit, called a nutlet, per flower. The nutlets have smooth, shiny backs and a closed groove above the scar. The lowermost leaves are more or less opposite, while the upper leaves are alternate. All leaves are narrow, linear or oblanceolate, and bristly-hairy; bristles on lower surface are bulbous-based. The branches stem is also bristly-hairy, and the bristles are mostly spreading, while some are ascending. This annual grows 5-30 cm tall. Look for Weak-stem Cryptantha in dry, open areas from early- to mid-July, with Mountain Tarweed (Madia glomerata) and Palmer’s Catchfly (Silene bernardina). In Greek, cryptantha means hidden flowers. This refers to the flowers of some species in this genus that are bud-like, un-opening, and usually self-pollinating. Flaccida is derived from the Latin word for flabby or feeble, referring to the weak stem of this slender annual. Pinewoods Cryptantha Cryptantha simulans E. Greene Native. Borage Family (Boraginaceae). Dainty, white, inconspicuous flowers adorn this annual. The inflorescence is an open cyme. The small radial flowers have 5 sepals that are strongly bristly-hairy, 5 petals that each measure 1-2 mm wide, 5 stamens, and 1 pistil. The mature fruits of flowers in the Borage family are referred to as nutlets. Pinewoods Cryptantha has 4 nutlets, each with a granular back and a closed groove above the scar. The lowermost leaves are more or less opposite, while the upper leaves are alternate. They are entire, and narrow with bristly hairs that are ascending to appressed. The stem, which is branched throughout, usually reaches 5-40 cm, and has bristly hairs that are ascending to appressed. You will need your hand lens to clearly see the hairs. Pinewoods Cryptantha blooms in dry, open areas from mid- to late July. Look for it along USFS Road 20 near the ski area parking lot, with Alice’s Daisy (Erigeron alicea) and Pineapple Weed (Chamomilla suaveolens) growing nearby. In Greek, cryptantha means hidden flowers. This refers to the flowers of some species in this genus that are bud-like, un-opening, and usually self-pollinating. Simulans is derived from the Latin for a likeness, or to imitate. It is uncertain to what this description refers. Torrey’s Cryptantha Cryptantha torreyana (A. Gray) E. Greene Also called Torrey’s Cat’s Eye. Native. Borage Family (Boraginaceae). Tiny white flowers are borne in a cyme that is coiled in flower and open in fruit. The radial flowers have 5 sepals that are bristly-hairy, 5 petals that each measure about 1 mm wide, 5 stamens, and 1 pistil. When the 4 nutlets are ripe, they each measure a little longer than 1 mm, have a shiny back, and a raised, closed groove above the scar. The entire nutlet is brown, mottled with black. The alternate leaves are entire. The branched stem and leaves have bristly hairs that are spreading to ascending. This annual grows 10-40 cm. Torrey’s Cryptantha is found in dry to moist open areas. Look for it especially around the ski lodge, growing next to Woolly Mullein (Verbascum thapsus) and Silverleaf Phacelia (Phacelia hastata). In Greek, cryptantha means hidden flowers, referring to the flowers of some species in this genus that are bud-like, un-opening, and usually self-pollinating. John Torrey (1796-1873), an American botanist who explored the west coast of North America and authored 5 major floras is honored in this species name. 92 Plant Descriptions © Stanford University Press Weak-stem Cryptantha (Cryptantha flaccida) © Stanford University Press Photo by Donald Myrick, California Academy of Sciences Pinewoods Cryptantha (Cryptantha simulans) Torrey’s Cryptantha (Cryptantha torreyanna) 93 Plant Descriptions Jessica’s Stickseed Hackelia micrantha (Eastw.) J. Gentry Also called Jessica’s Tickweed, and Jessica Stick-tight. Native. Borage Family (Boraginaceae). These flirty little flowers have five petals that are bluish-lavender, turning pink with age. The yellow center of this regular flower is surrounded by a white, textured ring. The inflorescence is a terminal panicle, with axillary flowers on the upper part of the stem. The simple, entire leaves are alternate and have no petiole. The stem is highly branched, and usually stands 25-60 cm tall, and usually grows in the shade of trees with Meadow Larkspur (Delphinium nuttallianum) and Showy Jacob’s Ladder (Polemonium californicum). Look for this perennial around Rabbit Ears and the Grouse Gap Shelter mid to late June, and in wetter areas through the end of July. The common name for this plant, Stickseed, is a result of the barbed spines that form a ring around the seeds. Because of these barbed spines, the seeds stick to everything! J. Hackel, a Czeck botanist born in 1783, lends his name to this genus. Appropriately, micrantha means small flower in Greek. Mustard Family (Brassicaceae) Spreading Pod Rockcress Arabis xdivaricarpa Nelson (pro sp.) [drummondii x holboellii] Native. Mustard Family (Brassicaceae). As all members of the Mustard Family, these radial flowers have four petals that are arranged in a cross-like fashion. The petals are pale pink to lavender. The inflorescence is a raceme, which typically matures from bottom to top. Because of this, you will often see flowers near the top of the plant and fruits below the flowers. The fruits of this perennial are elongated siliques, and they point upward in a spreading manner. The leaves are alternate and toothed to entire. The lower leaves are often sagittate and clasping the stem. The erect stem and leaves are sparsely hairy with branched hairs. This plant usually grows to 30-45 cm. Look for this one in dry, open areas around Rabbit Ears and upward to the summit, and on the upper portion of the ski slopes. This one blooms in June and early July and fruits can be seen through mid-August. Because the flower petals of most members in this family are arranged similar to a Maltese cross, the old name for this family was Cruciferae. Arabis is Latin, meaning of Arabia. Divaricarpa is a combination of Latin and Greek roots that mean “spreading fruits,” because the mature fruits are only slightly erect, spreading from the stem. The “x” in front of divericarpa indicates that this plant is a hybrid between two other species – Arabis drummondii and Arabis holboellii. Tower Rockcress Arabis glabra (L.) Benth. var. glabra Also called Smooth Rockcress. Native. Mustard Family (Brassicaceae). These radial flowers have 4 green sepals, 4 petals arranged similarly to a Maltese cross, 6 stamens (4 long and 2 short) and 1 pistil. The petals vary in color from cream to lavender. The alternate stem leaves are sessile and clasp the stem, and vary from lobed (sagittate) to entire to toothed. The stout stem is usually simple, rarely branched, and stands erect 40-120 cm tall. This biennial grows in open meadows. Look for it along the Pacific Crest Trail on the south side of Grouse Gap Shelter. Look for other Rockcresses (Arabis sp.) growing nearby, along with Spreading Ground Smoke (Gayophytum diffusum). Arabis is derived from the Latin word for “of Arabia.” Glabra is derived from the Latin word for smooth, referring to the absence of hairs. 94 © Stanford University Press Plant Descriptions Jessica’s Stickseed (Hackelia micrantha) © Stanford University Press Photo by Lee Dittman Spreading Pod Rockcress (Arabis xdivaricarpa) Tower Rockcress (Arabis glabra var. glabra) 95 Plant Descriptions Holboell’s Rockcress Arabis holboellii Hornem. var. retrofracta (Graham) Rydb. Also called Second Rockcress. Native. Mustard Family (Brassicaceae). The pale pink, lavender or white flowers of Holboell’s Rockcress are radial with four petals. If you look in the center of the flowers, you will find six stamens – four long and two short. This is a characteristic of the Mustard Family. The fruits of this perennial are strongly reflexed (pointing downward). The alternate sessile leaves are entire to shallowly toothed with minute multi-branched hairs hairs. The erect stem, also with multi-branched hairs, grows to 20-45 cm. Holboell’s Rockcress can be found in dry, open areas along the Pacific Crest Trail, and from Rabbit Ears to the summit. It blooms in June and the long, thin fruits, called siliques, are obvious through mid-August. Arabis is Latin, meaning of Arabia. Holboell's Rockcress was first described for science in 1828 by Jens Wilken Hornemann (1770-1841), professor of botany at Copenhagen. He dedicated the specific epithet to the eminent Danish ornithologist Carl Peter Holboell (1795-1856). Retrofracta is Latin for broken backwards, referring to the mature fruits which point down to the ground. Lyall’s Rockcress Arabis lyallii S. Watson var. lyallii Native. Mustard Family (Brassicaceae). Lyall’s Rockcress has radial flowers arranged in a raceme. The flowers have four spoon-shaped petals that vary in color from rose to lavender. When the fruits mature, they are straight and glabrous, and are held erect to spreading. These fruits are wider than most Rockcresses, and the seeds inside have a narrowly winged margin. The branched stems of this perennial are glabrous, while the alternate, sessile leaves have tiny multibranched hairs. These hairs can usually only be seen with a hand lens. Leaf margins are entire. This generally erect plant is low-growing, measuring less than 15 cm. Lyall’s Rockcress generally is found in rocky areas, and the best place to find it on Mount Ashland is in the bowl area on the north slopes, blooming in June. In and around the bowl, you might also find Drummond’s Anemone (Anemone drummondii) and Davidson’s Penstemon (Penstemon davidsonii). Arabis is Latin, meaning of Arabia. David Lyall (1817-1895) is remembered in this specific epithet. Lyall was a British surgeon and botanist who collected many valuable plant specimens from Antarctica, New Zealand, and western North America. Pioneer Rockcress Arabis platysperma A. Gray var. platysperma Also called Flat-pod Rockcress. Native. Mustard Family (Brassicaceae). These four-petaled flowers vary in color from pink to white. The few (if any) stem leaves are sessile and entire. The upper leaves are glabrous. The many basal leaves are entire and oblanceolate (widest just below the tip). The erect seedpods are broad (up to 5 mm wide), and are more like a legume than a typical silique. The base of these plants is woody. This one will be found only on the very dry, rocky slopes of the summit and in some places along the County Road 1151 between mile 8 and the ski lodge parking lot. Pioneer Rockcress blooms in June but the seedpods will be obvious through midAugust. Arabis is Latin, meaning of Arabia. Platysperma is Greek for broad, flat seeds. Indeed, this plant is most noticeable when the fruits are mature, and the wide, flat seed pods are obvious. 96 Photo by Br. Alfred Brousseau, St. Mary’s College Plant Descriptions Photo by Bob Vos Holboell’s Rockcress (Arabis holboellii var. retrofracta) Lyall’s Rockcress (Arabis lyallii) Pioneer Rockcress (Arabis platysperma) 97 Plant Descriptions Shepherd’s Purse Capsella bursa-pastoris (L.) Medikus Non-native from Europe. Mustard Family (Brassicaceae). A flower of disturbed sites, you will only find this annual in the gravel around the ski lodge. The small white to lavender flowers are radial with four petals. The inflorescence is a raceme. The alternate leaves are entire with bases that clasp the stem. Basal leaves are shallowly to deeply lobed with short hairs. The fruits of this plant are very distinctive – they are more or less heart shaped and roughly resemble the purse of a shepherd. In Europe, this plant has been widely used for its edible leaves and peppery-tasting seeds. Capsella is Latin for little box; bursa is Latin for purse, and pastoris is Latin for shepherd, all referring to the shape of the fruits. Nuttall’s Toothwort Cardamine nuttallii E. Greene var. gemmata (E. Greene) Rollins Also called Bittercress. Native. Mustard Family (Brassicaceae). Toothwort has four petals arranged radially. Their color varies from pale pink to lavender, and rarely white. The stem leaves are alternate and pinnately compound, with five to seven narrow leaflets. The basal leaves are quite different from the stem leaves. They are compound with three leaflets. Each leaflet is shallowly lobed, and the lower surface is purplish, while the upper surface is green. Basal leaves are not always obviously attached to the plant because they are attached to the underground rhizome. Fruits are slender, elongated siliques, and often appear with the flowers – flowers at the top of the raceme, fruits at the bottom. Look for Toothwort in moist, wooded areas on the Knoll. Cardamine is Greek for a species of cress. Nuttallii is derived Thomas Nuttall’s name. Nuttall (1786-1859) explored the largest portion of the U.S. than any other botanist in the 1800s. Originally from England, Nuttall came to the U.S. in 1807 and described many new plant species for North America. Gemmata is Latin for a bud. The common name, Toothwort, comes from the old Latin name for the genus, dentaria, which means tooth and refers to the tooth-like underground corms. Wort is an old English word for plant. Mountain Tansy Mustard Descurainia incisa (A. Gray) Britton ssp. incisa Native. Mustard Family (Brassicaceae). Many small, yellow flowers grow in tight racemes that elongate as the fruits mature. Each radial flower has four petals that surround six stamens – four long and two short. The long, slender fruits are called siliques. Because the flowers mature from the bottom of the raceme toward the top, you will often find mature fruits below the bright flowers. The green leaves are pinnately lobed or compound and grow alternately along a reddish-brown stem. The stem can be simple or branched. Tansy Mustard is most often seen in shaded areas and along the borders of wet areas on the south slopes. Growing nearby, you may find Orange Agoseris (Agoseris aurantiaca) or Siskiyou Penstemon (Penstemon anguineus). This genus is named after the French botanist, F. Descourain (1658 – 1740). Incisa is derived from Latin and means cut into, probably referring to the pinnately lobed or compound leaves. 98 Plant Descriptions © Stanford University Press Shepherd’s Purse (Capsella bursa-pastoris) Nuttall’s Toothwort (Cardamine nuttallii var. gemmata) Mountain Tansy Mustard (Descurainia incisa var. incisa) 99 Plant Descriptions Western Wallflower Erysimum capitatum (Douglas) E. Greene ssp. capitatum Also called Wallflower, Rough Wallflower, Sanddune Wallflower, and Douglas’s Wallflower. Native. Brassicaceae (Mustard family). These bright orange-gold-yellow flowers are radial with four petals and six stamens – four long and two short. The inflorescence is a terminal raceme. Most likely, you will find flowers along the upper portion of the raceme and ripening fruits along the lower portion of the raceme. The fruits are referred to as siliques, and are characteristic of the Mustard Family. The stem leaves are simple, alternate and very shallowly toothed, with the upper leaves sometimes appearing entire. The erect stem is round with minute bristly hairs, so that the plant feels rough when you rub fingers down the stem. This biennial grows to about 25-45 cm tall. Look for this flower around Rabbit Ears through June and July. In Greek, erysimum means to help, referring to the many medicinal species in this genus. Capitatum is Latin for head. Virginia Pepperweed Lepidium virginicum L. var. virginicum Also called Peppergrass. Subspecies probably not native to Oregon. Mustard Family (Brassicaceae). Small white flowers with four petals grow in a tight raceme that elongates as the fruits mature. The fruits are deep magenta to purple, circular, but flat in cross section. The basal leaves are fairly large and pinnately lobed to dissected. The alternate stem leaves are small and entire. The highly branched stems grow to 60 cm. I have found this annual growing only near the wooden fence along the parking lot at the ski area. Virginia Pepperweed blooms in early July. Growing nearby, look for another plant that is only found near the ski area parking lot, Bushy Blazing Star (Mentzelia dispersa), and the tiny belly flower, Red Sand Spurry (Spergularia rubra). Lepidium is Greek for little scale, referring to the shape of the fruit. Virginicum is of Greek derivation and means of Virginia. Probably, this flower was first identified in or near the state of Virginia. Bellflower Family (Campanulaceae) California Harebell Campanula prenanthoides Durand Also called California Bluebell. Native. Bellflower Family (Campanulaceae). Blue flowers are hard to find on Mount Ashland, but here is one that brightens the woods. Bright blue bell-shaped flowers have five reflexed petals that are backed by five green spreading sepals. The radial flower has five stamens and one long, blue pistil that is greatly exserted. California Harebell shows off its flowers in a loose, sometimes drooping raceme. Below the inflorescence, the linear, toothed leaves have a very short petiole (less than 5 mm), or none at all. The leaves are usually 1-6 cm long. The reclining to erect stem grows to 20-60 cm, and the leaves and stem can be glabrous or short-hairy. This perennial is found in dry, semi-open woods. Look for it on north slopes along the Bull Gap Trail in August. Rattlesnake Plantain (Goodyera oblongifolia) or White-veined Wintergreen (Pyrola picta) may be blooming nearby. Campanula is Latin for little bell, and is indicative of the shape of the flower. Prenanthoides is Greek for drooping flower, describing how the flower grows on the stem. 100 © Stanford University Press Plant Descriptions Western Wallflower (Erysimum capitatum) Photo by Bob Vos Virginia Pepperweed (Lepidium virginicum var. virginicum) California Harebell (Campanula prenanthoides) 101 Plant Descriptions Pale Bellflower Campanula scouleri A. DC Also called Scouler’s Bluebell. Native. Bellflower Family (Campanulaceae). Pale Bellflower has pale blue, to whitish bell-shaped flowers with 5 reflexed petals. Behind the petals are five green spreading sepals. The center of the flower bears five stamens and one strongly exserted pistil. One to five flowers are displayed on a one-sided raceme. The alternate leaves are sharply and coarsely toothed. The lower leaves are broadly oval or egg-shaped with long stalks, while the upper leaves become increasingly narrower and less stalked. The slender stem is either reclining or erect and usually 20-30 cm long. This perennial is generally found in dry woods. On Mount Ashland, look for it in the forest of the north slopes, perhaps along the Bull Gap Trail. It usually blooms around August, and may be found near its close relative, California Harebell (C. prenanthoides), or perhaps Pipsissewa (Chimaphila umbellata). Pale Bellflower is similar to California Harebell (C. prenanthoides), but there are some characteristics that make it easy to distinguish the two. The flowers of Pale Bellflower are a lighter blue than those of California Harebell. The leaves of Pale Bellflower are broad and have a long stalk, while those of California Harebell are narrow and more or less sessile. Campanula is Latin for little bell, and is indicative of the shape of the flower. Scouleri venerates Dr. John Scouler (1804-1871), a surgeon and naturalist from Glasgow, Scotland. He came to the Pacific Northwest in 1824 as the ship surgeon on a Hudson Bay Company ship. He collected plants in the Oregon and Washington area with David Douglas from 1824-1825. Honeysuckle Family (Caprifoliaceae) Twinflower Linnaea borealis L. var. longiflora Torrey Also called American Twinflower. Honeysuckle Family (Caprifoliaceae). Twinflower shyly displays two nodding flowers at the forked tip of a slender stem. The bellshaped, radial flowers have five lobes that can vary in color from white to pale pink to red. Small, opposite, evergreen leaves that are slightly shiny and sticky looking are at the base of the flowering stem. The margins of the oval leaves are primarily entire, but some may have a few shallow teeth. The flowering stems stand 5-20 cm and arise from a slender trailing runner. When the fruits mature, they form sticky burrs that are easily dispersed by animals and humans. Look for Twinflower in the moist, coniferous woods on the north slopes at lower elevations. One source claims that Native Americans made a decoction of the leaves to treat colds, but I can’t imagine anyone bold enough to deface this beauty by stealing its leaves. Gronovius, one of Carl Linnaeus’s teachers, honored Linnaeus (Carl von Linne, 1707-1778) by naming this flower after him because he knew this was Linnaeus’s favorite flower. Linnaeus is responsible for developing our current taxonomic scheme for naming plants. Borealis comes from Greek meaning north or northern, referring to the circumboreal range this plant. Twinflower can be found in Sweden, Russia, China, Alaska, and the Pacific Northwest. 102 Photo by Bob Vos Plant Descriptions Photo by Bob Vos Pale Bellflower (Campanula scouleri) Twinflower (Linnaea borealis) 103 Plant Descriptions Orange Honeysuckle Lonicera ciliosa (Pursh) Poiret Also called Trumpet Honeysuckle, Western Trumpet Honeysuckle, and Northwest Honeysuckle. Honeysuckle Family (Caprifoliaceae). Branch ends bear whorls of orange-yellow narrowly trumpet shaped flowers. Five lobes flare at the end revealing 5 stamens and 1 pistil that are slightly exserted from the petals. If you look inside the flower tube, you will discover a dense mess of tiny hairs. The opposite leaves have entire margins that are also covered with very fine hairs, and the end pair of leaves on each twig are united to form one round leaf that completely encircles the stem. The woody branches of this shrub are trailing or high climbing. In its effort to reach the light, Orange Honeysuckle often climbs into the tree tops, leaving behind scars of its stranglehold on the trees to which it clings. You will probably find Orange Honeysuckle at lower elevations in the forest. As expected, hummingbirds and swallowtail butterflies are attracted by the orange color, and are able to reach the nectar with their long tongues and proboscis. Adam Lonitzer, a 16th century German botanist, is honored by the generic name. Ciliosa means hairy or ciliate, referring to the hairs inside the flower and on the leaf margins. Wedded Honeysuckle Lonicera conjugialis Kellogg Also called Purple-flower Honeysuckle. Native. Honeysuckle Family (Caprifoliaceae). Wedded Honeysuckle is an erect shrub with small, paired, dark red flowers. The axillary flowers are bilateral with 4 lobes on the top lip and one down-turned lobe on the bottom lip. The style and anthers are exserted. Because the ovaries are fused, the two bright red fruits are also fused; hence, the name Wedded Honeysuckle. The opposite leaves are elliptic to round, and entire with hairs that are only visible through magnification. This shrub can be found along USFS Road 20 between the picnic area and Rabbit Ears. It flowers in May, and is often accompanied by Bitter Cherry (Prunus emarginata) and Wax Currant (Ribes cereum). Although none of the Oregon species of Lonicera have been reported as poisonous, they should be regarded with suspicion as berries of other species have caused fatalities. Adam Lonitzer, a 16th century German botanist, is honored by the generic name. Conjugialis is derived from the Latin word conjug, which means joined together, referring to the ovaries and fruits that are fused together. Blue Elderberry Sambucus mexicana C. Presl Native. Honeysuckle Family (Caprifoliaceae). At first glance, this deciduous shrub could be confused with Cascade Mountain Ash (Sorbus scopulina). The inflorescence of Blue Elderberry is a flatted-topped panicle, similar to that of Cascade Mountain Ash, but the flowers and leaves are slightly different. The flowers of Blue Elderberry are radial, white or cream with five petals, and protruding from the center are only five stamens (Cascade Mountain Ash has 15-20 stamens). The ripe fruits of Blue Elderberry are a glaucous blue, while the fruits of Cascade Mountain Ash are reddish-orange. The leaves are opposite and pinnately compound with 3-9 fairly large leaflets, while the leaves of Cascade Mountain Ash are alternate with 9-11 small leaflets. The margins of the leaflets of Blue Elderberry are irregularly toothed. Look for Blue Elderberry above 6500 feet along the road to Grouse Gap. It is usually in the middle of dry, open meadows. It blooms June through August, and the fruits ripen from August through September. Raw fruits are toxic if eaten in great quantities, but the fruits are non-toxic and quite tasty if cooked. The genus name, Sambucus, is Greek and is the name of a musical instrument made from the wood of this genus. Mexicana is New Latin for of Mexico, probably referring to the preferred southern range of this species. 104 © Stanford University Press Plant Descriptions Orange Honeysuckle (Lonicera ciliosa) Wedded Honeysuckle (Lonicera conjugialis) Blue Elderberry (Sambucus mexicana) 105 Plant Descriptions Red Elderberry Sambucus racemosa L. var. microbotrys (Rydb.) Kearney & Peebles Native. Honeysuckle Family (Caprifoliaceae). White radial flowers with five petals are found in a dome-shaped raceme at the branch tips of this tall shrub. The fruits ripen into small round shiny red berries that many birds find attractive. The large pinnately compound leaves are opposite, and the 5-7 leaflets, which are also opposite, are finely toothed. Red Elderberry grows in wet meadows and moist places along the Pacific Crest Trail at 6400 feet and lower. It flowers in June and the fruits ripen in August. It will be surrounded by Big-leaf Lupine (Lupinus polyphyllus) and Swamp White Heads (Sphensciatum capitellatum). Be careful not to confuse this with Blue Elderberry. The two shrubs are similar in many ways, but the two major distinctions are the inflorescence shape and the color of the fruit. The inflorescence of Red Elderberry is dome-shaped, while Blue Elderberry is flat-topped. The fruits of Red Elderberry are bright red, while the fruits of Blue Elderberry are bluish. Elderberry fruits make permanent inks, and a not-so-permanent violet dye. The ripe fruits are edible when cooked, but cause nausea when raw. The bark, stems, leaves, and roots are poisonous because they contain alkaloids and cyanide-producing glycosides. The genus name, Sambucus, is Greek and is the name of a musical instrument made from the wood of this genus. Racemosa is Latin for a cluster, referring to the inflorescence. Snowberry Symphoricarpos albus (L.) S.F. Blake var. laevigatus (Fern.) S.F. Blake Also called Common Snowberry, Upright Snowberry, and Waxberry. Native. Honeysuckle Family (Caprifoliaceae). This shrub is best appreciated in the fall and winter after the leaves fall off and clusters of white berries are exposed. White is an unusual color in the woods, so these fruits become quite obvious. In the spring, the opposite leaves appear before the tiny flowers. The leaf margins vary from entire to lobed. The tiny nodding, bell-shaped flowers have five lobes, and tend to grow in clusters of 416 flowers. These little pink blossoms are usually mature in July, and the waxy, white berries ripen around September, shortly before the deciduous leaves fall. On Mount Ashland, this erect shrub commonly grows 90-120 cm tall. Look for Snowberry in the forest understory below 6700 feet. In Oregon, it was reported that a child was poisoned after eating the berries of Snowberry. The child experienced nausea, delirium, and was in a semi-comatose state for awhile. Derived from several Greek words, symphoricarpos means to bear fruit together, referring to the clusters of berries. Albus is Latin for white, obviously referring to the color of the berries. Creeping Snowberry Symphoricarpos mollis Nutt. Also called Trip Vine. Native. Honeysuckle Family (Caprifoliaceae). A trailing shrub, Creeping Snowberry has reddish new stems that bear opposite leaves whose margins vary from entire to lobed. There are many soft hairs on the margins of the leaves. The small, nodding, urn-shaped flowers have five lobes. Two to eight pink-reddish urns bloom together in the leaf axils. If you use your hand lens to pear inside the flower, you’ll find that it is hairy inside. The flowers usually mature in July or August, while the fruits ripen in August and September. The deciduous leaves fall in September or October, revealing the white fruits. The creeping stems are usually about 60-90 cm long. Look for Creeping Snowberry in the forest understory on the south slopes along the Pacific Crest Trail. Nearby, you will probably see another shrub, Bush Chinquapin (Chrysolepis sempervirens) or White Hawkweed (Hieracium albiflorum). This plant looks like Common Snowberry whose habit is erect rather than trailing. Native Americans used the berries and roots for medicinal purposes. The leaves contain saponin, which is poisonous. There is some disagreement over whether the berries are poisonous; it seems to depend on the individual’s sensitivity. Symphoricarpos is Greek and means to bear fruit together, referring to the clusters of berries. Mollis is Latin for soft, perhaps referring to the soft hairs inside the flowers and on the leaf margins. 106 Plant Descriptions Photos by Charles Webber, California Academy of Sciences Red Elderberry (Sambucus racemosa) Snowberry (Symphoricarpos albus) Creeping Snowberry (Symphoricarpos mollis) 107 Plant Descriptions Pink Family (Caryophyllaceae) Ballhead Sandwort Arenaria congesta Nutt. var. congesta Also called Dense-flowered Sandwort. Native. Pink Family (Caryophyllaceae). At first glance, you may confuse this flower with Ball-Head Gilia, but if you look closely, you’ll easily see the differences. The radial flowers of Ballhead Sandwort have five petals, five sepals, 10 stamens and 3 styles. The stamens and styles are all greatly exserted. The flowers are arranged in a terminal, head-like cyme. The opposite leaves are simple and entire, resembling short blades of grass. The stem is glabrous and the nodes are slightly swollen. This erect perennial can grow 3-16 inches tall. Look for Ballhead Sandwort around Rabbit Ears and in other dry, open areas from late June to early July. The common name, Sandwort, refers to the fact that these plants are often found in loose, dry, sandy soils. “Wort” is old English for plant. Arenaria is Latin for sand, in reference to the soil types preferred by the members of this genus. Congesta is Latin for carried together, probably referring to the tight cluster of flowers. Large-leaf Sandwort Moehringia macrophylla (Hook.) Fenzl Also called Big-leaf Sandwort. Native. Pink Family (Caryophyllaceae). This low-growing perennial has one to two flowers at the top of each flowering stem. The small, radial flowers are white with five entire petals. Usually, flowers in this family have petals that are notched at the tip, but this flower is an exception to that rule. The petals open fully to reveal 10 stamens and 3 styles. The flowering stems are very leafy with narrow, opposite leaves that have entire margins. Flowering stems arise from an above-ground runner, which is why this plant grows in distinct patches. Large-leaf Sandwort is a short little flower, growing no more than 10 cm high in the moist, shady undergrowth of the forest. It is often found at the bases of trees. I found my first patch of this little flower on the Knoll at mile 8 along County Road 1151. It was on the north side of the peak of the knoll, growing with Davis’s Knotweed (Polygonum davisiae) and Showy Jacob’s Ladder (Polemonium californicum). It also grows along the Pacific Crest Trail in shaded areas. P.H.G. Moehring (1710-1791) was a naturalist from Danzig, Germany whose name is credited in this genus. Macrophylla is Greek for large leaf or long leaf, obviously referring to the leaves, which are larger in this species than in other species of this genus. Palmer’s Catchfly Silene bernardina S. Watson Native. Pink Family (Caryophyllaceae). Your lasting impression of this plant should be “sticky,” so get down there and touch it! The stem, leaves, and calyxes are covered with glandular hairs that hold sticky oils. The erect stem is usually branched, and bears opposite leaves and axillary flowers. The leaf margins are entire, while the five white to pinkish flower petals are so deeply lobed that one petals appears to be four very narrow petals. If you look closely (with a hand lens) at the center of the flower, you will see the throat is packed with 10 stamens and 3 or 4 styles. Palmer’s Catchfly usually grows 15-40 cm. This plant prefers open, dry meadows on the south slopes of Mount Ashland. Look for it in earlyto mid-July on the ridgeline behind Rabbit Ears and in the draw between Rabbit Ears and Grouse Gap Shelter. Frosted Indian Paintbrush (Castilleja pruinosa) and Worm-leaf Stonecrop (Sedum stenopetalum) may be growing nearby. One of the previous scientific names for this plant was Silene pamleri, after a gentleman whose last name was Palmer. This is where the common name comes from. One source suggests that the genus name, Silene, is from Silenus, the drunken foster-father of Bacchus in Greek mythology. The story goes that Silenus was covered with foam from the sticky secretions of many plants. Bernardina honors someone whose name is Bernard. 108 Plant Descriptions Ballhead Sandwort (Arenaria congesta var. congesta) © Stanford University Press Large-leaf Sandwort (Moehringia macrophylla) Palmer’s Catchfly (Silene bernardina) 109 Plant Descriptions Red Sand Spurry Spergularia rubra (L.) J.S. Presl & C. Presl Also called Ruby Sand Spurry, and simply Sand Spurry. Non-native from Europe. Pink Family (Caryophyllaceae). Ah! You’ve found another belly flower! Lay down on your belly, take out your hand lens, and prepare yourself for a little delight. The tiny flowers have five lavender/pink petals that are surrounded by five green sepals. The petals and the sepals are the same size and shape, but the outer surface of the sepals is glandular. They open wide to reveal 6-10 stamens and 3 styles. The terminal flowers top the stem that has many tiny, linear leaves in dense whorls. At the base of the entire leaves, there are shiny white stipules. The sprawling stems of Red Sand Spurry may measure 2-3 cm, but the flowers are no more than 1 cm from the ground. These delicate flowers grow only in open, dry, gravelly areas, blooming in early- to mid-July. Look for it in the ski area on the wide path just inside the wooden fence. A little closer to the fence, you will find two mustard plants: Virginia Pepperweed (Lepidium virginicum var. virginicum) and Mountain Tansy Mustard (Descurainia incisa var. incisa). Derived from Latin, spergularia means to scatter, referring to the practice in Europe of sowing the seeds of Spergula, another genus in this family, for early forage. Rubra is Latin for red and refers to the seeds which are reddish to dark brown at maturity. Curled Starwort Stellaria crispa Cham. & Schldl. Also called Crisped Starwort, and Crisp Sandwort. Native. Pink Family (Caryophyllaceae). This prostrate to trailing perennial has small, radial flowers, but there are no petals on these flowers! The five sepals are light green with whitish, papery margins. Inside the sepals, 10 stamens and three styles are protected. The flowers grow in the axils of opposite leaves. The evenly spaced leaves have margins that are entire but wavy, shiny and glabrous. There are occasional hairs on the stem between the leaves. The stems are often 10-30 cm long, but because they are prostrate, the flowers are never more than 6-8 cm from the ground. Curled Starwort grows in damp, shady areas. Stellaria is from the Latin for star, and refers to the shape of the flowers. Crispa is from the Latin for curled, and probably refers to the wavy leaf margins. The common name is based on the scientific name: curled refers to the leaf margins, star refers to the flower shape, and wort is old English for plant. Common Chickweed Stellaria media (L.) Villars Non-native from southwest Europe. Pink Family (Caryophyllaceae). Small, radial flowers have five white petals that are so deeply notched there appear to be 10 petals. The bushy center of the flower displays 10 stamens and 3 styles. The flowers are both terminal and axillary. The opposite leaves are flat and shiny and have entire margins. The weak, slender stems usually grow 7-40 cm, and they tend to use surrounding vegetation for support. You will find this little gem in the wet meadows on the south slopes of Mount Ashland. It blooms late-June through mid-July, and is often tangled with American Brooklime (Veronica americana) and Goosegrass (Galium aparine). I know from experience that Common Chickweed, which often is a weed in people’s gardens, is edible and makes a delicious addition to an early spring salad. Just pick, wash, and eat. Stellaria is from the Latin for star, and refers to the shape of the flowers. Media is Latin for in the middle or between; it is uncertain to what this refers. 110 Plant Descriptions Red Sand Spurry (Spergularia rubra) Photo by Jeff Abbas Curled Starwort (Stellaria crispa) Common Chickweed (Stellaria media) 111 Plant Descriptions Staff-Tree Family (Celastraceae) Oregon Boxwood Paxistima myrsinites (Pursh) Raf. Also called Oregon Boxleaf. Native. Staff-Tree Family (Celastraceae). This prostrate to ascending shrub is not abundant on Mount Ashland, so if you’ve found it, consider yourself lucky. The woody stems are very stiff and highly branched. The small, shiny, dark green, leathery leaves are opposite and finely toothed. They are usually about 1 cm long. In the leaf axils, tiny flowers with four petals are a delightful deep rosy color. Each petal is only 1-2 mm long. Four stamens and one short, stout pistil are at the flower center. The entire shrub grows to 30-60 cm. You will find Oregon Boxwood in the understory of the coniferous forest along the Pacific Crest Trail at lower elevations. It blooms from mid- to late June. Nearby, you may see Greenleaf Manzanita (Arctostaphylos patula) or Silverleaf Phacelia (Phacelia hastata). Paxistima is derived from Greek and means thick stigma, referring to the stigma on these flowers. Myrsinites comes from the Greek word for myrtle. Stonecrop Family (Crassulaceae) Cascade Stonecrop Sedum divergens S. Watson Also called Pacific Stonecrop, and Spreading Stonecrop. Native. Stonecrop Family (Crassulaceae). A matted and glabrous perennial, Cascade Stonecrop bears 3-17 yellow flowers. Each flower has 5 sepals, 5 petals, 10 stamens with yellow anthers, and 5 pistils. The extremely succulent leaves are opposite, sessile, nearly spheric, and have rounded tips. This sun-loving plant grows 5-12 cm tall in open, dry areas with gravelly soil. The succulent leaves of Cascade Stonecrop were eaten by several Native American groups in the Pacific Northwest. Other native groups used this plant as a medicine. Sedum is from the Latin word meaning to assuage. This refers to the healing properties of a plant called Houseleek (Sempervivum tectorum), to which Sedum was applied by some authors. Houseleek is found in the northeastern U.S., and is also in the family, Crassulaceae. Divergens is Lating for diverging; it is uncertain to what this refers. Spear-leaf Stonecrop Sedum lanceolatum Torrey Also called Lance-leaf Sedum. Native. Stonecrop Family (Crassulaceae). A striking plant, Spear-leaf Stonecrop sets deep yellow flowers atop a stem full of, and surrounded by, deep red leaves. Each yellow flower has 5 green sepals, 5 yellow petals that usually have a red mid-veins, 10 stamens with yellow anthers and 5 pistils. Each plant yields 3-24 flowers. The basal and stem leaves are very succulent, but the alternate, sessile stem leaves tend to fall when the plant is in flower. Often, you’ll find clumps of the red basal leaves before you see the flowers. The erect flowering stems usually reach 3-15 cm. This perennial grows in dry, gravelly soil from Rabbit Ears upward to the summit on the south slopes. It blooms from mid-June to early July with Rosy Pussytoes (Antennaria rosea) and Ball-head Ipomopsis (Ipomopsis congesta var. congesta). Sedum is from the Latin word meaning to assuage. This refers to the healing properties of a plant called Houseleek (Sempervivum tectorum), to which Sedum was applied by some authors. Houseleek is found in the northeastern U.S., and is also in the family, Crassulaceae. Lanceolatum is derived from the Latin word for lance, referring to the shape of the flower petals. 112 Plant Descriptions Oregon Boxwood (Paxistima myrsinites) Photo by Charles Webber, California Academy of Sciences Cascade Stonecrop (Sedum divergens) Spear-leaf Stonecrop (Sedum lanceolatum) 113 Plant Descriptions Cream Stonecrop Sedum oregonense (S. Watson) M. Peck Also called Creamy Stonecrop. Native. Stonecrop Family (Crassulaceae). The large substantial leaves make this Stonecrop fairly easy to identify. The thick, succulent basal and stem leaves are rounded or barely notched at the tip. The stem leaves are alternate and sessile, and the new leaves are glaucous. The leaves are primarily green, but some may be pink/red mottled. Each flowering stem bears 12-20 flowers, and each flower has 5 sepals, 5 pale yellow petals, 10 stamens with yellow to red-brown anthers, and 5 pistils. This perennial usually grows 4-20 cm tall. You will have the most luck finding this flower in and around the rocky outcrops on the south slopes from Rabbit Ears to the summit. It blooms from mid-June to early-July, next to Scabland Fleabane (Erigeron bloomeri) and Davidson’s Penstemon (Penstemon davidsonii). The leaves of all known Stonecrop species are edible. However, some species have emetic and cathartic properties and can cause headaches. Before eating any plant, please consult a botanist to confirm its correct identification. It is also important to research the plant thoroughly to determine which part of the plant is edible and whether it requires specific preparation. Sedum is from the Latin word meaning to assuage. This refers to the healing properties of a plant called Houseleek (Sempervivum tectorum), to which Sedum was applied by some authors. Houseleek is found in the northeastern U.S., and is also in the family, Crassulaceae. Oregonense is New Latin, meaning of Oregon, referring to the general range of this species. Worm-leaf Stonecrop Sedum stenopetalum Pursh Also called Narrow-Petaled Sedum. Native. Stonecrop Family (Crassulaceae). One small, pale yellow flower tops a non-branching flowering stem. The flower has 5 sepals, 5 narrow petals, 10 stamens with yellow or light brown anthers, and 5 pistils. The sessile, alternate leaves are narrow and long tapered, with papery bases. The leaves not very succulent, and the well-spaced, upper leaves appear withered. There are many leaves tightly packed on the lower portion of the stem. This glabrous perennial grows 5-16 cm tall in well-drained soils. Look for Worm-leaf Stonecrop on the south slopes in the dry soil behind Rabbit Ears and above USFS Road 20. It blooms in mid-July with Staining Collomia (Collomia tinctoria) and Palmer’s Catchfly (Silene Bernardina). Sedum is from the Latin word meaning to assuage. This refers to the healing properties of a plant called Houseleek (Sempervivum tectorum), to which Sedum was applied by some authors. Houseleek is found in the northeastern U.S., and is also in the family, Crassulaceae. Stenopetalum is derived from Greek and means narrow leaf, referring to the narrow petals and/or the narrow leaves. Heath Family (Ericaceae) Pinemat Manzanita Arctostaphylos nevadensis A. Gray Native. Heath Family (Ericaceae). The white to pale pink urn-shaped flowers of Pinemat Manzanita have five lobes, indicating that these flowers are made of five fused petals. The flowers and the inflorescence are nodding. This lowlying shrub has shiny, alternate, entire leaves, and the twigs often have fine, woolly hairs. Plants on the south slopes bloom fairly early – around mid-May – while plants on the north slopes and in the shade will bloom as late as early June. The bright red fruits usually ripen around late August. This plant is abundant on the ski slopes and along the ridge that runs northwest from Rabbit Ears. Some sources claim that Pinemat Manzanita was used by Native Americans for herbal teas and smoking mixtures similar to tobacco. Arctostaphylos is Greek for bear berries, perhaps referring to the fact that bears eat the berries of some of the species in this genus. Nevadensis is a Latin derivation of the state name Nevada. It is likely that the first of this species was found in or near Nevada. 114 Plant Descriptions Cream Stonecrop (Sedum oregonense) Worm-leaf Stonecrop (Sedum stenopetalum) Pinemat Manzanita (Arctostaphylos nevadensis) 115 Plant Descriptions Greenleaf Manzanita Arctostaphylos patula E. Greene Native. Heath Family (Ericaceae). This erect shrub has bright green, shiny, leathery leaves with entire margins. The flowers can vary from bright pink to pale pink to white, and the five petals are fused together to form an urn-shape. The flowers are clustered together toward the ends of the limbs. The bark is usually reddish-brown. The flowers appear in May while the dark brown fruits ripen around September. This shrub can be found around the stones of Rabbit Ears, and next to USFS Road 20 between the picnic area and Rabbit Ears. Although these berries are edible, they are not very tasty. Arctostaphylos is Greek for bear berries, perhaps referring to the fact that bears eat the berries of some of the species in this genus. Patula is Latin for open, spreading. This is probably in reference to the inflorescence which is open and spreading compared to the inflorescences of other species in this genus. Little Prince’s Pine Chimaphila menziesii (D. Don) Sprengel Native. Heath Family (Ericaceae). Each slender reddish-green stem produces one flower with white, reflexed petals. The tips of the petals may be tinged with pink. Because the petals are reflexed, it is easy to see the 10 stamens and one large pistil in the center. The opposite or whorled leaves are coarsely toothed and have white borders along the main veins. The flowers often grow in small clumps, and can give the impression of being a very low-growing shrub, usually shorter than 15 cm. You will find Little Prince’s Pine growing in the quiet of the coniferous forest. Look for it in late July along the Pacific Crest Trail along with Pipsissewa (C. umbellata) or White-veined Wintergreen (Pyrola picta). Little Prince’s Pine is very similar to Pipsissewa (C. umbellata), but Pipsissewa is larger, and has a completely green stem, and the leaves have no stems (petioles). Usually the flower of Pipsissewa is pink, and one plant produces several flowers. In the past, Native Americans have used Little Prince’s Pine to make a tea that relieved kidney problems. They believed this plant was good for all the vital organs. Chimaphila is Greek for winter-loving, referring to the fact that many plants in this genus have evergreen leaves. Archibald Menzies (1754-1842) was one of the first botanists from Scotland to reach the Pacific Coast of North America in 1792. He was the first to collect plants along the shore of what is now Washington, and the second to collect along the shore of California. Pipsissewa Chimaphila umbellata (L.) Bartram Also called Prince’s Pine, Wintergreen, Wax Flower, Ground Holly, Western Prince’s Pine, and Common Prince’s Pine. Native. Heath Family (Ericaceae). Pink, nodding flowers with spreading petals make Pipsissewa unique. The radial flowers have 5 petals that are mostly pink, but have white along the margins. Their spreading nature allows us easily to see the 10 stamens and one large green pistil in the center. The inflorescence, which is a raceme, rises above several sets of whorled evergreen leaves that are coarsely toothed. The stem is completely green, but the individual flower branches are reddish. Pipsissewa grows in the cool, moist, shaded woods. It blooms in late July, often next to its close relative, Little Prince’s Pine (C. menziesii), and perhaps with Baneberry (Acteae rubra). Pipsissewa once had a considerable medicinal reputation among whites and Native Americans. The leaves contain ericolin and tannin, and were used in infusions for bladder stones. The plant is still used in parts of Europe as a home remedy for wounds. The Cree Indians used the leaves as a tea substitute for healing internal organs. Once upon a time, it was also used to make Root Beer. A small percentage of people have reported cases of dermatitis after handling this plant. This won’t be a concern if you remember not to pick the flowers on Mount Ashland. Chimaphila is Greek for winter-loving, referring to the fact that many plants in this genus have evergreen leaves. Umbellata is derived from the Latin word for umbrella, in reference to the spreading petals that look like an umbrella. 116 Plant Descriptions Greenleaf Manzanita (Arctostaphylos patula) Little Prince’s Pine (Chimaphila menziesii) Pipsissewa (Chimaphila umbellatum) 117 Plant Descriptions Bog Laurel Kalmia polifolia Wangenh. Also called Pale Laurel, Western Bog Laurel, and Swamp Laurel. Native. Heath Family (Ericaceae). Bog Laurel has unmistakable pink to rose-purple flowers. Five sepals are fused toward the base and rose-colored near the top with white margins. The five petals are fused half way, forming an open cup-shape. Ten stamens have white, recurved filaments and the dark purple anthers are tucked into pockets in the corolla until the pollen is released, at which point the stamens stand erect. The stamens, thus, act as tripwires for insects. When an insect lands on the flower and trips the filaments, the anthers pop up to dust the insect with pollen. The style is white at the base, red at the top, and the stigma is white. Four to five flowers grow on one stem in the upper leaf axils. The leathery leaves can be opposite, alternate or whorled and are entire, with parts of the margins rolled under. The lower surfaces of the leaves have many white hairs. This shrub flowers for a very short period in May. I have seen this plant only in the wet meadow on the south slope below Rabbit Ears and above USFS Road 20. Bog Laurel contains a poisonous glycoside called andromedotoxin which, when ingested, lowers blood pressure and can cause breathing problems, dizziness, cramps, vomiting, diarrhea, and progressive paralysis. P. Kalm (1716-1779) is commemorated in this genus name. Kalm was a student of Linnaeus who traveled extensively in eastern North America. In Latin, polifolia means gray leaf, referring to the lower surfaces of the leaves that are covered with white hairs. Pinesap Monotropa hypopitys L. Native. Heath Family (Ericaceae). A fleshy saprophyte, Pinesap, is not a green plant because there is no chlorophyll in the cells. This perennial obtains its nutrients from the decaying organic matter in the forest soil. The unbranching flowering stems produce many bell-shaped flowers in a nodding, raceme-like inflorescence. The entire plant – the stem, leave, sepals, petals, etc. – are all the same color: yellowish to pink or red. After the plant goes to seed, it dries black. Each flower has 5 sepals, 5 petals, 10 stamens and 1 pistil. Each petal is slightly cup-shaped, bulging at the base. The narrow sepals and wider petals have very short hairs. The scale-like leaves are alternate and entire. Pinesap is usually 5-25 cm tall. On Mount Ashland, it will be found in moist, forested areas at elevations below 6000 feet. You may also find Woodland Pine Drops (Pterospora andromedea) or Rattlesnake Plantain (Goodyera oblongifolia) growing in the same type of habitat. Monotropa is Greek meaning one direction, referring to the one-sided inflorescence. Hypopitys also comes from Greek and means beneath the tree, referring to the forest habitat of this species. One-sided Wintergreen Orthilia secunda (L.) House Also called Sidebells Wintergreen, and One-sided Pyrola. Native. Heath Family (Ericaceae). Nodding flowers in a one-sided arcing raceme give this flower the first half of its name. Five small, green sepals are found at the base of the five greenish to cream-white petals that never fully open. The petals closed, protecting the 10 stamens and 1 pistil. Some anthers and the stigma usually poke out of the enclosed petals. Although the inflorescence arcs in flower, it straightens in fruit. The evergreen leaves give this perennial the second half of its name. Technically, the leaves are attached to the stem but they are so close to the base that they appear to be basal leaves. They are egg-shaped or widely oval, entire, and shiny green. The erect stem grows to 12-20 cm. One-sided Wintergreen generally grows in dry, shady forests. On Mount Ashland, I found it on the north slopes, west of the ski slopes on a rocky outcrop. Also on the rocks was a lot of Woodland Penstemon (Nothochelone nemorosus). You will find both blooming in late July to early August. In Greek, orthilia means straight spiral, referring to the one-sided raceme. In Latin, secunda means the second, in a row, or on one side, referring to the habit of the flowers to grow on one side of the stem only. 118 Photo by Br. Alfred Brousseau, St. Mary’s College Plant Descriptions Photo by Bob Vos Bog Laurel (Kalmia polifolia) © Stanford University Press Photo by Bob Vos Pinesap (Monotropa hypopitys) One-sided Wintergreen (Orthilia secunda) 119 Plant Descriptions Woodland Pine Drops Pterospora andromedea Nutt. Also called simply Pine Drops. Native. Heath Family (Ericaceae). This robust perennial is another saprophyte, and therefore, has no green structures. A saprophyte is a plant that obtains its nutrients from decaying organic material in the soil. There is no need for the chlorophyll that makes most plants green. The erect stem is reddish and has alternate, reddish, scale-like leaves. The long raceme bears many nodding flowers that each have 5 reddish sepals, 5 cream colored or yellowish petals that are almost completely fused and have reflexed lobes. Ten stamens are hidden inside the urn-shaped flowers. The stem and calyx are very sticky. Woodland Pine Drops can grow up to 120 cm in the moist shade of coniferous forests. Look for it at elevations below 6000 feet. In the same habitat, you may find California Harebell (Campanula prenanthoides) or Pinesap (Monotropa hypopitys). Greek for winged seed, pterospora describes the seeds, which have a wing-like margin on one side. Andromedea is Greek, and refers to the flowers, which closely resemble the flowers of another genus in this family, Andromeda. Andromeda of Greek mythology, was the daughter of Cassiopeia. The tales say that Andromeda was chained to a rock in the middle of the ocean. Similarly, plants in the genus Andromeda grow in the middle of swampy areas. White-veined Wintergreen Pyrola picta Smith Also called Whitevein Pyrola, and Whiteleaf Pyrola. Native. Heath Family (Ericaceae). Nodding white flowers on a naked stem are bilateral but may appear radial. Five short sepals are reddish-brown to greenish; the 5 petals are greenish, cream white or pink. The style is greatly exserted and curved, looking like an elephant’s trunk, while the 10 stamens are hidden inside the petals. The terminal raceme tops the reddish-brown, glabrous stem that usually reaches 10-30 cm tall. The leathery basal leaves are just as striking as the beautiful flowers. These egg-shaped leaves are deep green with prominent white markings around the veins, while the lower surface is often purple. They are entire to irregularly toothed. White-veined Wintergreen grows in moist wooded areas on the ski slopes and along the Pacific Crest Trail. Look for it blooming in late-July to early August. In nearby shaded areas, you may find White Hawkweed (Hieracium albiflorum) or Little Prince’s Pine (Chimaphila menziesii). At first glance, this plant resembles Rattlesnake Plantain (Goodyera oblongifolia) because of the similarity in size, habitat, and mottling of leaves. But look closely at the leaves and flowers. The leaves of Whiteleaf Pyrola are more rounded and have a thin petiole, and the flowers are more or less radial with 5 petals. The leaves of Rattlesnake Plantain are elongated and sessile, and the flower is orchid-shaped. Pyrola is from the Latin word, pyrus, which means pear. Sometimes the leaves are pear shaped. Picta is also from Latin, and means painted, referring to the white veins on the leaves. Thinleaf Huckleberry Vaccinium membranaceum Hook. Also called Black Huckleberry. Native. Heath Family (Ericaceae). This erect, glabrous shrub usually grows 1-2 meters. The deciduous leaves are very thin and membranous, rounded to elliptic, 2-5 cm long, and finely toothed. Each tooth is topped with a glandtipped hair. The lower surface of the alternate leaves has prominent veins. The little, pink, urn-shaped flowers grow in the axils of the lowest leaves of the youngest shoots. The flowers usually have no sepals, and five petals that are fused 2/3 of their length. Ten stamens and 1 pistil are hidden inside the urn. The fruits are black or dark red berries. Thinleaf Huckleberry is found at elevations below 6200 feet in the forest understory on the north slopes. You may find it surrounded by Engelmann Spruce (Picea engelmannii). The berries are reportedly the most delicious of Vaccinium species. Vaccinium is derived from the Latin word for cow; it is uncertain to what this refers. Membranaceum is Latin for membrane, referring to the very thin leaves. 120 Photo by Bob Vos Plant Descriptions Woodland Pine Drops (Pterospora andromedea) White-veined Wintergreen (Pyrola picta) Thinleaf Huckleberry (Vaccinium membranaceum) 121 Plant Descriptions Western Blueberry Vaccinium uliginosum L. ssp. occidentalis (A. Gray) Hulten Also called Bog Blueberry, Western Huckleberry, and Bog Bilberry. Native. Heath Family (Ericaceae). This erect, glabrous shrub is much shorter than its close relative, Thinleaf Huckleberry (V. membranaceum). This shrub only reaches 60-90 cm tall. The alternate, deciduous leaves are rather thick, elliptic to rounded, and entire. The flowers are usually borne in racemes, with 1-4 flowers. Each little urn-shaped flower has 4-5 sepals, 5 white petals that are fused 2/3 of their length, 10 stamens, and 1 pistil. The berries are blue and glaucous. Western Blueberry grows in the wet meadows on the south slopes. Look for it in wet areas along the Pacific Crest Trail. It blooms in June, and the fruits ripen in August. Around it, you may see Fringed Willow-herb (Epilobium ciliatum ssp. glandulosum), or Musk Monkey Flower (Mimulus moschatus). The blue berries can be eaten fresh, dried, or frozen. Vaccinium is derived from the Latin word for cow; it is uncertain to what this refers. In Latin, uliginosum means moisture, and refers to the moist habitat of this species. Occidentalis is Latin for western, referring to its range. Vaccinium species tend to be called blueberries if the fruit is blue, and huckleberry if the fruit is any other color. Legume Family (Fabaceae) Leafy Pea Lathyrus polyphyllus Nutt. Also called Oregon Pea, and Leafy Peavine. Legume Family (Fabaceae). Six to twelve bilateral flowers are displayed on one-sided axillary racemes. The flowers vary in color from pink to purple, to red-purple, to blue. The alternate leaves are pinnately compound with obvious alternate leaflets. Ten to sixteen leaflets are egg-shaped and entire. At the tip of the central leaf axis is a tendril that is either branched, coiled, or reduced to a bristle. At the base of the leaf stem, very large oval stipules can be mistaken for opposite leaves. This robust perennial has a glabrous stem that is angled or flanged, but not winged. You will find Leafy Pea in the forest and at the forest edge below 6200 feet. There are 21 species of Lathyrus in Oregon, all of which are toxic to humans and livestock. The seeds contain the toxin that in large quantities can cause serious convulsions and paralysis (lathyrism). Lathyrus is an ancient Greek name for a kind of vetch. Polyphyllus is derived from Greek and means many leaves, referring to the many leaflets that compose one leaf. Bird’s Foot Trefoil Lotus purshianus (Benth.) Clements & E.G.Clements var. purshianus Also called Pursh’s Lotus. Native. Legume Family (Fabaceae). A dainty, low-growing plant, this annual produces small, pale pink, bilateral flowers in the leaf axils. The light green sepals are long and narrow with fine, white hairs. The sepals remain when the flower matures to fruit. As the fruit matures, the base of the calyx turns deep pink. Because this plant is a legume, the fruits look like miniature green beans, only the color is different – these fruits are a rich brown. Alternate leaves are compound with three entire leaflets. The stems and leaves are covered with short, fine hairs. The stems are prostrate to erect and can be simple or branched. The overall height of the plant is usually no more than 15 cm. I have seen this plant only in the gravel along County Road 1151 between the marker for mile 8 and the ski area parking lot. Growing nearby, you may see Tall Annual Willow-Herb (Epilobium brachycarpum) or Shasta Tansy Aster (Machaeranthera canescens var. shastensis). The Greek derivation and meaning of lotus is unclear, but a similar Latin word means washing. Purshianus is derived from Frederick Pursh’s name. Pursh (1774-1829), a German botanist, was the first to publish information regarding the specimens collected by Lewis and Clark during their famous expedition. Pursh published Flora Americae Septentrionalis in 1814. 122 Plant Descriptions Photo by Br. Alfred Brousseau, St. Mary’s College Western Blueberry (Vaccinium uliginosum) Leafy Pea (Lathyrus polyphyllus) Bird’s Foot Trefoil (Lotus purshianus var. purshianus) 123 Plant Descriptions Sickle-keel Lupine Lupinus albicaulis Hook. Also called Pine Lupine. Native. Legume Family (Fabaceae). Terminal racemes bear whorls of bilateral flowers. The pea-like flowers vary in color from purple to white. Ten stamens, five long and five short, and 1 pistil are hidden inside keel. The keel tip shows between the wings. The rusty banner is reflexed. The alternate stem leaves are palmately compound with 5-10 entire leaflets. Narrow stipules, 5-18 mm long, are at the base of each leaf stem. The erect stem varies in color from very pale green (almost white) to reddish-brown. The stem, leaves, stipules and calyx have fine hairs that may only be seen with a hand lens. There seems to be two genetic variations of this plant on Mount Ashland. The plants with purple flowers have the reddish-brown stems, while the plants with white flowers have the whitish stems. I only observed three plants with white flowers, and they were all along USFS Road 20, very close to the confluence of Road 40S30 that leads to Grouse Gap Shelter. Because these white Lupines are so rare on Mount Ashland, PLEASE DO NOT PICK THESE PLANTS. Sickle-keel Lupine is found in dry, open areas. Look for it around Rabbit Ears, along the ridge behind Rabbit Ears, and along USFS Road 20. Scarlet Gilia (Ipomopsis aggregata) and Siskiyou Mountains Owl’s Clover (Orthocarpus cuspidatus var. cuspidatus) may be growing nearby. All plants require nitrogen, but no plant can absorb nitrogen from the atmosphere. Most members of the Legume Family have a very special relationship with a mycorrhizal fungus. This fungus grows on the fine root hairs and does the important job of converting atmospheric nitrogen into a form that plants can absorb. Thus, this fungus makes nitrogen available not only to the plant on which it grows, but for the surrounding plants, as well. In subalpine habitats with dry, porous soils, this extra bit of nitrogen is extremely important. The seeds, fruits, and young shoots of this plant have toxic alkaloids. These toxins are especially dangerous to livestock. Lupinus is Latin for wolf, from the mistaken idea that the plants in this genus robbed the soil of nutrients. Albicaulis is derived from Latin, and means white stem, referring to the stem color on some of these plants. Mt. Ashland Lupine Lupinus aridus ssp. ashlandensis Native. Legume Family (Fabaceae). If you’ve discovered this flower, you’ve stumbled upon a real jewel: this flower is found only on Mt. Ashland and in small populations on Red Mountain. These small, irregular, lavender to bluish flowers grow in a spike-like raceme. The erect stem has long white hairs, and no leaves. The stems can grow from 6-12 inches tall. Each stem is surrounded by many palmately compound basal leaves. Each leaf contains 5 to 7 leaflets that are keeled, so that each leaflet appears to fold upward. The leaves and leaf petioles are covered with long, fine, white hairs often appearing gray or silvery. Run your fingers over these leaves for a wonderful tactile treat! Mt. Ashland Lupine grows profusely around Stonehenge in early July, but you may see only the basal leaves much earlier than this. From 1980 to 1996, Mt. Ashland Lupine was a Category 2 candidate for listing under the U.S. Endangered Species Act. This category included species for which sufficient data was not available to confirm vulnerability. In 1996, this category of species was discontinued, and Mt. Ashland Lupine was dropped. In September 1999, the U.S. Fish and Wildlife Service received a formal petition from the Rogue Group of the Sierra Club of Ashland to list Mt. Ashland Lupine as endangered or threatened. In June 2000, the U.S. Fish and Wildlife Service announced that it would conduct a comprehensive review of Mt. Ashland Lupine to determine if it should be protected under the ESA. Since 1995, the U.S. Forest Service has considered this plant a sensitive species. Lupines generally can withstand dry, poor soil. The name Lupinus comes from the Latin word for wolf, lupus, because the plant was thought to “devour” the soil. Ironically, lupines, like many others in the pea family, actually enrich the soil by fixing nitrogen. The name aridus comes from the Latin word for dry, and refers to the general habitat for this species. Ashlandensis is New Latin and means of Ashland, referring to the location of this subspecies. 124 Plant Descriptions Sickle-keel Lupine (Lupinus albicaulis) Mt. Ashland Lupine (Lupinus aridus var. ashlandensis) 125 Plant Descriptions Brewer’s Lupine Lupinus breweri A. Gray var. breweri A. Gray Native. Legume Family (Fabaceae). This low, mat-forming plant has irregular lavender-bluish flowers with palmately lobed leaves that appear silvery as a result of fine, soft, white hairs. The five to seven leaflets usually appear to be folded longitudinally down the center. There are alternate leaves on the stem and masses of basal leaves. Most of the above-ground stem is woody. These perennials or subshrubs are generally about 5-8 cm high. You’ll find Brewer’s Lupine in open areas around Rabbit Ears, up toward the summit, around Grouse Gap Shelter, on the ski slopes, and on the Knoll, blooming throughout June. Look for Cobwebby Paintbrush (Castilleja arachnoidea) and Spreading Phlox (Phlox diffusa) growing nearby. Lupinus is Latin for wolf, from the mistaken idea that the plants in this genus robbed the soil of nutrients. Breweri honors William Henry Brewer (1828-1910), a geologist and botanist. Brewer was the botanist for the California State Geological Survey from 1860-1864. Big-leaf Lupine Lupinus polyphyllus Lindley var. pallidipes (A.A. Heller) C.P. Smith Also called Large-leaf Lupine, and Blue-Pod Lupine. Native. Legume Family (Fabaceae). This large, showy plant will be growing in wet meadows in late June, early July. The stout stems can grow to heights of 120 cm. The lavender, irregular flowers grow in whorls on a long spike. The typical palmately lobed lupine leaves are alternate, with 10-17 entire leaflets. There are obvious stipules at the base of each leaf. Unlike the Brewer’s Lupine and Mt. Ashland Lupine, this perennial is glabrous – no hairs. Big-leaf Lupine grows in moist soil in the wet meadows on the south slopes of Mount Ashland, tangled in with Western Columbine (Aquilegia formosa) and Giant Red Indian Paintbrush (Castilleja miniata ssp. miniata). Lupinus is Latin for wolf, from the mistaken belief that these plants robbed the soil of nutrients. Polyphyllus is Greek for many leaves, and refers to the very leafy stems. Pallidipes means pale; it is uncertain to what this refers. Long-stalk Clover Trifolium longipes Nutt. Native. Legume Family (Fabaceae). Clovers are not at all abundant on Mount Ashland. This low-growing perennial has many small, irregular flowers packed tightly into several terminal, head-like inflorescences. The flowers vary in color from dull white, to pale lavender. Five calyx lobes are long and very narrow with a few hairs along the margins. Ten tiny stamens and one small pistil are all hidden inside the keel. The palmately compound leaves have three narrow, finely-toothed leaflets. There are some alternate stem leaves and many basal leaves. The prostrate stems may be 2-8 cm long, but the flowers are never more than 4 cm from the ground. Long-stalk Clover grows in moist areas, next to running water. You will find it on Mount Ashland along USFS Road 20 between Road 300 and Road 40S30 that leads to Grouse Gap Shelter. It blooms in June, along with Small White Violet (Viola macloskeyi) and Alpine Shooting Star (Dodecatheon alpinum). In Latin, trifolium means three leaves, and refers to the number of leaflets on each leaf. Longipes means long-stalked, referring to the stems which are longer than usual for this genus. 126 © Stanford University Press Plant Descriptions Brewer’s Lupine (Lupinus breweri) Big-leaf Lupine (Lupinus polyphyllus var. pallidipes) Long-stalk Clover (Trifolium longipes) 127 Plant Descriptions White Clover Trifolium repens L. Non-native from Eurasia. Legume Family (Fabaceae). This is the clover with which we are all so familiar! Small white or greenish flowers grow in many head-like inflorescences. The flowers become reflexed with age. The typical clover leaves are palmately compound with three leaflets. They grow at the bottom of the flowering stem, so they appear to be basal leaves. White, papery stipules are at base of the leaf stems. The flowering stems may be 5-8 cm long, but because of the decumbent nature of this plant, the flowers are usually only 4-5 cm from the ground. White Clover is usually found in disturbed areas, and on Mount Ashland, you will find it along County Road 1151 at 6200 feet and lower. It blooms in late July and early August with Idaho Gum Weed (Grindelia nana) and Dalmatian Toadflax (Linaria genistifolia var. dalmatica). In Latin, trifolium means three leaves, and refers to the number of leaflets on each leaf. Repens is Latin for creeping and refers to the creeping nature of the above-ground stems. Oak Family (Fagaceae) Golden Chinquapin Chrysolepis chrysophylla (Hook.) Hjelmq. var. minor (Benth.) Munz Native. Oak Family (Fagaceae). This evergreen shrub has leathery leaves that are green on the upper surface and yellow or golden on the lower surface. The entire, alternate leaves are pointed at the tip, and appear to be folded along the main vein with the margins turned upwards lengthwise. The flowers are fairly plain and inconspicuous, and grow in short, spike-like clusters. The tiny flowers do not have petals, but they have six sepals. Some of the flowers have only stamens and some have only pistils. The fruits, however, are interesting and obvious. One nut is surrounded by spiny, bur-like bracts. Usually, several fruits mature together in one, large, spiny clump. Golden Chinquapin grows at the forest edge and around rocky outcrops. Look for it at Rabbit Ears, and along USFS Road 20. This shrub flowers sometime in July, and the fruits begin to appear in early August. In Greek, chrysolepis means golden scale, referring to the lower surface of the leaves. Chrysophylla is Greek for golden leaf, again referring to the lower surface of the leaves. Bush Chinquapin Chrysolepis sempervirens (Kellogg) Hjelmq. Native. Oak Family (Fagaceae). Leathery, evergreen leaves grow alternately on this shrub. The upper surface of the leaves are green, but the lower surface is a muted gold color. The leaves are entire and rounded at the tip, and the leaf blade is flat. The flowers are rather inconspicuous, growing on a short stiff spike. The flowers do not have petals, but have six small sepals. Some of the flowers have only stamens, and some have only pistils. The fruits make Bush Chinquapin easy to identify. One to three nuts closely surrounded by spiny, bur-like bracts. Several fruits usually mature together in a tight, spiny cluster. Look for Bush Chinquapin in the understory of the forest along the Pacific Crest Trail. The fruits begin to mature in August. Bush Chinquapin is very similar to Golden Chinquapin, but on Mount Ashland, you will find Bush Chinquapin growing in the understory of the forest along the Pacific Crest Trail, while Golden Chinquapin is often in the full sun along rocky outcrops. The leaves are also different. The leaves of Bush Chinquapin are rounded at the tip, and the entire leaf is flat. The leaves of Golden Chinquapin are pointed at the tip, and the leaf appears to be loosely folded along the main vein. In Greek, chrysolepis means golden scale, referring to the lower surface of the leaves. Sempervirens is Latin for always green, referring to the evergreen leaves. 128 Plant Descriptions Photo by Br. Alfred Brousseau, St. Mary’s College © Stanford University Press White Clover (Trifolium repens) © Stanford University Press Photo by Charles Webber, California Academy of Sciences Golden Chinquapin (Chrysolepis chrysophyllus var. minor) Bush Chinquapin (Chrysolepis sempervirens) 129 Plant Descriptions Geranium Family (Geraniaceae) Richardson’s Geranium Geranium richardsonii Fischer & Trautv. Native. Geranium Family (Geraniaceae). Five white petals with magenta lines surround the 10 stamens with pink anthers and the 5-lobed pistil. The alternate leaves are palmately lobed with usually 5 primary segments that are lobed again. At the base of the leaves and any place where the stem branches, there are red stipules. The branching stems reach to heights of 20-45 cm. The fruits are erect, elongated capsules with a beak at the tip. Richardson’s Geranium is found in wet, shady areas, surrounded by Arrowhead Butterweed (Senecio triangularis) and Sitka Valerian (Valeriana sitchensis). Geranium is Greek for crane, referring to the beak-like fruits. Richardsonii commemorates Dr. John Richardson who collected plants in the Pacific Northwest with Scouler, Nuttall, Drummond, and others. He was a naturalist and a surgeon. Gooseberry Family (Grossulariaceae) Wax Currant Ribes cereum Douglas var. cereum Also called Squaw Currant. Native. Gooseberry Family (Grossulariaceae). Wax Currant is an abundant shrub on Mount Ashland. The flowers vary in color from white to pink, but the colored part of the flower is actually the five sepals that are fused into a bell-shape. The petals are very tiny and white, and are attached in the sinuses of the calyx lobes. What, at first glance, appears to be green sepals are actually bracts at the base of the inflorescence. The leaves are alternate with three lobes, and the margins are irregularly toothed. The bark is usually gray, and there are no spines on this plant. Look for Wax Currant on the dry hillsides around Rabbit Ears and on the ski slopes. This deciduous shrub flowers in May and the red fruits mature around late August to early September. Growing nearby, you will probably find Greenleaf Manzanita (Arctostaphylos patula) and Tobacco Brush (Ceanothus velutinus). Ribes is an old Arabic name for the plants of this genus. Cereum comes from the Latin word for wax, and refers to the glossy upper surface of the leaves. Swamp Currant Ribes lacustre (Pers.) Poiret Also called Prickly Currant, Swamp Gooseberry, Black Gooseberry, and Black Swamp Gooseberry. Native. Gooseberry Family (Grossulariaceae). Ouch! The stems are prickly! This prostrate to ascending shrub has 3-9 spines at every node, and lots of prickly bristles between the nodes. The small flowers have five sepals that vary in color from green to purple. The sepals, which are 1.5 mm long, are larger than the tiny purple petals that are only 1 mm long. The sepals and petals open to reveal 5 stamens and one pistil with two styles. The flowers hang in racemes of 5-15 flowers. When mature, the shiny black berries have glandular hairs all over them. The alternate, deciduous leaves are somewhat maple-leaf shaped. They are deeply 3-7 lobed, toothed, and have just a few, sparse hairs, if any at all. The upper leaf surface is dark green, while the lower surface is lighter green. The older stems are covered with a beautiful cinnamon colored bark. On Mount Ashland, this shrub usually less than 1 meter tall but could be as tall as 2 meters. Swamp Currant grows along creeks and seeps. Look for it on the north slopes, just below the ski runs, along little streams. It may be surrounded by Mountain Alder (Alnus incana ssp. tenuifolia) and Columbia Wind Flower (Anemone deltoidea). The prickles of Swamp Currant have been known to cause allergic reactions in some individuals. Because of its prickly nature, some groups thought this shrub had special powers in warding off evil influences. Ribes is an old Arabic name for the plants of this genus. Lacustre is New Latin meaning of a lake, and refers to this species’ preference to grow near water. 130 Plant Descriptions Photo by Elizabeth Wasserman Richardson’s Geranium (Geranium richardsonii) Wax Currant (Ribes cereum var. cereum) Swamp Currant (Ribes lacustre) 131 Plant Descriptions Gummy Gooseberry Ribes lobbii A. Gray Also called Pioneer Gooseberry. Native. Gooseberry Family (Grossulariaceae). These scarlet pendant flowers will surely catch your eye during the month of May, but this thorny shrub might keep you from getting too close! The scarlet part of the flower is actually the five sepals that are reflexed when the flower is mature. The five petals are white and curled into tubes, and the five red anthers are strongly exserted. Flowers grow either singly or in small clusters of two or three. The leaves have three lobes that have shallow, rounded teeth. The bottom surface of the leaves is sticky from glandular hairs. There are usually three thorns at each node on the stems. The fruits are oblong, red berries with glandular bristles that ripen around late August to early September. Look for this shrub in the understory of the forest, especially along Road 40S30 beyond Grouse Gap Shelter. It is often found in the same areas as Sticky Currant (R. viscosissimum). Ribes is an old Arabic name for the plants of this genus. Lobbii commemorates William Lobb (1809-1863). From Cornwall, England, Lobb collected plants in the California and Oregon area from 1849-1856 for a company in Exeter, England. Red Flowering Currant Ribes sanguineum Pursh Also called Red-flower Currant, and Red Currant. Native. Gooseberry Family (Grossulariaceae). Because of the variability of this shrub, Red Flowering Currant can be difficult to identify. The flowering racemes can be erect or drooping, and usually hold 10-20 flowers. Five sepals are 4-5 mm long and can be white, pink, or red. The five smaller petals are about 2-3 mm long, and can be white, pink, or red. Five stamens and one pistil are protected by the sepals and petals. The glaucous berries are blueblack, and have glandular hairs. Alternate, deciduous leaves are more or less 5 lobed, with each lobe irregularly toothed. The upper surface has minute hairs that can only be seen with a hand lens, and the lower surface can be sparsely hairy or white from many woolly hairs. This shrub is usually only one meter tall at higher elevations, but at lower elevations can grow to nearly four meters tall. Ribes is an old Arabic name for the plants of this genus. Sanguineum is from the Latin word for blood, referring to the usual color of the flowers. Sticky Currant Ribes viscosissimum Pursh Native. Gooseberry Family (Grossulariaceae). Sticky is the best word to describe this deciduous shrub. The leaves, stems and pedicels have glandular hairs that produce a sticky oil. The flowers have five creamy white sepals that recurve as the flowers mature, and five creamy white petals that remain erect protecting the styles and stamens within. Between 4 and 15 flowers grow in loose axillary racemes, often nodding. The inflorescence is subtended by several green, spreading bracts. The simple, alternate leaves have three to five lobes, and each lobe is crenate. The leaves are also highly fragrant. The bark is gray to brown, and the new growth often appears to be pealing. There are no spines on this shrub. This shrub flowers in May or early June and the black fruits ripen around late August to early September. Look for Sticky Currant in the understory along the Pacific Crest Trail and along Road 40S30 beyond Grouse Gap Shelter. You may find Gummy Gooseberry (Ribes lobbii) growing nearby, or Large-leaf Sandwort (Moehringia macrophylla). Ribes is an old Arabic name for the plants of this genus. Derived from Latin, viscosissimum means the stickiest. Indeed, it is! 132 Plant Descriptions Photo by Br. Alfred Brousseau, St. Mary’s College Gummy Gooseberry (Ribes lobbii) Red Flowering Currant (Ribes sanguineum) Sticky Currant (Ribes viscosissimum) 133 Plant Descriptions Waterleaf Family (Hydrophyllaceae) White Waterleaf Hydrophyllum fendleri (A. Gray) var. albifrons (A.A. Heller) J.F. Macbr. Also called Fendler’s Waterleaf. Native. Waterleaf Family (Hydrophyllaceae). The color of these flowers varies from white to lavender, or white with lavender marks. The nodding cluster of several regular flowers sometimes towers over the leaves, and sometimes is hidden beneath them. The bell-shaped flowers have 5 petals, 1 pistil, and 5 stamens that are strongly exserted. The pinnately compound leaves are alternate and have 7-13 leaflets that are coarsely toothed. The lower pairs are often smaller, and distinctly separate from the other pairs. The stem and leaves are short-hairy. White Waterleaf can grow up to 45 cm tall, but is usually a bit smaller than that. You will find this abundant plant blooming in mid to late June in shaded areas and moist areas on the south slopes. Look for it near the picnic area along the Pacific Crest Trail. Nearby, you may find Starry False Solomon’s Seal (Smilacina stellata) or Showy Jacob’s Ladder (Polemonium californicum). The genus name, Hydrophyllum, is Greek for waterleaf, and refers to the large number of plants in this genus that grow near water. Augustus Fendler (1813-1883), a German botanist who collected many plants that were later identified by Dr. Asa Gray, lends his name for this specific epithet. Small-flower Nemophila Nemophila parviflora Benth. Native. Waterleaf Family (Hydrophyllaceae). This slender, little annual bears 3-7 tiny bell-shaped, axillary flowers. The flowers have 5 green sepals with short, stiff hairs, 5 white or pale blue petals, 5 stamens, and 1 pistil. The basal and opposite stem leaves have 5 lobes with pointed tips. The lobes are sometimes dissimilar. The two lower-most basal leaves are entire to crenate. This diminutive early bloomer grows only 3-8 cm tall. Small-flower Nemophila blooms mid-May through early June in wet meadows, beneath the large, pleated leaves of California False Hellebore (Veratrum californicum var. californicum), and tangled with another tiny annual, Low Mountain Bedstraw (Galium bifolium). Nemophila, from the Greek words nemos (grove) and philos (lover), means grove-loving or woodland-loving, and refers to its preferred habitat. In Latin, parviflora means small flower, and refers to the tiny flowers of this little annual. Silverleaf Phacelia Phacelia hastata Lehm. ssp. hastata Native. Waterleaf Family (Hydrophyllaceae). This perennial has an erect stem with many stiff hairs. It stands about 10-14 inches tall. The alternate leaves are highly variable. They are generally entire, but could be hastate with two to four lobes, or compound with 3 to 5 leaflets. The regular flowers have 5 white to purple petals with 5 greatly exserted stamens. The inflorescence is a one-sided, coiled cyme. You’ll find this one growing around Rabbit Ears, the ski area, and other gravelly slopes from mid-June through July. You may find Dusty Maidens (Chaenactis douglasii) or Siskiyou Penstemon (Penstemon anguineus) blooming nearby. Phacelia is Greek for bundle or cluster, referring to the dense inflorescence of most plants in this genus. Hastata is from the Latin for a spear, or spear-shaped, and refers to the arrowhead shape of some of the leaves. 134 Plant Descriptions Photo by Br. Alfred Brousseau, St. Mary’s College White Waterleaf (Hydrophyllum fendleri var. albifrons) © Stanford University Press Small-flower Nemophila (Nemophila parviflora) Silverleaf Phacelia (Phacelia hastata ssp. hastata) 135 Plant Descriptions Vari-leaf Phacelia Phacelia heterophylla Pursh Also called Variable Phacelia. Native. Waterleaf Family (Hydrophyllaceae). The lower leaves of this biennial are hastate with 3 lobes, while the upper leaves are entire. All alternate leaves have stiff hairs. The stem is erect with stiff hairs, growing 20-60 cm tall. The regular flowers have 5 creamy white to purple petals with 5 greatly exserted stamens. The inflorescence is a onesided, coiled cyme. Vari-leaf Phacelia may be found around Rabbit Ears, the ski area, along the road, and in dry, open areas along the Pacific Crest Trail through July. Phacelia is Greek for bundle or cluster, referring to the dense inflorescence of most plants in this genus. Heterophylla is also derived from Greek and means different leaves, referring to the two different kinds of leaves found on the same plant: entire, and hastate. Changeable Phacelia Phacelia mutabilis E. Greene Native. Waterleaf Family (Hydrophyllaceae). You will find this stiff-hairy biennial or short lived perennial by listening for the buzzzzz… Changeable Phacelia has lots of bumblebee visitors, as it is one of the few fresh flowers late in the season on Mount Ashland. Some plants are erect and tall, while others may be leaning toward the ground. The plant bears many one-sided, coiled cymes made up of white or cream radial, bell-shaped flowers. Five sepals and five petals cannot hide the five hairy stamens that are greatly exserted. The pistil, with its twolobed style, is usually not as tall as the stamens. Most of the alternate leaves are entire, but some of the lower leaves may be compound with 3 leaflets. This late-season plant is usually 10-30 cm tall. Changeable Phacelia prefers the dry soil of roadsides, and ridges. It blooms in September, along USFS Road 20 between Road 300 and Road 40S30 to Grouse Gap Shelter. Growing nearby, you should see Slender Tarweed (Madia gracilis), and further down the road, closer to road 40S30 you will find white specimens of Sickle-keel Lupine (Lupinus albicaulis). Phacelia is Greek for bundle or cluster, referring to the dense inflorescence of most plants in this genus. In Latin, mutabilis means changeable generally in reference to color, but it is uncertain exactly to what this refers. Tall Phacelia Phacelia procera A. Gray Also called Mountain Phacelia. Native. Waterleaf Family (Hydrophyllaceae). The radial flowers are usually white, cream colored, or brownish white. The bottlebrush-like cluster bears many of these flowers, which have five sepals, five petals, five stamens that are greatly exserted, and one pistil with a two-lobed style. Alternate leaves are lobed, and the lobes vary from entire to irregularly toothed. The erect stem, which grows 10-90 cm tall, is covered with hairs. This perennial blooms in late June through early July at the margins of wet areas, often tangled in with Big-leaf Lupine (Lupinus polyphyllus) and Western Bistort (Polygonum bistortoides). Phacelia is Greek for bundle or cluster, referring to the dense inflorescence of most plants in this genus. Procera is derived from the Latin word for tall, or high, referring to the usual height of this plant. 136 Photo by Bob Vos © Stanford University Press Plant Descriptions © Stanford University Press Vari-leaf Phacelia (Phacelia heterophylla) Photo by Br. Alfred Brousseau, St. Mary’s College Changeable Phacelia (Phacelia mutabilis) Tall Phacelia (Phacelia procera) 137 Plant Descriptions St. John’s Wort Family (Hypericaceae) Tinker’s Penny Hypericum anagalloides Cham. & Schldl. Also called Bog St. Johns Wort. Native. St. John’s Wort Family (Hypericaceae). To thoroughly appreciate this minute little flower, you will need a hand lens, and you will have to lie down on your belly. Tiny flowers like this are generically referred to as “belly flowers” because you have to lay on your belly to savor them. Five bright, yellow petals surround many stamens and a threelobed pistil. One radial flower blooms atop each short, prostrate stem. The tightly packed opposite leaves are entire and surprisingly fleshy for such a small plant. The bases of the leaves more or less clasp the stem. Many of these little gems usually grow together because as a result of many matted stolons – thin stems that runs along the ground sprouting roots and flowering stems as it grows. Tinker’s Pennies are found only on moist soil. While you are on your belly, you should also look for Primrose Monkey Flower (Mimulus primuloides) and Small-flower Nemophila (Nemophila parviflora), two other great belly flowers that grow in wet areas. The scientific name of this plant is a mystery. Hypericum is derived from Greek, but its meaning is unclear. Anagalloides is also derived from Greek and means Anagallis-like. Anagallis is the Greek word for pimpernel, and is a genus in the Primrose Family (Primulaceae). Klamath Weed Hypericum perforatum L. Also called Common St. John’s Wort, God’s Wonder Plant, Devil’s Scourge, Goat Weed, Grace of God, Rosin Rose, Amber Touch and Heal, Terrestrial Sun, and Hundred Holes. Non-native from Europe. St. John’s Wort Family (Hypericaceae). Many stamens and one three-lobed pistil make the center of this yellow flower appear bushy. The radial flower has five petals, the margins of which are often fringed. The petals and the anthers are dotted with black glands. Clusters of flowers grow in the axils of the opposite, entire leaves. The margins of the leaves are often rolled under. The stems are erect and usually green, mottled with red. One stem can produce 25-100 flowers. Klamath Weed grows 45-75 cm tall. On Mount Ashland, you will find it blooming in late June through early July along USFS Road 20 below the ski lodge. Growing nearby, you may find Hairy Cat’s Ear (Hypochaeris radicata) or Sheep Sorrel (Rumex acetosella). Klamath Weed has become very popular as a homeopathic treatment for mild depression. Recent research has also revealed that the toxin, hypericin, may inhibit human immunodeficiency virus. In spite of these benevolent characteristics, the plant is considered a noxious weed in the western United States because it is toxic to livestock. In order to control its spread, the flea beetle, Chrysolina quadrigemina, has been introduced to affected areas. The beetles eat the plant, preventing its growth and the development of more seed. In England, one of its native countries, Klamath Weed was hanged over doorways to bring prosperity. The genus name, hypericum, is derived from Greek, but its meaning is unclear. Perforatum is Latin and means to bore through, perhaps referring to the black dots on the leaves which might look similar to many small holes. 138 Plant Descriptions Tinker’s Penny (Hypericum anagalloides) Klamath Weed (Hypericum perforatum) 139 Plant Descriptions Iris Family (Iridaceae) Yellow-flowered Iris Iris chrysophylla Howell Also called Yellow-leaf Iris, and Slender-tubed Iris. Native. Iris Family (Iridaceae). I found only one small clump of these jewels on the second ski run. Although common in southwest Oregon, Yellow-flowered Iris is a rarity on Mount Ashland. These flowers have three long, white, narrow sepals that may be mistaken for petals. The three petals are whitish, perhaps with a yellow or bluish tinge, with dark purple veins. In the center of the flower, there are three style branches that are petal-like: long, white, and narrow. Each style branch arches over a stamen. There are many grass-like basal leaves that do not appear to be attached to any particular flowering stem. There are also a few stem leaves that fold protectively around the stem. Leaves typical of Irises have an interesting habit that Linnaeus termed equitant. The older leaves fold lengthwise around younger leaves, “like one astride a horse’s back.” As a general rule of thumb, irises are poisonous to humans, but the ancient Greeks valued them for their medicinal properties. Irises cured spleens, coughs, bruises, fits, dropsy, snakebites, and anger. Orrisrott, used in perfumes, comes from the roots of several Iris species. Iris, the winged messenger of the gods, was symbolized by the rainbow, and for her colorful cloak, the iris flowers were named. Thomas Howell, an Oregon botanist, named this plant in 1901 in A Flora of Northwest America. Although he chose the specific epithet chrysophylla, which means golden leaf, the leaves have very little yellow coloration. Blue-eyed Grass Sisyrinchium bellum S. Watson Also called Western Blue-eyed Grass. Native. Iris Family (Iridaceae). These delightful flowers seem to float in a sea of grass, but upon closer inspection, you will discover that this “grass” is actually the leaves of this plant. Each plant bears one or two deep blue-purple or blue-violet flowers, with 3 sepals that are identical to the 3 petals, 3 stamens and 1 pistil. The petal tips are sometimes notched. Two leaf-like bracts subtend the radial flowers. Grass-like leaves appear to be nearly opposite, and are folded along the mid-rib. The glabrous, erect stem is usually 10-20 cm tall, and generally flat. These delicate perennials prefer open, moist, grassy areas. Look for them in the wet meadows along the Pacific Crest Trail as you get close to Grouse Gap. I found them blooming in late June to early July between two clumps of Quaking Aspen (Populus tremuloides). Nearby, Tinker’s Pennies (Hypericum anagalloides) were also in bloom. Theophrastus called Irises by the name of Sisyrinchium. Theophrastus, who died in 287 BC, was a Greek philosopher and botanist. He was the favorite student of and immediate successor of Aristotle. Bellum is derived from the Latin word for beautiful. Ah! Mint Family (Lamiaceae) Nettle-leaf Giant Hyssop Agastache urticifolia (Benth.) Kuntze Also called Horsemint, and Nettle-leaf Horsemint. Native. Mint Family (Lamiaceae). This is one of only three mints found on Mt. Ashland above 6,200 feet. The creamy-white flowers are irregular, and have two stamens greatly exserted, and two stamens just barely exserted. From a distance, the flower heads may appear pinkish to purplish because of the colored sepals and bracts. The leaves are opposite, coarsely toothed, and slightly cordate at the base. The erect, square stem reaches heights of 45-60 cm. Nettle-leaf Giant Hyssop is common around the rock wall by the ski area restaurant, at the picnic area, along USFS Road 20, and along the Pacific Crest Trail in sunny areas. You’ll find this perennial from June through August, often growing near Coyote Mint (Monardella odoratissima) and Oregon Sunshine (Eriophyllum lanatum). Agastache is Greek for many spikes; urticifolia is Latin for nettle-leaf, referring to the shape of the leaf, which must have reminded someone of the nettle leaves. 140 © Stanford University Press Plant Descriptions © Stanford University Press Yellow-leaf Iris (Iris chrysophylla) © Stanford University Press Blue-eyed Grass (Sisyrinchium bellum) Nettle-leaf Giant Hyssop (Agastache urticifolia) 141 Plant Descriptions Coyote Mint Monardella odoratissima Benth. Also called Western Balm, Mountain Monardella, and Mountain Balm. Native. Mint Family (Lamiaceae). Try rubbing the leaves between your fingers and then smell your fingers...very aromatic. The flowers appear to have 5 petals, but actually the flowers are irregular bilabial: the upper lip has 2 lobes, and the lower lip has 3 lobes. The soft lavender petals are complemented by 4 dark purple stamens. Each flower is surrounded by a fuzzy calyx consisting of 5 fused lobes. The dense head of flowers is subtended by several purplish bracts that reflex with maturity. The leaves are opposite and entire. The erect stem is square and glabrous to hairy. This plant stands 25-35 cm tall. You’ll find this perennial blooming around Rabbit Ears in late June, along the road close to the ski area in early July, and in the picnic area through July. Milk Kelloggia (Kelloggia galioides) or Mountain Tarweed (Madia glomerata) may be growing nearby. In the days of outhouses, many flowers in the mint family were used as “strewing herbs” – herbaceous flowers that were scattered on the floors of outhouses so that when they were crushed by people’s feet, their aroma would enhance the outhouse experience. Monardella is Latin for small monarda. Monarda is another Mint genus, named for Nicolas Monardes (1493-1588), a Spanish physician and botanist. Derived from Latin, odoratissima means the smelliest – in a fragrant way, of course. Rough Hedge Nettle Stachys ajugoides Benth. var. rigida Jepson & Hoover Also called Rigid Hedge Nettle, and Hedge Nettle. Mint Family (Lamiaceae). A long, terminal spike bears whorls of sessile, bilateral flowers. Leaf-like bracts subtend each whorl, and the entire spike may have 6-16 flowers. The pink, magenta to purplish flowers have two lips: the upper lip is shorter than the tongue-like lower lip. The lower lip has magenta dots around its margin. Four stamens are tucked beneath the upper lip, and the pistil is hidden in the throat of the flower. The opposite leaves have coarsely toothed or scalloped margins, and the leaf bases are rounded to cordate. The leaves and the simple, erect stem are soft-hairy. Rough Hedge Nettle is usually 10-25 cm tall. This perennial grows in moist to dry places, usually along the margins of wet areas. It blooms from early- to mid-July through early August. You may find Large-flowered Boykinia (Boykinia major) or Tall Sneezeweed (Helenium bigelovii) growing nearby. Stachys is Greek for spike or ear of corn, referring to the inflorescence. Ajugoides means “Ajugalike.” Ajuga is another genus in the Mint family. Rigida is Latin for stiff, and refers to the erect stem. Lily Family (Liliaceae) Pacific Onion Allium validum S. Watson Also called Tall Swamp Onion. Native. Lily family (Liliaceae). Like many lilies, these radial flowers have six tepals that can vary in color from light lavender to pale pink to white. Six stamens are exserted from each corolla. Many flowers are arranged in densely clustered umbel. The number of basal leaves can vary from three to six; they are narrow and grass-like (linear), and sheathe the stem at the base. The glabrous stem rises 40-80 cm. Pacific Onion is a perennial that can be found in moist areas at the base of the ski slopes, and along the Pacific Crest Trail between USFS Road 20 and Grouse Gap. On the south slope, it grows near White-flowered Bog Orchid (Platanthera leucostachys). At the base of the ski slopes, look around for Columbia Windflower (Anemone deltoidea). Allium is Latin for garlic or onion; validum is Latin for strong, in reference to the strong onion smell of this plant. 142 Plant Descriptions Coyote Mint (Monardella odoratissima) Rough Hedge Nettle (Stachys ajugoides var. rigida) Pacific Onion (Allium validum) 143 Plant Descriptions Pussy Ears Calochortus tolmiei Hook. & Arn. Also called Tolmie’s Pussy Ears, and Tolmie’s Star Tulip. Native. Lily Family (Liliaceae). These low-lying but erect flowers might be missed at first glance. They often grow in the middle of other vegetation, so it’s difficult to distinguish which leaves belong to this little plant. The unique, regular flowers have 3 wide petals and 3 narrow sepals that are all similarly colored. The outer margins of the petals are white and hairy. The center of the flower has no hairs and is purplish. There are usually one to three flowers per plant. There is only one basal leaf that is entire, simple, and grass-like, and sometimes there will be one stem leaf. The flower is only about 4-5 cm from the ground, but the basal leaf usually extends over the flower by 15 cm. Look for Pussy Ears around Rabbit Ears and on other dry, grassy slopes from June through July. You may also find Common Blue Flax (Linum lewisii) or Torrey’s Blue-Eyed Mary (Collinsia torreyi var. latifolia). In Greek, calochortus means “beautiful grass” because the beautiful flowers have slender, grasslike leaves. Tolmiei honors William Fraser Tolmie (1812-1886) from Scotland. Tolmie collected plants primarily in Washington and Oregon from 1833-1859. Bride’s Bonnet Clintonia uniflora (Schultes) Kunth Also called Queen’s Cup, Queen Cup, Blue Bead, Queen’s Cup Beadlily, and Bead Lily. Native. Lily Family (Liliaceae). The solitary waxy white flower sits atop a leafless, glabrous stem that rises 7-15 cm. The regular flower has six tepals, six stamens with yellow anthers that rise like a crown from the center of the flower. One pistil has a three-lobed stigma. Tulip-like basal leaves are entire and shiny with hairy margins. The fruit is a single blue berry, hence Blue Bead, one of the common names. Bride’s Bonnet is found in coniferous forests. The Bella Coola Indians in British Colombia, Canada toasted the berries and the leaves and made a poultice that was used to ease eye soreness and to aid in the healing of cuts. However, they considered the berry poisonous to humans if eaten, hence their name for this flower, Wolf’s Berry. The generic name, Clintonia, commemorates DeWitt Clinton (1769-1828), governor of New York and botanist. The specific epithet, uniflora, is Latin for one flower. This indicates that each flowering stem will only produce one flower. Hooker’s Fairybells Disporum hookeri (Torrey) Nicholson Also called Drops of Gold. Native. Lily Family (Liliaceae). These nodding, narrowly bell-shaped flowers grace shady forested areas. The greenish, radial flowers have 6 tepals, 6 stamens that are as long as or longer than the petals, and 1 pistil with a 3-lobed stigma. The alternate leaves are sessile, entire and glabrous, but may be densely hairy on the veins on the underside of the leave. The stem, which is usually hairy above, grows 20-60 cm. Many of the northwest coastal tribes considered the berries of this plant poisonous, and associated it with snakes or ghosts. Disporum is Greek for double seed, perhaps meaning that each ovary chamber bears two ovules/seeds. Hookeri is named after Dr. William Jackson Hooker (1785-1865) who wrote Flora BorealiAmericana which was published in England in 1825. 144 Plant Descriptions Photo by Bob Vos Pussy Ears (Calochortus tolmiei) Photo by Bob Vos Bride’s Bonnet (Clintonia uniflora) Hooker’s Fairybells (Disporum hookeri) 145 Plant Descriptions Glacier Lily Erythronium grandiflorum Pursh Also called Yellow Avalanche Lily, Snow Lily, Adder’s Tongue, Yellow Adder’s Tongue, Lamb’s Tongue, Dog -Toothed Violet, Fawn Lily, Western Fawn Lily and Yellow Fawn Lily. Native. Lily Family (Liliaceae). This bright yellow flower has 6 tepals, a term referring to the petal-like structures when the sepals and petals are identical in appearance. The tepals are usually recurved displaying a long, white pistil with a three-lobed stigma and six yellow stamens. The flowers are either solitary at the end of a leafless stem, or two flowers can grow together on one stem. This plant has only two lanceolate basal leaves with entire but wavy margins. The leaves are solid green with no mottling. The entire plant is glabrous and can be as small as 10 cm tall and as large as 25 cm tall. Look for Glacier Lily in wet areas in May. The bulbs of this beautiful flower were prized by the Salish Indians in British Columbia. They might harvest up to 200 lbs per year, which they cooked and ate immediately or threaded and hung up to dry for winter use. Small rodents store them, and black and grizzly bears dig and eat them fresh. Because digging up the bulbs means the death of a lily, wholesale harvesting by humans is no longer practiced. The common names, Avalanche Lily and Glacier Lily, refer to this plant’s tendency to bloom at the edge of melting snow banks. Erythros is Greek for red. In this genus, the anthers are often red, but this particular species has anthers that can vary from yellow to creamy white. Grandiflorum means large flowered in Latin. Chocolate Lily Fritillaria atropurpurea Nutt. Also called Spotted Fritillary, Mission Bells, and Spotted Mountain Bells. Native. Lily Family (Liliaceae). This is the only flower on Mount Ashland that has brown and green/yellow mottled tepals. The six petal-like structures are called tepals because the sepals are identical to petals in appearance. The leaves are alternate and linear (grass-like). The stem is somewhat succulent and can grow from six to 18 inches tall. The entire plant is glabrous. There are six stamens and one pistil with a three-lobed stigma. Look for this one in May and early June in dry areas – near Rabbit Ears and just outside the wet meadows on the south slope. In Latin, fritillaria means dice box and refers to the shape of the fruit. Atropurpurea is Latin for blackish-purple, referring to the color of the tepals. Wiggin’s Lily Lilium pardalinum Kellogg ssp. wigginsii (Beane & Vollmer) M. Skinner Also called Leopard Lily. Native. Lily Family (Liliaceae). These nodding, striking yellow-orange flowers are a feast for the eyes. Six long tepals recurve displaying an array of reddish-purple dots toward the center of the flower. If you look closely, you will see the tip of each petal has short, fuzzy hairs. Six large orange-yellow anthers protrude from the center, surrounding the single stout, green pistil. The entire leaves grow in whorls on a tall, glabrous stem. Each stem produces one to several flowers. Wiggin’s Lily can grow as tall as one meter, but is usually about 60 cm. This perenial is found in wet areas, often next to running water. It may be surrounded by Oregon Checkerbloom (Sidalcea oregana) and Columbian Monskhood (Aconitum columbianum). Because of the spots on these flowers, there is a superstition in the northwest: anyone who smells Wiggin’s Lily or Leopard Lily will get freckles! Lilium is the Greek word for lily, and pardalinum is a derivative of Greek meaning spotted net or leopard cloth, obviously referring to the spots on the orange-yellow petals. 146 Photo by Bob Vos Plant Descriptions Glacier Lily (Erythronium grandiflorum) © Stanford University Press Chocolate Lily (Fritillaria atropurpurea) Wiggin’s Lily (Lilium pardalinum ssp. wigginsii) 147 Plant Descriptions Washington Lily Lilium washingtonianum Kellogg Also called Shasta Lily, Cascade Lily, and Mountain Lily. Native. Lily Family (Liliaceae). Large, waxy, white flowers have reddish-purple spots and mottling, and the petals turn completely purple with age. The nodding flowers have 6 tepals that are usually 8-12 cm long. The tepals recurve about 1/3 their length, displaying the single dirty yellow pistil and six stamens with large pale yellow anthers. The entire leaves grow in whorls around the tall stem that reaches 1-2 m high. Washington Lily is usually found in open woods and thickets in loose, dry soil at moderate altitudes. Although the bulbs are edible and were widely used by Native Americans, please leave these gorgeous flowers for nature. Lilium is the Greek word for lily, and washingtonianum means of Washington, referring to the general range of this species. False Solomon’s Seal Smilacina racemosa (L.) Link or Maianthemum racemosum (L.) Link ssp. racemosum Also called Western False Solomon’s Seal, Branched Solomon’s Seal, and ZigZag Solomon’s Seal, Feathery False Lily-of-the-Valley, and Scurvy Berries. Native. Lily Family (Liliaceae) Many small regular flowers create a dense terminal panicle that causes the top of the plant to nod or lean sideways. Upon close examination, you will find that the flowers have six whitish-yellow tepals. The leaves are alternate, simple, and entire with wavy margins. The parallel venation in the leaves indicates that this plant is a monocotyledon. The stem and leaves are glabrous, and on Mount Ashland most of the stems are no taller than 30 cm, although the plant never stands that tall as it is always bent in a graceful arc. In late summer, the bright red berries become obvious. Look for this one in shaded areas along the Pacific Crest Trail, at the picnic area and around Stonehenge – in the understory or beneath shrubs. It is often found along with Starry False Solomon’s Seal (Smilacine stellata) or Meadow Larkspur (Delphinium nuttallianum). The berries of False Solomon’s Seal are edible, and the name Scurvy Berries suggests that they contain an ample amount of vitamin C. Centuries ago, in New England, this plant was known as treacle berries because the berries taste similar to an English treat called treacle, a type of molasses. False Solomon’s Seal has been relied on for centuries as a food source and as a valuable medicinal plant. This plant is called False Solomon’s Seal because of its resemblance to Solomon’s Seal (Polygonatum biflorum). The underground portion of both of these plants is called a rhizome. At the end of the season, the above ground stem breaks off and leaves a scar on the rhizome. This round stem scar resembles the stamped-wax envelope seals of old. Stories claim that these seals were made by Solomon, a man who “knew the diversities of plants and the virtues of roots,” as a testimony of its value as a medicine. Smilacina is Greek for little smilax. Smilax is another genus in the Lily family; the Greek meaning of this word is unknown. Racemosa is derived from the Latin word for cluster and refers to this flower’s inflorescence, which is a panicle – a compound raceme. 148 Photo by Bob Vos Plant Descriptions Photo by Bob Vos Washington Lily (Lilium washingtonianum) False Solomon’s Seal (Smilacina racemosa) 149 Plant Descriptions Starry False Solomon’s Seal Smilacina stellata (L.) Desf. or Maianthemum stellatum (L.) Link Also called Star Solomon’s Seal, Few-flowered False Solomon’s Seal, and Starry False Lily-ofthe-Valley. Native. Lily Family (Liliaceae). A short raceme of 5-15 starry flowers sit atop a straight or zig-zapped stem. The white, radial flowers have 6 tepals, 6 stamens, and 1 pistil with 3-lobed stigma. Alternate leaves are sessile, entire, and minutely hairy below (you will need a hand lens to see the hairs). This perennial often is 15-35 cm tall. Starry False Solomon’s Seal blooms in mid- to late-June next to Brewer’s Mitrewort (Mitella breweri) and Baneberry (Acteae rubra) in moist, shaded areas. Look for these flowers along forested portions of the Pacific Crest Trail. Rhizomes were cooked and eaten by Native Americans. Tea from roots helped colds and increased appetites. Boiled leaf tea was used to prevent conception by drinking ½ cup daily for two weeks. Raw berries were eaten as a laxative. Smilacina is Greek for little smilax. Smilax is another genus in the Lily family; the Greek meaning of this word is unknown. Stellata is derived from the Latin word for star, and refers to the shape of the flowers. Western Tofieldia Tofieldia occidentalis S. Watson ssp. occidentalis Also called Sticky False Asphodel. Native. Lily family (Liliaceae). Small, white flowers are borne in a dense, terminal raceme. Each little flower has 6 tepals, 6 stamens with yellow anthers, and 1 pistil with 3 styles. Dense hairs at the top of the erect stem are tipped with black glands that contain sticky oils. The linear, grass-like leaves are mostly basal, but there may be 2 or 3 leaves at the bottom of the stem. This perennial is 20-45 cm tall. Western Tofieldia grows in wet meadows on the south slopes. Be careful not to mistake it for Western Bistort (Polygonum bistortoides). The easiest way to tell the difference is to feel the stem. If the stem is sticky, it is Western Tofieldia. If the stem is not sticky, it is Western Bistort. It blooms in early to mid-July with Mountain Larkspur (Delphinium glaucum) and Tall Sneezeweed (Helenium bigelovii). Thomas Tofield (1730-1779), a British botanist, is commemorated in this genus name. Occidentalis is Latin for western, referring to the range of this species. Western Trillium Trillium ovatum Pursh Also called Pacific Trillium, Western Wake Robin, Wood Trillium, and White Trillium. Native. Lily Family (Liliaceae). All parts of the trillium come in threes or multiples of three: three styles (and therefore three berries in fruit), three petals, three sepals, three leaves, six stamens. The common name, Wake Robin, comes from the fact that it is one of the earliest spring bloomers, coming with the early bird. Color changes from white to pink or rose with maturity. This is a signal to insects that the flower has been pollinated and the insects shouldn’t bother with it. Native women used trillium root to stop bleeding after childbirth, or to stop hemorrhaging. Recent studies of this plant indicate that one Western Trillium plant can live from 30-50 years. Western Trillium has an interesting relationship with ants. The seeds have oil-rich appendages that the ants love to eat. So, the ants drag the seeds to their nests where they eat the appendages, and then discard the seed outside of the nest. As a result of this mutualistic relationship, the ants receive food while the seeds of Western Trillium are carefully dispersed. This handsome plant was first recorded in 1806 by Capt. Meriweather Lewis (of the Lewis and Clark expedition), who found it near rapids on the Columbia River. It was officially named in 1814 by Frederick Pursh, a Polish botanist, who depended on Lewis’s description of the plant. Derived from Latin, trillium means three, and refers to the number of petals, sepals, and leaves. Ovatum is also Latin and means egg-shaped, probably referring to the leaf shape. 150 Plant Descriptions Starry False Solomon’s Seal (Smilacina stellata) © Stanford University Press Photo by Bob Vos Western Tofieldia (Tofieldia occidentalis) Western Trillium (Trillium ovatum) 151 Plant Descriptions California False Hellebore Veratrum californicum Durand var. californicum Also referred to as Corn Lily, and California Corn Lily. Native. Lily Family (Liliaceae). A stout plant of wet meadows with very distinct leaves, California False Hellebore is hard to miss. The huge, plaited leaves are obvious as early as late May, but the large, triangular panicle does not appear until July. Many white to greenish-white flowers have 6 tepals, 6 stamens, and 1 pistil with a 3lobed stigma. Using your hand lens to look at the ovary, you will find that it is glabrous. At the base of each petal is a dark green gland in the shape of a Y or V. Those alternate leaves are entire and glabrous above but tomentose beneath. California False Hellebore grows 1-2 m high in wet meadows, next to Oregon Checkerbloom (Sidalcea oregana) and Cow Parsnip (Heracleum lanatum). All False Hellebores contain toxic alkaloids. Ingestion of any part of the plant could result in extreme salivation, prostration, depressed heart action, difficult breathing, burning sensations in the mouth and throat, hallucinations, and headaches. Most poisonings result from misuse of the medicinal properties of the plant. Veratrum is derived from Latin and means dark root, referring to the black rhizome. Calfornicum is New Latin and means of California, referring to the general range of this species. Siskiyou False Hellebore Veratrum insolitum Jepson Also called Siskiyou Corn Lily. Native. Lily Family (Liliaceae). Siskiyou False Hellebore looks almost exactly like California False Hellebore (V. californicum), except for the ovary. Siskiyou False Hellebore has a woolly ovary that is easily seen with a hand lens, while California False Hellebore has a glabrous ovary. Also, the petals of Siskiyou False Hellebore are often irregularly toothed, almost appearing fringed. Siskiyou False Hellebore grows interspersed among California False Hellebore. Veratrum is derived from Latin and means dark root, referring to the black rhizome. Insolitum means unfamiliar or strange, referring to the lack of abundance of this species. There are very few of them on Mount Ashland. Green False Hellebore Veratrum viride Aiton Also referred to as Green Corn Lily, White Hellebore, Indian Poke, Devil’s Bite, and Indian Hellebore. Native. Lily Family (Liliaceae). This is a coarse, perennial with a leafy stem and greenish-yellow flowers in a large, loosely branched panicle, with the lower branches of the panicle drooping. The regular flowers have six tepals with a dark green V at it base. The large, alternate, entire leaves are vertically ribbed and clasp the hollow stem. This highly conspicuous plant grows 1-2 m high. Although not as common as California False Hellebore (V. californicum), you will see the leaves of this large plant as early as June 1st, long before the flowers appear in mid to late July. Look for it in wet seeps and wet meadows. Green False Hellebore contains toxic alkaloids that make this one of the most poisonous plants in the Pacific Northwest. To eat just a small portion of this plant would result in loss of consciousness and death. It is so powerful that people who have drank the water in which Green False Hellebore grows have reported stomach cramps, frothing at the mouth, blurred vision, lockjaw, vomiting, and diarrhea. Because of the alkaloids, this plant can also be used medicinally, and it was an important and respected medicine of the PNW coastal tribes. However, most poisonings result from misuse of the medicine. True hellebores are species of Helleborus in the Buttercup Family (Ranunculaceaea), and do not bear much resemblance to Veratrum species. Veratrum is derived from Latin and means dark root, referring to the blackish rhizome. Viride is Latin for green, and refers to the flower color. 152 Plant Descriptions California False Hellebore (Veratrum californicum var. californicum) © Stanford University Press Photo by Bob Vos Siskiyou False Hellebore (Veratrum insolitum) Green False Hellebore (Veratrum viride) 153 Plant Descriptions Flax Family (Linaceae) Common Blue Flax Linum lewisii Pursh Also called Wild Blue Flax, Western Blue Flax, and Prairie Flax. Native. Flax Family (Linaceae). These delightful flowers are regular with 5 sky blue petals that are veined in a deeper shade of cobalt blue, and are yellow in the center. Five stamens and five pistils protrude from the center of the flower. Each flower persists for only one day. They open in the morning, and the petals fall to the ground the following day. The inflorescence is a very loose, terminal cyme. The small leaves are simple, entire, and alternate. This perennial has a glabrous stem that grows to about 45 cm. It flowers in late June through early July, and can be found on the slopes around Rabbit Ears growing next to Siskiyou Mountains Owl’s Clover (Orthocarpus cuspidatus) and Spreading Ground Smoke (Gayophytum diffusum). The cultivated flax, L. usitatissimum, is not found on Mount Ashland. It is cultivated for flaxseed, or linseed, oil obtained from the seeds. For centuries, Egyptian “cotton” made from this plant clothed the Pharaohs, the priests, and the common man. Flaxseed capsules were found in a tomb dating back to 3100 BC. The genus name, linum, could be derived from the Greek word linon, and the Celtic word, lin, which mean thread, and refer to the fibers of the plant that are used to make linen. Or could come from the Latin word, lineus, which means flax blue. Lewisii honors Meriweather Lewis, of the famous Lewis and Clark expedition. Loasa Family (Loasaceae) Bushy Blazing Star Mentzelia dispersa S. Watson Native. Loasa Family (Loasaceae) For a plant with such a striking name, the radial flowers are unexpectedly small. A long slender calyx tube covered with barbed hairs is topped with 5 tiny lobes, about 1-2 mm long. Five small, yellow petals, about 2-3 mm, open fully to reveal many stamens and one pistil. In the cyme, a small, green, leaflike bract subtends each flower. The alternate stem leaves are sessile and entire, although a few may be shallowly toothed. Basal leaves are entire to lobed. This erect annual usually stands 7-35 cm tall, and the branching stem is a white/cream color with pink mottling. Stiff, barbed hairs that cover the stem, leaves, and calyxes make this plant very clingy – like velcro. In general, Bushy Blazing Star prefers sandy or rocky soils. On Mount Ashland, I have only seen this plant growing near the ski area parking lot, just inside the wooden fence next to a large rock. It blooms late July to early August with Mountain Tarweed (Madia glomerata) and Virginia Pepperweed (Lepidium virginicum var. virginicum). The derivation and meaning of the genus name, Mentzelia, is uncertain. Dispersa is derived from Latin and means to scatter, perhaps referring to the plant’s seed dispersal mechanism. 154 Plant Descriptions Common Blue Flax (Linum lewisii) Bushy Blazing Star (Mentzelia dispersa) 155 Plant Descriptions Mallow Family (Malvaceae) Oregon Checkerbloom Sidalcea oregana (Torrey & A. Gray) A. Gray Also called Oregon Sidalcea, and Spiked Sidalcea. Native. Mallow Family (Malvaceae). Lovely pink, radial flowers are a delightful surprise in the wet meadows on Mount Ashland. Protected by five sepals, the five rose-pink to purple petals are notched at the tip. Many stamens and one pistil make the flower center appear slightly bushy. The terminal spike-like raceme can be dense or open. The hairy, alternate leaves vary in shape along the stem. The lowest leaves are usually crenate, while the upper-most leaves are more or less palmately compound. The segments of the upper leaves are either entire or deeply lobed. The erect stem is also hairy and usually grows 30-150 cm tall. There are two subspecies of this perennial on Mount Ashland. Those with an open inflorescence are ssp. oregana, while those with a dense inflorescence are ssp. spicata (Regel) C. Hitchc. Oregon Checkerbloom brightens the wet meadows on the south slopes in July and August. You will find them entangled with Western Columbine (Aquilegia formosa) and Tall Sneezeweed (Helenium bigelovii). The derivation and meaning of the genus name, Sidalcea, is uncertain. Oregana is New Latin for of Oregon, referring to the general range of this species and subspecies. Spicata is Latin for spike, referring to the dense inflorescence of this subspecies. Evening Primrose Family (Onagraceae) Fireweed Epilobium angustifolium L. ssp. circumvagum Mosq. Also called Willow Herb, Blooming Sally, and Red Fireweed. Native. Evening Primrose Family (Onagraceae). Many people think Fireweed grows only in recently burned areas. Although it is true that this hardy pioneer species can grow in recently burned areas, it also flourishes in any open place, particularly along roadsides and on gravel bars. On Mount Ashland, you’ll find these familiar plants in various places along USFS Road 20. The bright magenta flowers have four wide, spoon-shaped petals and four narrow, linear reddish sepals. The 8 stamens mature before the long, stout pistil, which has a 4-lobed stigma. The flowers grow in a long raceme, with nodding buds at the top and the large, open flowers below. The alternate, entire leaves are long, narrow and lanceolate. The stout stem is primarily green with reddish-purple mottling, and grows from 60-120 cm. Fireweed spreads by seeds and by buds on the underground stem. Native Americans used the tough fibers in the stem as rope or twine for fishnets. They also ate the young leaves and new shoots as salad greens. This plant is also prized for its nectar. Many beekeepers seek out populations of Fireweed for the production of premium honey. The common name alludes to this plant’s amazing capacity to quickly colonize in a newly burned area. Epilobium is derived from two Greek words, epi – upon, and lobos – pod, and refers to the inferior ovary. Angustifolium is taken from two Latin words, angustus – narrow, and folium – leaf, and describes the shape of the leaves. Circumvagum is Latin for wandering around, and possibly refers to the tendency of this plant to appear in various open places. 156 Plant Descriptions ssp. oregana ssp. spicata Oregon Checkerbloom (Sidalcea oregana) Fireweed (Epilobium angustifolium var. circumvagum) 157 Plant Descriptions Tall Annual Willow-herb Epilobium brachycarpum C. Presl Also called Parched Fireweed. Native. Evening Primrose Family (Onagraceae). Four white, deeply notched petals distinguish these radial flowers. In the center of the petals, there are 8 stamens with yellowish anthers and one pistil. The flowers bloom in a very open raceme, usually mature fruits are obvious below the flowers. The fruits are slender, and elongated with four obvious angles. Alternate leaves are long and narrow and generally folded along the mid-vein. Tall Annual Willow-herb prefers dry, open hillsides on the south slopes. Growing nearby, look for its close relatives, Spreading Ground Smoke (Gayophytym diffusum) and Fireweed (Epilobium angustifolium var. circumvagum). Epilobium is a Greek derivative meaning upon the pod, and refers to the petals which sit upon the inferior ovary. Brachycarpum is Greek for short fruit, indicating that the fruits of this species are shorter than the fruits of many of the other species in this genus. Fringed Willow-herb Epilobium ciliatum Raf. ssp. glandulosum (Lehm.) P. Hoch & Raven Also called Common Willow-herb. Native. Evening Primrose Family (Onagraceae). A slender plant of wet meadows, Fringed Willow-herb has several small, pink to rose-purple flowers borne in a raceme. Each radial flower has 4 sepals, 4 petals that are generally notched at the tip, 8 stamens that are shorter than or equal to the one pistil, which is club-shaped. The leaves are generally opposite below, entire, and have conspicuous veins. The erect stem grows 15-90 cm, and is usually hairy. This perennial blooms in wet meadows from late June through early August, with Candy Flower (Claytonia sibirica) and American Brooklime (Veronica americana). Epilobium is a Greek derivative meaning upon the pod, and refers to the petals which sit upon the inferior ovary. Ciliatum is derived from the Latin word for small hair and refers to the hairs on the stem, inflorescence, and fruit. Also derived from Latin, glandulosum means full of glands and refers to the hairs in the inflorescence which often bear glands filled with sticky oil. Hornemann’s Willow-herb Epilobium hornemannii Reichb. ssp. hornemannii Native. Evening Primrose Family (Onagraceae). These pink-purple flowers have four petals that are notched about half way to the base. Each half of the petal has dark purple lines. When the flower starts to wither, the color turns to deep purple. The center of the flower holds 8 stamens and one long club-shaped pistil. Four sepals vary from green to reddish-purple, depending on the maturity of the plant, and are shorter than the petals. The flowers bloom in an open raceme, and usually there are mature fruits below the delicate flowers. The fruits are elongated and square in cross-section. The leaves are opposite below the inflorescence, and alternate through the inflorescence. They are shallowly toothed and on more mature plants, the leaf margins are often red. Look for this perennial in June in wet meadows, growing close to the water. You may find Candy Flower (Claytonia sibirica) or Three-leaf Wood Sorrel (Oxalis trilliifolia) growing nearby. Epilobium is a Greek derivative meaning upon the pod, and refers to the petals which sit upon the inferior ovary. Hornemannii honors Jens Wilken Hornemann (1770-1841) professor of botany and amateur ornithologist at Copenhagen, Denmark. 158 Plant Descriptions Tall Annual Willow-herb (Epilobium brachycarpum) © Stanford University Press Fringed Willow-herb (Epilobium ciliatum) Hornemann’s Willow-herb (Epilobium hornemannii var. hornemannii) 159 Plant Descriptions Milkflower Willow-herb Epilobium lactiflorum Hausskn. Also called Milkflower Willow-weed, and White-Flowered Willow Herb. Native. Evening Primrose Family (Onagraceae). Milkflower Willow-herb has several small to moderate sized white or pink flowers arranged in a nodding or erect raceme. The inflorescence is glandular. Each radial flower has 4 sepals, 4 petals that are generally notched at the tip, 8 stamens, and 1 pistil. The leaves are generally opposite below, entire, and petioles are more or less winged. The erect stem grows 15-50 cm, and is usually minutely hairy. This perennial blooms in wet meadows. Epilobium is a Greek derivative meaning upon the pod, and refers to the petals, which sit upon the inferior ovary. Derived from Latin, lactiflorum means milk-white flower and refers to the usual white color of the flower. Oregon Willow-herb Epilobium oregonense Hausskn. Native. Evening Primrose Family (Onagraceae). A solitary, terminal white blossom is held on a red, thread-like, short stem. The flower has 4 redpink sepals that are erect in flower, 4 white petals that are notched at the tip, 8 stamens, and 1 club-shaped pistil. Tiny leaves are opposite and entire. They are tightly packed at the base of the stem, and sparse above. The nodes along the stem are slightly swollen. The stem and leaves are glabrous, but the inferior ovary and the sepals have short, fine hairs that can only be seen with a hand lens. This slender, erect plant only grows 4-6 cm tall. This little waif of a perennial blooms in late July in saturated soils next to Tinker’s Penny (Hypericum anagalloides) and Primrose Monkey Flower (Mimulus primuloides ssp. primuloides). Epilobium is a Greek derivative meaning upon the pod, and refers to the petals, which grow above the inferior ovary. Oregonense is New Latin and means of Oregon, referring to the range of this species. Spreading Ground Smoke Gayophytum diffusum Torrey & A. Gray Also called Spreading Gayophytum. Native. Evening Primrose Family (Onagraceae). I love this common name! Tiny white flowers with 4 petals grow in the leaf axils of these highly branched plants. Eight stamens are exserted from the corolla. The small leaves are alternate and entire, and may appear to be opposite near the base of the stem. The slender, thread-like stems are glabrous, and can stretch as high as 60 cm. These delicate, whimsical flowers are abundant in the dry soils of the open meadows on the south slopes. You will find them tucked between larger, more obvious plants such as Common Blue Flax (Linum lewisii) and Yarrow (Achillea millefolium). Gayophytum is a Latin derivative meaning Gay’s plant. C. Gay (1800-1873) was the French author of the Flora of Chile. Diffusum is Latin for spreading, and refers to the branching nature of the plant. 160 © Stanford University Press Photo by Br. Alfred Brousseau, St. Mary’s College Plant Descriptions © Stanford University Press Milkflower Willow-herb (Epilobium lactiflorum) Oregon Willow-herb (Epilobium oregonense) Spreading Ground Smoke (Gayophytum diffusum) 161 Plant Descriptions Orchid Family (Orchidaceae) Calypso Orchid Calypso bulbosa (L.) Oakes Also called Venus’ Slipper, Lady’s Slipper, False Lady’s Slipper, Deer-head Orchid, Cytherea, Hider-of-the-North, Pink Slipper Orchid, and Fairy Slipper. Native. Orchid Family (Orchidaceae). As you are walking through the woods, you may see one small, bilateral pink flower atop a leafless stalk. The erect, glabrous stem is fairly slender and only grows up to 15 cm tall, with most of these plants only growing 8-10 cm tall. Sometimes, the color changes from pink, to yellow to white with maturity, making some people think it is a different species. The stem is pink or mauve with several pink bracts. This gorgeous plant has only one spherical basal leaf that becomes ridged and tinged with purple as it matures. Look for this one in wooded areas along the Pacific Crest Trail at lower elevations. This exquisite flower is disappearing at an accelerated rate as a result of urban encroachment and picking. Adding to its decline is the little-known fact that it was prized by Chinese herbalists. When it was plentiful, Chinese immigrants dug up the bulb and sent it to their homeland. This practice ceased long ago, but left its mark on the populations. Calypso was a sea nymph, a goddess, and queen of the island of Ogygia. To readers of Homer’s Odyssey, it was she who waylaid Ulysses for seven years. Like the goddess for whom it is named, the Calypso Orchid finds secluded places to its liking. In Latin, bulbosa means bulb, referring to the root, which is bulb-like. Phantom Orchid Cephalanthera austinae (A. Gray) A.A. Heller Also called Snow Orchid. Native. Orchid Family (Orchidaceae). This entirely white plant often becomes yellowish or brown with age. A terminal raceme bears bilateral, white flowers that are each subtended by one bract. The alternate leaves are scale-like, entire, and the leaf bases sheathe the stem. The glabrous stem grows 20-55 cm in decomposed litter of rich soil in coniferous forests. Look for Phantom Orchid in the forested areas at lower elevations. The only true saprophytic orchid, Phantom Orchid is dependent upon its relationship with a subterranean mycorrhizal fungus for nutrients. Cephalanthera is Greek for head-like anther, referring to the shape of the anthers. Austinae is from someone’s name, perhaps Stafford Wallace Austin (18601931), who collected and studied plants in the California area. Spotted Coralroot Corallorhiza maculata Raf. Native. Orchid Family (Orchidaceae). The clown-like, rounded lower petal of these irregular flowers characterize this plant. The lower petal is white with many small, purple dots. The upper two petals are yellowish with a few purple dots on them. The three sepals are yellowish with a reddish-purple margin. The flowers grow in a raceme along the upper half of the reddish-brown stem. The stem has no leaves, but there are some scale-like, reddishbrown sheaths that may be mistaken for leaves. On Mount Ashland, Spotted Coral-root usually grows between 12 and 25 cm tall. This perennial prefers the cool, moist shade of wooded areas where there is lots of organic material in the soil. Growing nearby, you may find a close relative, Striped Coralroot (Corallorhiza striata), or perhaps Pipsissewa (Chimaphila umbellata). All coralroots lack green stems and leaves and, therefore, do not photosynthesize. These plants are called saprophytes, which means that they obtain nutrients from decaying organic matter in the soil. Corallorhiza comes from the Greek words for coral and root, in reference to the branched root system which resembles coral. Maculata is Latin for spotted, and refers to the purple spots on the white flower petals. 162 Photo by Bob Vos Plant Descriptions Photo by Bob Vos Calypso Orchid (Calypso bulbosa) Phantom Orchid (Cephalanthera austiniae) Spotted Coralroot (Corallorhiza maculata) 163 Plant Descriptions Western Coralroot Corallorhiza mertensiana Bong. Also called Pacific Coralroot, Merten’s Coralroot, and Purple Coralroot. Native. Orchid Family (Orchidaceae). Most botanical references report that Western Coralroot’s rounded lower petal is white with two purple dots. But the plants that I have seen on Mount Ashland have an elongated lower petal with purple markings that look smeared. The two upper petals are yellowish with purple markings on the margins, and the three sepals are yellowish with red margins on the inner surface and reddish on the outer surface. The reddish-purple stem has no leaves; instead, there are brown, papery, sheaths. Western Coralroot grows to about 12 to 20 cm in shaded forests. Look for it along the Pacific Crest Trail and on the north slopes in the trees between the ski runs. Close by, there may also be Rattlesnake Plantain (Goodyera oblongifolia) or White-veined Wintergreen (Pyrola picta). Coralroots tend to grow in clumps. At times you will see flowers blooming next to the dried stalks of last years plants. Other times, last years stalks will not be accompanied by fresh plants of the current year. That is because these flowers sometimes “take a rest” for one year. But the roots below the dried stalks of last year will be alive and healthy. Corallorhiza comes from the Greek words for coral and root, in reference to the branched root system which resembles coral. Franz Carl Mertens (1764-1831), a German botanist, is honored in the species name. Striped Coralroot Corallorhiza striata Lindley Also called Madder Stripes. Native. Orchid Family (Orchidaceae). Think candy-striper. The three sepals and two upper petals are creamy white with narrow pink longitudinal lines. The lower petal is mostly purplish-pink with some white markings. The lower petal often looks toothed or fringed around the margin. The leafless stem is pinkish-purple and grows between 15 and 50 cm. As a saprophyte, Striped Coralroot is found in the cool, moist forest where there is lots of organic litter on the ground. Corallorhiza comes from the Greek words for coral and root, in reference to the branched root system which resembles coral. Striata is derived from the Latin word for streaked, referring to the purple lines on the sepals of most of these plants. The alternative common name, Madder Stripes, referring to the purple color, which is sometimes called madder-purple. 164 Plant Descriptions Photo by Bob Vos Western Coralroot (Corallorhiza mertensiana) Striped Coralroot (Corallorhiza striata) 165 Plant Descriptions Rattlesnake Plantain Goodyera oblongifolia Raf. Also called Western Rattlesnake Plantain, Large Rattlesnake Orchid, Rattlesnake Orchid, and Lattice Leaf. Native. Orchid Family (Orchidaceae). Instead of a flamboyant flower surrounded by modest leaves, here we have a spike of fairly inconspicuous orchid flowers surrounded by ornate basal leaves – deep green with white veins. The venation on the leaves may be highly variable. The veins are white because they do not contain chlorophyll. The creamy white flowers are borne on a fairly tall, green stem that has small, bract-like alternate leaves. Look for this one in the shade of the conifers on the ski slopes and along the Pacific Crest Trail on the north slopes. At first glance, the leaves may resemble White-veined Wintergreen (Pyrola picta). However, Rattlesnake Plantain leaves are longer, pointed, and have flat petioles. White-veined Wintergreen flowers are radial, not orchid-like. The Doctrine of Signs was an old theory subscribed to by many early herbalists. Because the markings on the leaves of this plant reminded early settlers of rattlesnakes, they believed that this plant would successfully treat rattlesnake bites. This species is not a true plantain, and it is not effective against snakebites, but it does have some medicinal value. A leaf rubbed between the fingers will separate into two halves. The inner moist surface soothes mosquito bites, burns, cuts, or scratches. John Goodyer was a 17th century English botanist who is remembered in this genus name. Oblongifolia is derived from the Latin, oblongus, which refers to the oblong basal leaves. Broad-leaved Twayblade Listera convallarioides (Sw.) Nutt. Also called Broad-lipped Twayblade. Native. Orchid Family (Orchidaceae). Small bilateral flowers are displayed in an open raceme at the top of an erect stem that bears only two stem leaves. The sepals and petals are brownish-maroon-greenish, and the lower petal is notched at the tip. The two stem leaves are more or less opposite, rounded, entire, and sessile. Broad-leaved Twayblade grows 10-25 cm tall in the moist shade of coniferous forests. Look for it on Mount Ashland below 6200 feet. This genus owes its name to Martin Lister (1638-1711), an English naturalist. Convallariodes is derived from the Latin word for valley, referring to the preferred habitat of this species. Green-flowered Bog Orchid Platanthera hyperborea (L.) Lindley Also called Northern Green Orchid. Native. Orchid Family (Orchidaceae). Because the flowers are small and green, this plant is often difficult to see in a wet meadow. But after you find the first one, you will see lots of them. The green bilateral flowers have a tongue-like lower lip. This lip is descending to slightly upcurved and measures 4-7 mm long. This lip is spurred from the back, and the spur is slightly shorter than the lip, slightly curved, and has a blunt tip. Many of these flowers grow in a dense spike-like inflorescence that is usually 5-15 cm tall. Alternate leaves are linear to elliptic, entire, and grow smaller toward the top of the stem. The overall plant height is 15-75 cm Green-flowered Bog Orchid generally prefers wet coniferous forests. On Mount Ashland, look for it in the wet meadows along the Pacific Crest Trail below 6400 feet from mid- to late-June. It may be surrounded by White-flowered Bog Orchid (Platanthera leucostachys) and American Brooklime (Veronica americana). Green-flowered Bog Orchid is very similar to Sparse-flowered Bog Orchid (Platanthera sparsiflora), which also grows in wet meadows. To distinguish the two, look at the inflorescence, the lip, and the spur. Green-flowered Bog Orchid has a dense inflorescence that is 5-15 cm tall, a lip that is 4-7 mm long, and a spur with a blunt tip. Sparse-flowered Bog Orchid has an inflorescence that is 15-40 cm long with fewer well-spaced flowers, a lip that is 6-10 mm long, and a spur with an acute tip. In Greek, platanthera means wide anther, referring to the morphology of the anthers in this genus. Also in Greek, hyperborea means of the extreme north, referring to the preferred range of this species. 166 Plant Descriptions Photo by John Gane Rattlesnake Plantain (Goodyera oblongifolia) Broad-leaved Twayblade (Listera convallarioides) Green-flowered Bog Orchid (Platanthera hyperborea) 167 Plant Descriptions White-flowered Bog Orchid Platanthera leucostachys Lindley Also called Sierra Crane Orchid, Sierra Bog Orchid, Cascade Crane Orchid, Tall White Bog Orchid, Rein Orchid, White Rein Orchid, Scent Candle, and Fragrant White Orchid. Native. Orchid Family (Orchidaceae). All orchids are gorgeous, exotic flowers, and this one is no exception. The white irregular flowers are quite unusual. There are three sepals that are the same color as the three petals. The upper sepal and two lateral petals form a “hood” over the stamen and style that are fused into a column. The two lower sepals and one lower petal, or lip are free and spreading, and the petal has a long spur on its underside. The flowers grow in a spike. The glabrous stem has alternate, entire leaves that are linear, or grass-like. This perennial can be found in wet meadows on the south slopes from late-June through July. Nearby, you may find Oregon Saxifrage (Saxifraga oregana) or Arrowhead Butterweed (Senecio triangularis). As some of the alternative common names suggest, this species is very fragrant – often smelled before it is seen. One person described its heady scent as a combination of cloves, vanilla, and mock orange. Mmmm. I’ll add a reminder here that none of the flowers on Mount Ashland should be picked. Orchids are especially fragile and will not withstand the rigors of being transplanted. Orchids are fairly rare in the wild because their seeds require very specific conditions for germination: the seeds must be penetrated by microscopic threads of a specific species of fungus. Until this occurs, the seeds will not germinate. Although one plant may contain millions of seeds, only a small percentage will reach maturity. Please do not pick or dig these beauties. Instead, try a camera or sketch pad. In Greek, plantanthera means wide anther, referring to the morphology of the anthers in this genus. Also in Greek, leucostachys means white spike, referring to the inflorescence. Sparse-flowered Bog Orchid Platanthera sparsiflora (S. Watson) Schltr. Native. Orchid Family (Orchidaceae). Sparse-flowered Bog Orchid is very similar to Green-flowered Bog Orchid (P. hyperborea), which grows in the same habitat. To distinguish the two, look at the inflorescence, the lip, and the spur. Sparse-flowered Bog Orchid has an inflorescence that is 15-40 cm long with fewer well-spaced flowers, a lip that is 6-10 mm long, and a spur with an acute tip. Green-flowered Bog Orchid has a dense inflorescence that is 5-15 cm tall, a lip that is 4-7 mm long, and a spur with a blunt tip. In Greek, plantanthera means wide anther, referring to the morphology of the anthers in this genus. Sparsiflora is Latin for scattered flowers, referring to the open, few-flowered inflorescence. Hooded Ladies’ Tresses Spiranthes romanzoffiana Cham. Also called Twisted Orchid. Native. Orchid Family (Orchidaceae). Oh, what a gem! These unusual, bilateral flowers are slightly modified from other orchid flowers. The upper sepal is fused to the lateral petals forming a hood that encloses the column. The margins of the petals and sepals are wavy, and the violin-shaped lower lip is deeply grooved below the middle with a glabrous tip on the upper surface. The white or cream flowers rise in spirals along the dense spike. A green, leafy bract subtends each flower. The basal and stem leaves are linear to oblong, but the stem leaves are generally shorter than the basal leaves. These erect perennials grow 17-30 cm tall. You will find Hooded Ladies’ Tresses in wet meadows. In Greek, spiranthes means coiled flowers, referring to the inflorescence. Nikolei Rumliantzev, or Count Romanzoff (1754-1826), is honored by this specific epithet. A Russian, Rumliantzev was an avid supported of the sciences, and he sent Kotzebue to explore Alaska. 168 Plant Descriptions Photo by Bob Vos White-flowered Bog Orchid (Platanthera leucostachys) Photo by Bob Vos Sparse-flowered Bog Orchid (Platanthera sparsiflora) Hooded Ladies’ Tresses (Spiranthes romanzoffiana) 169 Plant Descriptions Broom-rape Family (Orobanchaceae). California Ground Cone Boschniaka strobilacea A. Gray Native. Broom-rape Family (Orobanchaceae). The casual observer might mistake this plant for a fallen pine cone. But upon closer observation, the observer would probably notice two things: first, that there are no pine trees nearby, and second, that there are small, bilateral flowers between the numerous brown bracts. The flowers are usually purplish, but fade to a cream color with maturity. This plant has no leaves, and no stem. In fact, you will notice that there are no green structures on it at all. This indicates that the plant does not photosynthesize, and therefore, it does not make its own food. How does it obtain nutrients? It grows as a parasite on the roots of chaparral plants, especially on Greenleaf Manzanita (Arctostaphylos patula). This genus is very small with a total of only four species, 3 of which are native to North America. The fourth species is native to the Himalayan Mountains. This species was first officially described by the American botanist, Asa Gray in the Pacific Railroad Reports published in 1856. The genus name is from a fairly obscure Russian botanist named Boschniak. The specific epithet is Greek for pine cone. Broom-rape, the common family name, describes the tendency for species in this family to parasitize Scotchbroom plants. Clustered Broomrape Orobanche fasciculata Nutt. Native. Broom-Rape Family (Orobanchaceae). Clustered Broomrape is usually pale yellow but may have a pink or purple tinge. Many bilateral flowers grow in a cluster. The unusual aspect of this plant is that there are no green structures – there are no leaves, and the stem is either bright yellow (when the flowers first appear) or brownish. The bases of the yellow or brown sepals are fused into a tube with 5 pointed lobes at the mouth of the tube. The lobes are shorter than the tube. Look for Clustered Broomrape in the draw between where the road splits to go to the summit and Grouse Gap. It is found only on dry, bare soil. The absence of green structures indicates that this plant does not photosynthesize. Instead, it obtains its nutrients from the roots of the nearby Sulfur Flower (Eriogonum umbellatum). Clustered Broomrape is a parasite. The genus name, orobanche, is Greek for vetch strangler and refers to this plant’s parasitic nature. The specific epithet, fasciculata, is derived from Latin and means little bundle, referring to the tendency of these flowers to grow in clusters. Broom-rape, the common family name, describes the tendency for species in this family to parasitize Scotchbroom plants. 170 © Stanford University Press Photo by Br. Alfred Brousseau, St. Mary’s College Plant Descriptions © Stanford University Press California Ground Cone (Boschniaka strobilacea) Clustered Broomrape (Orobanche fasciculata) 171 Plant Descriptions Naked Broomrape Orobanche uniflora L. Also called One-Flowered Cancer Root, One-flowered Broomrape, and simply Broomrape. Native. Broom-Rape Family (Orobanchaceae). The single pinkish, purplish, or bluish flowers have 5 lobes. The flower may appear to be radial but it is truly bilateral with an upper lip that has two lobes and a lower lip that has 3 lobes. The base of the sepals are fused into a tube with 5 pointed lobes at the mouth of the tube. The lobes are longer than the tube. The delicate stem is whitish, bluish or brownish, and there are no leaves. You have to search for these flowers as they only grow from ½ to 2 inches tall. Look for them in the wet meadows along the Pacific Crest Trail in late June and July. This interesting flower has a unique life cycle compared to most of the other flowers that surround it. You’ll notice that there are no leaves on this plant. In fact, there are no green structures at all on this plant, because it does not photosynthesize. Instead, this flower obtains its nutrients from the roots of surrounding flowers. That is, this flower is a parasite that feeds, perhaps, on the roots of the Oregon Saxifrage (Saxifraga oregana) or the Alpine Shooting Star (Dodecatheon alpinum) that are also found in the wet meadows where this little flower appears. This flower has been known in Europe since 1578, when it was called Rapum genistae. It was known then that this flower was a parasite on the roots of broom plants (genus Genista), and other legumes. Hence the name, broomrape. The Greek, orobos, means vetch, or clinging plant, and ancho, means to strangle; this refers to the parasitic nature of this plant. Uniflora means one flower, referring to the habit of this plant to have only one or two flowers. Oxalis Family (Oxalidaceae). Three-leaf Wood Sorrel Oxalis trilliifolia Hook. Also called Trillium-leaf Sorrel, and Great Oxalis. Native. Oxalis Family (Oxalidaceae). You’ll probably notice the familiar basal leaves of this plant before you see the flower. These typical, heart-shaped sorrel leaves have three entire leaflets that fold down the center. If you push some of the leaves aside, you will find the coy little flowers. Three to nine nodding flowers grow in umbels. The radial flowers have 5 white to cream colored petals with greenish-lines guiding the pollinators to the center of the flower. The flowering stem does not have leaves. Three-leaf Wood Sorrel prefers moist, shaded areas. I’ve only seen it in the wet meadow below the ski area on the south side right along the Pacific Crest Trail. It grows at the foot of Candy Flower (Claytonia sibirica) and Big-leaf Lupines (Lupinus polyphyllus var. pallidipes). Oxalis is Greek for sour, referring to the oxalic acid, which gives the leaves a pleasantly sour taste. Trilliifolia is Latin for three leaves, referring to the three leaflets. Poppy Family (Papaveraceae) Steer’s Head Dicentra uniflora Kellogg Also called Longhorn Steer’s Head. Native. Poppy Family (Papaveraceae). These unusual, precious flowers are not at all abundant on Mount Ashland. Solitary, nodding flowers are considered biradial, meaning they can be divided into mirror-image halves in two ways, as opposed to only one way for bilateral flowers. The two ephemeral sepals are shed just after the flower appears. Four white to pink to lavender petals are divided into two pairs: the outer pair is free and have recurved tips, while the inner pair is fused at the tip, hiding the 6 stamens and 1 pistil. There are usually 1-3 basal leaves that are 2 or 3 times ternately dissected. The leafless flowering stem is glabrous, sometimes glaucous, and grows 3-7 cm tall. Steer’s Head blooms in June in dry, gravelly soil. So far, it has only been spotted near the picnic area, below USFS Road 20. Species of Dicentra contain poisonous alkaloids such as protopine. If any part of the plant is ingested, labored breathing and convulsions may occur. Dicentra is Greek for twice spurred, referring to the outer petals that are recurved. Uniflora is Latin for one flower, and refers to the solitary flower on each plant. 172 Plant Descriptions Naked Broomrape (Orobanche uniflora) Photo by Bob Vos Three-leaf Wood Sorrel (Oxalis trilliifolia) Steer’s Head (Dicentra uniflora) 173 Plant Descriptions Plantain Family (Plantaginaceae) English Plantain Plantago lanceolata L. Also called Narrow-Leaf Plantain, Ribwort, and Buckhorn. Non-native from Europe. Plantain Family (Plantaginaceae). Long stamens with white anthers make this plant conspicuous. The tall, leafless flowering stem bears one terminal, dense spike of tiny, radial flowers. You’ll need your hand lens to clearly see the 4lobed calyx, and the papery, colorless, 4-lobed corolla. The 4 stamens and 1 pistil are obvious. The erect stem is grooved or angled and has messy, short-hairs. The basal leaves are lanceolate, and taper gradually to the petiole. Leaf margins are entire, and conspicuous veins run lengthwise. This weedy perennial grows 15-45 cm tall. Look for English Plantain along County Road 1151 and other roadsides or disturbed areas below 6300 feet in late July through early August. It may be blooming next to Hairy Cat’s Ear (Hypochaeris radicata) or Yellow Salsify (Tragopogon dubius). In Europe, English Plantain was a highly prized medicinal herb of old. Plantago is Latin for sole of foot, referring to how easily this plant spreads to new areas “from the shoes of white man.” Lanceolata is Latin for spear and refers to the shape of the leaves. Phlox Family (Polemoniaceae) Large-flowered Collomia Collomia grandiflora Lindley Also called Grand Collomia. Native. Phlox Family (Polemoniaceae). This fairly common flower is a treat because of the subtle salmon colored flowers. Each regular flower has five petals, and the flowers grow in a tight terminal cluster. The alternate leaves are simple and entire. The stem is glabrous toward the base, and fuzzy toward the top, growing to heights of 10-20 cm. Look for this annual in dry soils, especially in open areas along the Pacific Crest Trail and along USFS Road 20 between Road 300 (leads to the summit) and Grouse Gap Shelter during July. You may also see Mountain Tarweed (Madia glomerata) or Common Blue Flax (Linum lewisii). Collomia is derived from the Greek word for glue, and refers to the wet seed surface. Grandiflora is Latin for large flower, and indicates that these flowers are large compared to the flowers of other species in this genus. Tiny Trumpet Collomia linearis Nutt. Native. Phlox Family (Polemoniaceae). Small pink and white radial flowers grow in a tight terminal cluster above many long, alternate leaves. The flower is shaped like a tiny trumpet. The five petals are mostly pink with some white at the base. Inside the yellow tube that makes up the center of the flower are five stamens and one pistil. The alternate leaves are entire and vary in shape from lanceolate (widest at the base and tapering to the tip) to linear (nearly parallel sides). The lower surface of the leaves is covered with many short hairs. These annuals usually grow to about 9-20 cm. Tiny Trumpets bloom in July in and at the edges of wet areas. Growing nearby, you may find the large leaves of California False Hellebore (Veratrum californicum var. californicum), or Rough Hedge Nettle (Stachys ajugoides var. rigida) in bloom. Collomia is derived from the Greek word for glue, and refers to the wet seed surface. Linearis is derived from the Latin work for line, and refers to the leaves which are long and grass-like, otherwise called linear. 174 Plant Descriptions English Plantain (Plantago lanceolata) Large-flowered Collomia (Collomia grandiflora) Tiny Trumpet (Collomia linearis) 175 Plant Descriptions Staining Collomia Collomia tincturia Kellogg Native. Phlox Family (Polemoniaceae). These diminutive annuals have pale pink, radial flowers with 5 petals. The petals often turn white with maturity. In the center of the flower are five delightful blue anthers exserted at different levels from the flower. The leaves are entire, opposite, and linear to lanceolate. The stems and the margins of the leaves are often reddish-brown. The stems and leaves have glandular hairs and, therefore, are sticky. Because these plants only grow to 4-7 cm, you must look closely to find them. They are most often found on open, dry, gravelly soil. Growing nearby, you may find Torrey’s Blue-eyed Mary (Collinsia torreyi) or Pussy Paws (Calyptridium umbellatum). Collomia is derived from the Greek word for glue, and refers to the wet seed surface. The Latin derivative, tinctoria, means that the plant produces a dye or a stain. Scarlet Gilia Ipomopsis aggregata (Pursh) V. Grant Also known as Skyrocket, Desert Trumpet, Fire Pink, Foxfire, Polecat Plant, and Skunk Flower. Native. Phlox Family (Polemoniaceae). These showy red-orange, trumpet-like, regular flowers consist of five petals that are fused together to form a tube. Some flowers grow in the axils along the stem and some are terminal, but all flowers grow along one side of the sticky stem. The simple, deeply lobed leaves are alternate, at times appearing opposite. This beautiful plant stands 20-40 cm tall, and can be found growing profusely on the south slopes of Mount Ashland from early June through August. Look for it along USFS Road 20, around Rabbit Ears, and in other open areas, blooming with Nettle-leaf Giant Hyssop (Agastache urticifolia) or Scouler’s Hawkweed (Hieracium scouleri). This plant is a biennial or perennial that dies after flowering only once. Hummingbirds pollinate these gorgeous flowers. Pink and yellow forms are also known, and at least one yellow plant grows on Mount Ashland – along Road 20 between the picnic area and the fork in the road where spur 300 goes to the summit. In the Rocky Mountains, plants that bloom later in the season are known to be white. No one is quite sure why this is so. One source claims that the Okanagan Indians prized the leaves of this plant for their tonic properties. Steeped until hot water turns bright green, the leaves were then removed and the medicine taken in small doses. Many other sources report that this plant is poisonous. Although it is true that many poisonous plants yield valuable medication if they are correctly prepared and administered, we should leave this plant alone. Scarlet Gilia (I. aggregata) was originally described by Frederick Pursh, a Polish botanist, in 1814. This confusing species has had up to twenty different scientific names since then. The common name, Gilia, honors eighteenth-century Spanish botanist, Felipe Luis Gil. Those who adhere to the Spanish pronounce the “g” softly – “hilia.” In Greek, ipomopsis means striking appearance, an appropriate designator for this flower and many of the others in this genus. Aggregata in Latin means to bring together, referring to the many flowers that bloom on one plant. 176 Plant Descriptions © Stanford University Press Staining Collomia (Collomia tinctoria) Scarlet Gilia (Ipomopsis aggregata) 177 Plant Descriptions Ball-head Ipomopsis Ipomopsis congesta (Hook.) V. Grant ssp. congesta Also called Ball-head Gilia. Native. Phlox Family (Polemoniaceae). These little flowers grow in dense, terminal clusters that look like little cotton balls. Occasionally, there are small axillary flowers just beneath the terminal cluster. The regular flowers have five white petals, and five stamens with delightful blue anthers. The stems are reddish with soft, white fuzz. The green leaves are alternate and simple, but deeply lobed, with three leaflets. There are white hairs on the leaves, as well. Usually, many flower stems grow together in a clump. The stems can be prostrate or erect, reaching 12-20 cm. Look for these perennial flowers at Rabbit Ears and on the summit in late June through early July. They will probably be blooming next to Ballhead Sandwort (Arenaria congesta var. congesta) or Rosy Pussytoes (Antennaria rosea). In Greek, ipomopsis means striking appearance, an appropriate designator for many of the flowers in this genus. Congesta is Latin for carried together, referring to the flowers that grow in tight clusters. Spreading Phlox Phlox diffusa Benth. Native. Phlox Family (Polemoniaceae). These low-lying, mat-forming showy, regular flowers with five petals vary in color from white, to magenta, to lavender. The simple, entire, needle-like leaves are tightly packed, so it’s difficult to tell whether they are opposite or alternate, but in fact, they are opposite. These perennial flowers have a woody stem beneath all the foliage. The flowers grow only 3-6 cm from the ground. Look for Spreading Phlox around Rabbit Ears, the Ski area, and other dry, open areas in early to mid June. Growing nearby, you may find Brewer’s Lupine (Lupinus breweri) or Pussy Paws (Calyptridium umbellatum). Phlox is a Greek word meaning flame, referring to the bright colors of the flowers. Diffusa is derived from Latin and means to spread out or apart, referring to the sprawling nature of this plant. Slender Phlox Phlox gracilis (Hook.) E. Greene Native. Phlox Family (Polemoniaceae). A low-growing annual, Slender Phlox bears small bright pink to white radial flowers with 5 sepals, 5 petals, 5 stamens, and 1 pistil. The stamens are all attached at different levels inside the corolla tube. The entire leaves are opposite below and alternate above. All upper parts of this little plant (leaves, flowers, tip of stem) are glandular. The growth habit of Slender Phlox is highly variable. It can be decumbent or erect, and highly branched or simple, but it usually only reaches 4-12 cm. This one grows in dry to moist areas, and blooms in early to mid-June. You may find Torrey’s Blue-Eyed Mary (Collinsia torreyi var. latifolia), or Staining Collomia (Collomia tinctoria) growing nearby. Slender Phlox can be easily confused with Staining Collomia (C. tinctoria). The best way to tell the difference is to pluck one flower from each plant and use your hand lens to look inside the corolla throat. If the stamens are all attached at different levels, the plant is Slender Phlox. If the stamens are attached at the same level, the plant is Staining Collomia. Another way to distinguish the two is to pluck a leaf and crush it. If it leaves a pink or purple color on your fingers, it is Staining Collomia. Phlox is a Greek word meaning flame, referring to the bright colors of the flowers. Gracilis is Latin for slender, referring to the delicate appearance of this plant. 178 Plant Descriptions Ball-head Ipomopsis (Ipomopsis congesta var. congesta) Photo by Willaim R. Hewlett, California Academy of Sciences Spreading Phlox (Phlox diffusa) Slender Phlox (Phlox gracilis) 179 Plant Descriptions Showy Jacob’s Ladder Polemonium californicum Eastw. Also called Moving Polemonium. Native. Phlox Family (Polemoniaceae). These flowers are usually found in the shade of conifers where the microhabitat is a little cooler and more moist than surrounding areas. The five petals of this regular flower are a soft blue, with a yellow center. The five white stamens are obvious. The five green sepals are fuzzy with soft, white hairs. The inflorescence is an open panicle. The usually erect stem is also fuzzy with dense, white hairs. The alternate leaves are pinnately compound with 11-25 leaflets; leaflets are entire, and also have soft, white hairs. These flowers stand about 15-25 cm tall, and often grow in dense clumps. These perennials are often found alongside Jessica’s Stickseed (Hackelia micrantha) and Meadow Larkspur (Delphinium nuttallianum). Look for this flower in any shady area with dry soil from mid to late June. Polemonium is Greek and could be named after Polemon, an Athenian philosopher, or it could be a derivative of the word, polemos, meaning war or strife. Californicum means of California and refers to the general range of this species. Buckwheat Family (Polygonaceae) Jayne’s Canyon Buckwheat Eriogonum diclinum Rev. Also called Buckwheat-Knotweed. Native. Buckwheat Family (Polygonaceae). These flowers superficially look like miniature versions of Sulfur Flower (Eriogonum umbellatum). They are bright yellow, with six petals and grow in a very compact umbel of umbels. The reddish-green flowering stalk rises above a mat of small basal leaves. The leaves are more or less spoonshaped and are covered with many short, white hairs, making them greenish-gray in appearance. The flowering stem is also covered with short, white hairs. The entire plant hugs the ground; the leaves are only 2-3 cm above the ground, while the flowering stem may be 7-10 cm long. Janyne’s Canyon Buckwheat grows only in the open, dry, gravelly soil on the summit of Mount Ashland. It blooms through July and early August. Nearby, you will probably see Silky Raillardella (Raillardella argentea) or Cobwebby Paintbrush (Castilleja arachnoidea). True buckwheat (Fagopyrum) that was cultivated in Europe for animal fodder has similar flowers to the species in the genus Eriogonum. Eriogonum is Greek meaning woolly knees, referring to the hairy nodes of some of the species in this genus. Diclinum is Greek for two beds, indicating that the flowers of this plant are usually imperfect - the stamens and pistils are in separate flowers. Naked Buckwheat Eriogonum nudum Benth. Also called Naked Eriogonum, and Barestem Buckwheat. Native. Buckwheat Family (Polygonaceae). This is a taller, spindly version of the ubiquitous Sulfur Flower (Eriogonum umbellatum). The tiny flowers have six bright yellow petals, with six yellow stamens exserted. The flowers are borne in cymes atop tall, slender, naked stems. The leaves are mostly basal, but there may be a few leaves at the base of the stems. Basal leaves have long stems and an egg-shaped blade with wavy margins. The lower surface of the leaves has many woolly, white hairs. The stems are highly branched and glabrous, and there is a reddish-brown ring at every node. Naked Buckwheat prefers dry, open areas. Look for it along County Road 1151 at 6200 feet and lower. Idaho Gum Weed (Grindelia nana) or Douglas’s Knotweed (Polygonum douglasii var. douglasii) may be growing nearby. Eriogonum is Greek meaning woolly knees, referring to the hairy nodes of some of the species in this genus. Nudum is Latin for naked, referring to the leafless stems. 180 Plant Descriptions © Stanford University Press Showy Jacob’s Ladder (Polemonium californicum) Jayne’s Canyon Buckwheat (Eriogonum diclinum) Naked Buckwheat (Eriogonum nudum) 181 Plant Descriptions Sulfur Flower Eriogonum umbellatum Torrey Also called Sulfur Eriogonum, Sulfur Flower Buckwheat, and Sulfur Buckwheat. Native. Buckwheat Family (Polygonaceae). These small bright yellow flowers are regular with 6 petals with 6 exserted stamens. The inflorescence is a terminal umbel, more or less flat across the top. The stem is very fuzzy with no leaves, however there is a whorl of bracts directly below the flowers. The bracts are simple, entire to very shallowly toothed. The fuzzy basal leaves are simple, entire, darker green on top, almost whitish/grayish below. This perennial sometimes forms dense mats with many basal leaves and erect flower stems. This plant grows to about 15-20 cm tall in dry, gravelly soils. Look for this flower along roadsides, at the ski area, and at Rabbit Ears from June through September. Early in the season, you may find Common Blue Flax (Linum lewisii) growing nearby. Later in the season, look for Mugwort (Artemisia douglasiana). There are many different varieties of this species on Mount Ashland, so don’t be confused if some of the plants are slightly different from each other. Eriogonum is Greek meaning woolly knees, referring to the hairy nodes of some of the species in this genus. Umbellatum Latin for umbrella, and refers to the inflorescence, an umbel, which is reminiscent of an upturned umbrella. Western Bistort Polygonum bistortoides Pursh Also called Mountain Meadow Knotweed, and American Bistort. Native. Buckwheat Family (Polygonaceae). These tiny radial flowers have 5 white or pink petals with 3-8 stamens that are barely exserted. The inflorescence is a dense, round to oblong panicle. The unbranched stem is glabrous, and grows to about 20-60 cm. The stem leaves are alternate, entire, and linear with no petiole. Basal leaves are petioled and narrowly lanceolate. Look carefully and you’ll find a brownish, papery sheathing stipule at the base of each leaf. This perennial is very common in wet meadows. It flowers in July with Gray’s Lovage (Ligusticum grayi) and Seep Monkey Flower (Mimulus guttatus). Polygonum is derived from the Greek words for many knees, referring to the swollen nodes of some species in this genus. Bistortoides means bistortus-like, referring to the species Polygonum bistortus. Davis’ Knotweed Polygonum davisiae A. Gray Native. Buckwheat Family (Polygonaceae). The radial flowers of this plant are small, creamy-white with five petals, and 2-5 flowers grow together in the leaf axils. The alternate leaves are lanceolate to linear with entire margins and short, fine hairs. They have very short petioles, if any at all. If you look closely at the base of the leaves, you will see a papery, sheathing stipule that is characteristic of this family. The lower half of the stem is usually red, while the upper half varies from red, to pink, to light green. The glabrous stem is often zig-zagged at the nodes. This plant usually grows erect, but at times appears decumbent. This perennial grows in gravelly soil, in open, sunny areas. It blooms in early to mid-June with Western Wallflower (Erysimum capitatum) and Spreading Phlox (Phlox diffusa). Polygonum is derived from the Greek words for many knees, referring to the swollen nodes of some species in this genus. The specific epithet, davisiae, honors John Jefferson Davis (1852-1937), an American physician and mycologist. 182 Plant Descriptions Sulfur Flower (Eriogonum umbellatum) © Stanford University Press Western Bistort (Polygonum bistortoides) Davis’ Knotweed (Polygonum davisiae) 183 Plant Descriptions Douglas’ Knotweed Polygonum douglasii E. Greene Native. Buckwheat Family (Polygonaceae). These very inconspicuous annuals need dry soil. The pink or white radial flowers are very small, and often the 5 petals do not open. The petal margins are usually red or green. Eight stamens and 1 pistil with 3 styles remain hidden inside. The alternate, entire leaves are sessile, and grow shorter toward the top of the stem. The leaves have one prominent vein. The stem is erect and more or less angled. It can be simple or branching. There are two subspecies of this plant on Mount Ashland. If the plant is branching and the inflorescence is open, you are looking at subspecies douglasii. If the stem is not branching and the flowers grow in the leaf axils, you have subspecies johnstonii (Munz) J. Hickman, commonly called Johnston’s Knotweed. Subspecies johnstonii blooms in June in the open forest, next to Staining Collomia (Collomia tinctoria) and Torrey’s Blue-Eyed Mary (Collinsia torreyi var. latifolia), while subspecies douglasii blooms along County Road 1151 in August next to Bird’s Foot Trefoil (Lotus purshianus var. purshianus) and Shasta Tansy Aster (Machaeranthera canescens var. shastensis). Polygonum is derived from the Greek words for many knees, referring to the swollen nodes of some species in this genus. Douglasii commemorates the Scottish botanist, David Douglas (1798-1834). Douglas explored and collected plants in what is now Oregon, Washington, and Idaho from 1825-1827. Johnstonii honors someone whose last name was Johnston. Sheep Sorrel Rumex acetosella L. Also called Common Sorrel, Common Sheep Sorrel, Sour Grass, Sour Weed, Red Sorrel, Cow Sorrel, Mountain Sorrel, Field Sorrel, or Horse Sorrel. Non-native from Europe. Buckwheat Family (Polygonaceae). It’s easy to miss this inconspicuous perennial amidst all the showy flowers. The tiny, regular flowers have six red tepals (it’s difficult to determine which is a petal and which is a sepal because they are all the same color). The branching plants can have many panicles of these red flowers. This plant has basal leaves and stem leaves. The lower stem leaves and basal leaves are similar in shape and size. They have a very distinct shape, referred to as hastate or sagittate. The smaller, upper leaves often appear entire. All stem leaves are alternate, and each leaf has a papery, sheathing stipule at its base. These plants can grow 10-25 cm tall. Look for them in July along the parking lot at the ski area, and roadside between the ski area and Grouse Gap Shelter. This plant is eaten by wildlife – birds eat the seeds, rabbits and deer eat the leaves, mice and squirrel eat seeds and leaves. Humans can also throw some seeds in a salad or soup. This plant is an indicator of acidic soil. Its flavor is distinctly acid, hence its specific name, acetosella, from the Latin acetum, acid. Pliny applied the name rumex for sorrel, which means sour. Although the leaves contain vitamin C, they also contain oxalic acid, which, in high quantities, is dangerous. The oxalic acid produces oxalate salts that prevent the body from absorbing calcium. 184 Plant Descriptions Douglas’ Knotweed (Polygonum douglasii var. douglasii) Sheep Sorrel (Rumex acetosella) 185 Plant Descriptions Willow Dock Rumex salicifolius J.A. Weinm. Native. Buckwheat Family (Polygonaceae). Willow Dock is a rare plant on Mount Ashland. Typical of all docks, the flowers are small and not at all showy. Six petal-like structures are arranged in two whorls: the outer three structures are small, about 1 mm, and green, while the three inner lobes are larger, about 4mm, and green-pink. Six stamens and 1 pistil with 3 fringed stigmas are at the flower center. Many flowers are packed tight in the erect, narrow panicle. Below the inflorescence, the stem is unbranched, ridges, and swollen at the nodes. There are only a few alternate stem leaves and several large basal leaves. The stem leaves have stipules that are fused, sheathing the stem above the nodes. All leaves are somewhat leathery with very wavy margins that are entire to crenate. This stout perennial grows 50-150 cm in wet meadows. I have only seen Willow Dock in one area on Mount Ashland. If you park at the confluence of USFS Road 20 and Road 40S30 and hike in the opposite direction of the road, you will walk over a saddle and downhill into a hidden wet meadow in the Ashland watershed. I found Willow Dock blooming in mid- to late-July next to California False Hellebore (Veratrum californicum). Rumex is Latin for sorrel, which means sour and refers to the oxalic acid in these plants, which give them a sour taste. Salicifolius is derived from the Latin words for willow and leaf, referring to the shape of the leaves. Purslane Family (Portulacaceae) Pussy Paws Calyptridium umbellatum (Torrey) E. Greene Also called Mount Hood Pussy Paws. Native. Purslane Family (Portulacaceae). If you see a gravelly hillside that doesn’t appear to have vegetation on it, look again. Often you will find this perennial growing in prostrate clumps on rocky soils. The regular flowers are difficult to discern, but they have four petals that are usually a creamy white color, turning to a deep rose color with age. The petals are surrounded by two conspicuous tissuey sepals that are the same color as the flowers. The inflorescence is a tight, terminal umbel. The stems are red and glabrous with no leaves. The green, basal rosette of leaves are simple, entire, and somewhat fleshy. Look for this flower anywhere there is loose, gravelly soil from May through August. It is usually found along with Brewer’s Lupine (Lupinus breweri), and some small annual flowers like Blue-eyed Mary (Collinsia parviflora). Calyptridium is Greek for small veil, referring to the petals, which form a veil, or covering over the fruit. Umbellatum is Latin for umbrella. The inflorescence of this plant is an umbel, which looks like an umbrella that has been pushed inside-out by the wind. Heart-leaf Spring Beauty Claytonia cordifolia S. Watson Also called Broad-leaf Montia. Native. Purslane Family (Portulacaceae). Delicate, white flowers have five pink-striped petals that are notched at the tip. The base of each petal has a yellow dot. In the center of the radial flower, 5 stamens with pink anthers surround a threelobed stigma. The raceme of flowers grows above one pair of opposite leaves that are wide, nearly heartshaped, and sessile. Several long-stemmed basal leaves are also present. The leaves are entire, and the leaves and stem are glabrous. These plants grow 10-40 cm in cool, shaded areas, and in moist areas. This genus is named after John Clayton (1685-1773), a colonial American botanist, who studied the plants in the eastern U.S., and contributed to Gronovius’s Flora Virginica. Cordifolia means heartshaped leaf in Latin. 186 © Stanford University Press Plant Descriptions Photo by Elizabeth Wasserman Willow Dock (Rumex salicifolius) Pussy Paws (Calyptridium umbellatum) Heart-leaf Spring Beauty (Claytonia cordifolia) 187 Plant Descriptions Miner’s Lettuce Claytonia perfoliata Willd. ssp. perfoliata Also called Gray Miner’s Lettuce. Native. Purslane Family (Portulacaceae). These small, delicate, solid white flowers have five petals that are notched at the tips. The center of the flower holds 5 stamens and one pistil with a three-lobed stigma. Several flowers rise above two opposite leaves that are fused together to form one complete leaf that is pierced through the middle by the stem. Basal leaves are long-stemmed and elliptic. All leaves and stems are glabrous. Look for these in the shade of trees along the Pacific Crest Trail. Growing nearby, you may find Wild Ginger (Asarum caudatum) or Heart-leaf Spring Beauty (Claytonia cordifolia). Local lore claims that the miners ate these plants as salad greens, hence the name, Miner’s Lettuce. This genus is named after John Clayton (1685-1773), a colonial American botanist, who studied the plants in the eastern U.S., and contributed to Gronovius’s Flora Virginica. Perfoliata is Latin meaning through the leaf, in reference to the stem, which appears to pierce through the middle of the leaves. Actually, two opposite leaves are fused together, completely surrounding the stem. Red-stem Spring Beauty Claytonia rubra (Howell) Tidestrom Native. Purslane Family (Portulacaceae). These low-growing, prostrate plants bear small white flowers with 5 notched petals. In the center of the flower, the three-lobed stigma is obvious. The 5 stamens are short, and not very noticeable. One to several flowers bloom above one pair of opposite leaves, and many basal leaves surround the flowering stem. The stems are brownish-red and the leaves vary from green to brownish-red. The upper surfaces of the leaves have short, white, woolly hairs. Look for this small annual in rich, moist soils of shady areas. I found it on the Knoll in early June. Growing nearby was Nuttall’s Toothwort (Cardamine nuttallii var. gemmatta) and Meadow Larkspur (Delphinium nuttallianum). This genus is named after John Clayton (1685-1773), a colonial American botanist, who studied the plants in the eastern U.S., and contributed to Gronovius’s Flora Virginica. Rubra is Latin for red, probably referring to the stems, which are often reddish-brown. Candy Flower Claytonia sibirica L. Also called Western Spring Beauty, Siberian Spring Beauty, and Siberian Miner’s Lettuce. Native. Purslane Family (Portulacaceae). The medium-sized white flowers of Candy Flower are obvious in shaded, wet meadows. The five petals are plain white with a yellow base. The tips of the petals vary from entire to notched. Five stamens with fairly large, pink anthers are obvious in the center of the flower, surrounding the three-lobed stigma. The terminal raceme rises above several pairs of large, opposite leaves. The leaves are entire and glabrous. The glabrous stem usually grows to 30-45 cm tall. Candy Flower blooms in June. Growing nearby, you may find Hornemann’s Willow-herb (Epilobium hornemannii var. hornemannii) and Western Bistort (Polygonum bistortoides). Native peoples of the Pacific Northwest used the leaves of this annual to treat syphilis, constipation, headaches, and female ailments. Before using any plant as a medicinal treatment, please consult an ethnobotanist, and research the plant and its uses as thoroughly as possible. This genus is named after John Clayton (1685-1773), a colonial American botanist, who studied the plants in the eastern U.S., and contributed to Gronovius’s Flora Virginica. The derivation and meaning of sibirica is uncertain. 188 Photo by Bob Vos Plant Descriptions Miner’s Lettuce (Claytonia perfoliata var. perfoliata) Red-stem Spring Beauty (Claytonia rubra) Candy Flower (Claytonia sibirica) 189 Plant Descriptions Primrose Family (Primulacaceae) Alpine Shooting Star Dodecatheon alpinum (A. Gray) E. Greene Native. Primrose Family (Primulacaceae). This glabrous plant has one to three nodding flowers that bloom at the end of a leafless stalk. Each flower has four magenta to lavender petals with an arc of white and yellow at the center. The petals are strongly recurved. The style is completely surrounded by four large, erect purplish-black anthers that appear wrinkled at the base. The erect stem is reddish-brown and grows to about 30 cm tall. The basal leaves are linear and grass-like, narrowed gradually at the base. Alpine Shooting Star flowers in May and early June in wet areas. Look for it in any wet meadow on Mount Ashland. Mr. Edward Greene saw in these blooms a convention of gods seated around Olympus; hence the genus name, Dodecatheon, from the Greek dodeka, twelve, and theoi, gods. Alpinum is Latin for mountains, referring to the habitat of this species. Western Starflower Trientalis latifolia Hook. Also called Pacific Starflower, Broad-leaved Starflower, and Indian Potato. Native. Primrose Family (Primulaceae). White, to pink, to rose colored flowers are radial with a variable number of parts: 5-7 sepals, petals, and stamens. If there are 5 sepals, there will be 5 petals and 5 stamens, but if there are 7 sepals, there will be 7 petals, and 7 stamens. There is always only one pistil. One flower rises on a very slender pedicel (flowering stalk) from a whorl of 4-8 leaves. These leaves are egg-shaped to elliptic, and entire. Other leaves lower on the stem are scale-like. The erect simple stem rises 5-15 cm from the forest floor. Look for this below 6200 feet in forest openings, or along the semi-shaded forest edge. The enlarged tubers of Western Starflower are edible, hence the alternate common name, Indian Potato. But let me add another reminder not to pick any part of the plants on Mount Ashland. Derived from Latin, trientalis means 1/3 of a foot, referring to the typical height of this plant. Also derived from Latin, latifolia means broad leaf and refers to the wide leaves. Buttercup Family (Ranunculaceae) Columbian Monkshood Aconitum columbianum Nutt. Also called Western Monkshood, Blue Monkshood, Western Aconite. Native. Buttercup Family (Ranunculaceae). The alternate leaves of this tall perennial have three to five deep palmate lobes, and the lobes are toothed. The inflorescence is a long, terminal raceme with many bluish-purple bilateral flowers. What appears at first glance to be petals are actually 5 petal-like sepals. The uppermost sepal is larger than all the others and forms a hood that protects the 2 blue to whitish petals. In the center are many stamens and 3 pistils. The plant grows to 150 cm, and is found in wet meadows. Look for this plant in July in the wet meadows on the south slopes. It will be surrounded by Bigleaf Lupine (Lupinus polyphyllus var. pallidipes) or Mountain Larkspur (Delphinium glaucum). This highly toxic plant is poisonous for both livestock and humans. The tubers contain an alkaloid, aconitine, that is paralytic, as well as a temperature and blood pressure depressant. It has also been known to cause respiratory failure. Aconitum is Latin for the monk’s hood, in reference to the shape of the upper petal which forms a hood-like structure over the top of the flower. Columbianum is derived from the Latin word for leadcolored with a sheen, possibly referring to the color of this plant. 190 Plant Descriptions Photo by Bob Vos Alpine Shooting Star (Dodecatheon alpinum) Western Starflower (Trientalis latifolia) Columbian Monkshood (Aconitum columbianum) 191 Plant Descriptions Baneberry Actaea rubra (Aiton) Willd. Also called Western Baneberry, and Red Baneberry. Native. Buttercup Family (Ranunculaceae). This bushy-looking perennial has one stem that produces one very large, ternately compound leaf. The pinnately branched leaflets are coarsely-toothed. Because there is only one leaf, this is classified as alternate. Don’t let the leaflets, which are opposite, confuse you. The flowering stalk rises from a node below the large compound leaf. The terminal raceme, which is subtended by a small compound leaf with three toothed leaflets, is a tight cluster of many white radial flowers. The flowers appear to be very “bushy” because of the many stamens. The small petals are difficult to find beneath the stamens. The number of petals can vary from four to ten, but all are small and spoon-shaped. This plant can grow up to 60 cm tall. Look for this one in the understory of forested areas along the Pacific Crest Trail below the ski area. It may be neighbored by Heart-leaf Spring Beauty (Claytonia cordifolia) or Wild Ginger (Asarum caudatum). This plant is probably most striking when in fruit – the berries are bright, shiny red. But beware: they are highly toxic to humans. In fact, the entire plant is toxic. Actaea is an ancient Greek name that means wet habitat, referring to the moist forests where these species grow; rubra is Latin for red, referring to the shiny, bright red berries. Columbia Windflower Anemone deltoidea Hook. Also called Three-leaf Anemone, Windflower, and Western White Anemone. Native. Buttercup Family (Ranunculaceae). This lovely flower is like a breath of fresh air. The five startling white “petals” are actually sepals. This flower has no true petals! The center of the flower is bushy with many pistils and stamens, a characteristic of the Buttercup Family. There is a whorl of three leaves several centimeters below the flower. These leaves are toothed and sometimes shallowly lobed. One compound basal leaf, with three toothed leaflets, will be found next to the flower. To find this perennial, you will have to search under shrubs near wet areas in early July. It grows in the same areas as Mountain Alder (Alnus incana), and Pacific Onion (Allium validum) may be nearby. A tea was made of this plant to treat tuberculosis. Some garden species of Anemone are toxic due to high levels of protoanemonin, an irritant. It is not yet know whether this is true of wild anemone species. Anemone is Greek for flower shaken by wind. This could refer to several different characteristics of this flower. It could refer to the fruits, which rely on the wind for dispersal, or it could refer to the petals which easily fall off as a result of a strong gust of wind. It is also possible that this word refers to the fact some of the flowers in this genus are found on exposed, wind-swept ridgelines. Deltoidea is Greek for triangular, perhaps because the three leaves form the overall shape of a triangle. 192 Plant Descriptions Baneberry (Actaea rubra) Columbia Windflower (Anemone deltoidea) 193 Plant Descriptions Drummond’s Anemone Anemone drummondii S. Watson Also called Alpine Anemone. Native. Buttercup Family (Ranunculaceae) There are no petals on this flower! Those lovely white structures are sepals that mimic petals. This perennial has between 5 and 8 white sepals that are sometimes tinged with blue, and have a softhairy lower surface. The flowers are radial and open completely, forming a flat surface. The center of the flower appears “bushy” because of the many yellow pistils and stamens. The flowering stem usually yields only one flower, but sometimes there will be two. There is a whorl of compound leaves around the middle of the stem. The leaflets are long and narrow. There are also several to many basal leaves that are similar to the stem leaves, but the stems are longer and they are covered with soft hairs. These plants reach 10 to 25 cm. Drummond’s Anemone grows on the north aspect of the summit of Mount Ashland, around the mouth of what many people call the bowl. The bowl area is open, dry, steep, and gravelly. Growing nearby will be Davidson’s Penstemon (Penstemon davidsonii) and Ballhead Sandwort (Arenaria congesta). There are nine species of anemone in Oregon. Some of them are capable of producing dangerous amounts of protoanemonin, a toxic irritant. Anemone is Greek for flower shaken by wind. This could refer to several different characteristics of this flower. It could refer to the fruits which rely on the wind for dispersal, or it could refer to the petals which easily fall off as a result of a strong gust of wind. It is also possible that this word refers to the fact some of the flowers in this genus are found on exposed, wind-swept ridgelines. Drummondii honors Dr. Thomas Drummond (ca 1790-1835), a Scottish naturalist. Drummond visited North America twice to make plant collections: the first trip to western Canada, and the second visit to Texas where he collected approximately 750 plant species and 150 bird species. White Pasque Flower Anemone occidentalis S. Watson Also called Western Pasque Flower, Western Anemone, Tow-headed Baby, and Moptop. Native. Buttercup Family (Ranunculaceae). If this is the first anemone you have seen, then you probably are not aware that those beautiful white structures are not petals! They are sepals that look like petals. Each flower can have between 5 and 8 sepals that are usually white, but sometimes tinged with purple. These radial flowers do not open completely, so the sepals form a bowl-like shape. The center of the flower is “bushy” as a result of many yellow pistils and stamens. The flowering stem generally produces only one flower. There is a whorl of three compound leaves at the center of the stem; the leaves are highly dissected and the leaflets are short and narrow. Several basal leaves look similar to the stem leaves but have soft hairs. White Pasque Flower is a perennial that reaches 20-75 cm and grows on open, rocky slopes. Look for this one on an open ridge along USFS Road 20, just beyond Grouse Gap. Anemone is Greek for flower shaken by wind. This could refer to several different characteristics of this flower. It could refer to the fruits, which rely on the wind for dispersal, or it could refer to the petals, which easily fall off as a result of a strong gust of wind. It is also possible that this word refers to the fact some of the flowers in this genus are found on exposed, wind-swept ridgelines. However, Pliny claimed that anemones are called windflowers because the flowers do not open when it is windy. Occidentalis is Latin for western, because this plant either grows only in the western United States, or was first discovered in the western United States. 194 Photo by Bob Vos Plant Descriptions Drummond’s Anemone (Anemone drummondii) Photo by Bob Vos fruits White Pasque Flower (Anemone occidentalis) 195 Plant Descriptions Little Mountain Thimbleweed Anemone lyallii Native. Buttercup Family (Ranunculaceae). These tiny flowers have no petals! Those white structures that look like petals are showy sepals. There are usually 5-6 sepals, but sometimes as many as 9. They can vary in color from white, to pale blue or pale pink. There are many stamens and pistils in the center of the flower, giving it a bushy appearance. One to few basal leaves and the whorl of 3 stem leaves are dissected into many narrow segments. These perennials grow at lower elevations on Mount Ashland. Anemone is derived from Greek and means flower shaken by wind. This could refer to several different characteristics of this flower. The fruits rely on the wind for dispersal, and the petals easily fall off as a result of a strong gust of wind. It is also possible that this refers to the fact some of the flowers in this genus are found on exposed, wind-swept ridgelines. However, Pliny claimed that anemones are called windflowers because the flowers do not open when it is windy. David Lyall (1817-1895), a British surgeon and botanist is honored in this species name. Lyall collected many valuable plant specimens from Antarctica, New Zealand, and western North America. Western Columbine Aquilegia formosa Fischer Also called Crimson Columbine, or simply Columbine. Native. Buttercup Family (Ranunculaceae). This beauty is not likely to be confused with any other plant on Mount Ashland. The distinctive regular red flower of this perennial has 5 petals, each with a spur that protrudes upward from the nodding flower. The inflorescence is a raceme of just a few flowers. The plant has basal and stem leaves that are lobed; each lobe has shallow teeth. This erect plant grows to 20-75 cm tall, and is usually found in moist areas. Look for it in late July in the wet meadow near Grouse Gap. The nectaries are in the back of the long spur, therefore, only long-tongued insects and hummingbirds can partake and act as pollinators. There are many species of Columbine throughout the U.S. At one time, there was a movement afoot to make the Columbine the national flower, but it never came to pass. Some Columbine species are also native to England. In medieval times, these columbines were thought to be a remedy against the plague. Derivation of the word aquilegia is uncertain. It could be Latin for eagle, the spurs being likened to talons, or it could be Latin for water-drawer in reference to the wet habitats where these flowers are most often found. Formosa is Latin for graceful or beautiful. The common name, Columbine, comes from columba, which means dove. The person who conferred this name on the plant must have seen a congregation of five doves in a circle. White Marsh Marigold Caltha leptosepala DC. var. biflora (DC.) G. Lawson Also called White-flowered Marigold, Alpine White Marsh Marigold, Twin-flowered Marigold, Broad-leaved Marigold, Broad-leaved Caltha. Native. Buttercup Family (Ranunculaceae) This plant is a trickster that might fool you! Starting with the common name – this plant does not have gold or yellow flowers. These beautiful flowers are white with many yellow stamens and pistils. And those showy white things that look like petals aren’t petals at all. This flower has no petals; those are actually sepals. The number of sepals can vary from 5 to 12. This plant has only basal leaves that are simple and entire with wavy margins. The shape of the leaves is called reniform, which means “kidney shaped.” You have to be on the mountain in May to see these plants in bloom, but you can find the fruits (follicles) towering over the leaves through the summer. Look for White Marsh Marigold along streams; they are often found next to its relative, Plantainleaf Buttercup (Ranunculus alismifolius var. alismifolius). The family name, Ranunculaceae, is a derivative of the Latin word, rana, which means frog. Several species in this family, like Marsh Marigold, are found in areas frequented by small frogs. Caltha is Latin for marigold; leptosepala is Greek for fine sepals, referring to the showy, petal-like sepals. 196 Photo by Bob Vos Plant Descriptions Little Mountain Thimbleweed (Anemone lyallii) Western Columbine (Aquilegia formosa) White Marsh Marigold (Caltha leptosepala var. biflora) 197 Plant Descriptions Anderson’s Delphinium Delphinium andersonii A. Gray Also called Anderson’s Larkspur. Native. Buttercup Family (Ranunculaceae). 10-25 bilateral flowers bloom in an open raceme. Each flower has 5 petal-like sepals that are dark blue. The uppermost sepal has a long spur on the back. The 4 petals are much smaller than sepals but similar in color to the sepals. The petals are obviously hairy. Many stamens and 3 pistils give the center of the flower a bushy appearance. Alternate leaves grow more or less on the lower half of the stem. They are glabrous and palmately lobed with 7-30 lobes. The erect stem grows 30-60 cm tall. Anderson’s Delphinium prefers dry soils. Delphinium comes form the Greek, delphin, or Latin, delphinus, which means dolphin, and could refer to the supposed resemblance of the long spur to the dolphin, or to the shape of the flower bud. Dr. Charles Lewis Anderson (1827-1910), physician and naturalist of western Nevada and California, is remembered in this specific epithet. Born in Virginia, he moved west and was one of the first to collect extensively in Nevada. He sent his collected specimens to Asa Gray of Harvard for identification, and many proved new to science. He compiled the first flora of Nevada. Yellowtinge Larkspur Delphinium decorum Fischer & C. Meyer Also called Coast Delphinium, Coastal Larkspur, and Low Larkspur. Native. Buttercup Family (Ranunculaceae). 2-20 bilateral flowers bloom in an open raceme. Each flower has 5 petal-like sepals that are dark blue-purple. The uppermost sepal has a long spur on the back. The 4 petals, which are the same color as the sepals, are much smaller than sepals. The petals are obviously hairy. Many stamens and 3 pistils give the center of the flower a bushy appearance. Leaves are mostly basal and are palmately lobed with 5 lobes. They are glabrous on the upper surface and minutely hairy on the lower surface. The erect stem grows 30-60 cm tall, and is hairy toward the base. Yellowtinge Larkspur grows in dry soils. Delphinium comes form the Greek, delphin, or Latin, delphinus, which means dolphin, and could refer to the supposed resemblance of the long spur to the dolphin, or to the shape of the flower bud. derived from the Latin word for dolphin, and refers to the shape of the of the flower bud. Decorum is Latin for elegant. Need I say more? Mountain Larkspur Delphinium glaucum S. Watson Also called Tower Delphinium, Pale Larkspur, and Sierra Larkspur. Native. Buttercup Family (Ranunculaceae). A tall, towering plant that grows in wet soil, Mountain Larkspur has many gorgeous purple-blue bilateral flowers. What appear to be five deep-purple petals are actually sepals that create a long spur at the back of the flower. The outer surface of the calyx is covered with many white, very short hairs. The four petals are much smaller than the sepals. The two upper petals are usually white with some purple at the tips, and the two lower petals are mostly purple with some white at the base. The lower petals cover the many stamens. The tall stems are sometimes branched toward the top, bearing the flowers in one to several racemes. The lower portion of the stem is glabrous and glaucous. Below the inflorescence, the deeply lobed leaves are coarsely toothed and grow alternately along the stem. The upper-most leaves are either lobed or entire. Mountain Larkspur prefers wet meadows, where it can grow to 150-180 cm. Growing nearby, you may find Western Columbine (Aquilegia formosa), or Columbian Monkshood (Aconitum columbianum). There are about two dozen species of Delphinium in Oregon. The young foliage and seeds are poisonous, as they contain various alkaloids. The toxicity varies with each species. Delphinium comes form the Greek, delphin, or Latin, delphinus, which means dolphin, and could refer to the supposed resemblance of the long spur to the dolphin, or to the shape of the flower bud. Glaucum is derived from the Latin word for gray, referring to the bloom (pronounced blome) on the lower half of the stem, making it glaucous. 198 Photo by Jeff Abbas Plant Descriptions Yellowtinge Larkspur (Delphinium decorum) © Stanford University Press Photo by Bob Vos Anderson’s Delphinium (Delphinium andersonii) Mountain Larkspur (Delphinium glaucum) 199 Plant Descriptions Meadow Larkspur Delphinium nuttallianum Walp. Also called Bilobed Delphinium, Dwarf Larkspur, Two-lobed Larkspur, Slim Larkspur, and Sonne’s Larkspur. Native. Buttercup Family (Ranunculaceae). This tall, showy flower is hard to miss in dry, shady areas. The five deep purple-blue petal-like structures are actually sepals. These can have a slightly fringed appearance, and they form a distinct spur at the back of this bilateral flower. The four petals are much smaller than the sepals, and are usually white with purple-blue veins. The upper two petals are notched at the tip, and the lower two petals are long-hairy and also notched. The lower two petals form a cover for the many stamens with yellow and purple anthers. The inflorescence is an open raceme, with a leaf-like bract subtending every flower. The alternate, glabrous leaves are simple, but so deeply palmately lobed that they may appear to be compound. The larger basal leaves are also deeply palmately lobed. These erect perennials have reddishbrown stems that grow 20-60 cm. Look for this flower in June dry butshady areas around Rabbit Ears and along the Pacific Crest Trail. All parts of this beautiful plant are poisonous. It contains a number of alkaloids, delphinine being among the worst. When cattle eat the fresh green plants in the spring, they grow weak, suffer from nausea, and at times, die. But, interestingly, other sources says that this plant does not harm sheep. Larkspur has been used as an insecticide, and soldiers once applied it externally to kill body lice. The common name, Larkspur, probably refers to the long hind toe of larks. Because these birds are ground walkers, they require a long toe (the spur) for balance. Delphinium comes form the Greek, delphin, or Latin, delphinus, which means dolphin, and could refer to the supposed resemblance of the long spur to the dolphin, or to the shape of the flower bud. Thomas Nuttall (1786-1859), a British-born, Philadelphia-raised botanist, is commemorated in this specific epithet. Nuttall traveled west three times to collect plants, from 1811-1812, 1818-1820, 1834-1836. He was lecturer and curator of the Botanic Garden at Harvard University from 1825-1834. Plantain-leaf Buttercup Ranunculus alismifolius Benth. var. alismellus A. Gray Also called Gray Plantain-leaved Buttercup, and Water Plantain Buttercup. Native. Buttercup Family (Ranunculaceae). One of the earliest flowers to bloom on Mount Ashland, this is also one of the brightest. These radial flowers have 5-7 bright, shiny yellow petals that hide 5 reflexed sepals. Many yellow stamens and many pistils mark the center of the flower. There are alternate stem leaves on the lower portion of the stem and many basal leaves. Stem leaves tend to be linear and entire, while basal leaves tend to be spoon-shaped or lanceolate and entire, with a long petiole. Stems are either decumbent to erect, and these flowers tend to grow in large clumps. This perennial blooms in May along streambanks next to White Marsh Marigold (Caltha leptosepala var. biflora). Ranunculus is Latin for little frog, referring to the wet habitats of most species in this genus. Alismifolius is Greek for Plantain leaf, referring to the shape of the leaves. Popular Buttercup Ranunculus populago E. Greene Native. Buttercup Family (Ranunculaceae). Popular Buttercup is a radial flower with 5 green sepals, 5 shiny yellow petals, many stamens, and many pistils. There are basal leaves and alternate stem leaves that vary in shape from heart-shaped to egg-shaped. They are entire to barely toothed. The erect stem of this perennial is few-branched and glabrous, and grows 8-30 cm tall. This one grows in wet meadows and along streambanks. The sap of most flowers in the Buttercup family contains irritants that blister the skin with prolonged contact. Ranunculus is Latin for little frog, referring to the wet habitats of most species in this genus. Populago is derived from the Latin word for the poplar, referring to the leaves which are shaped like poplar leaves. 200 Plant Descriptions Meadow Larkspur (Delphinium nuttallianum) Plantain-leaf Buttercup (Ranunculus alismifolius var. alismellus) Popular Buttercup (Ranunculus populago) 201 Plant Descriptions Fendler’s Meadow Rue Thalictrum fendleri A. Gray var. fendleri Native. Buttercup Family (Ranunculaceae). This is a very special plant on Mount Ashland because it is dioecious. That is, each plant has only staminate (male) flowers or only pistillate (female) flowers. Dioecious is derived from Greek and means two houses, referring to the separate plants for staminate flowers and pistillate flowers. If you find a male plant, the radial flowers will have 4 or 5 greenish-white to purplish sepals, no petals, and 15-28 stamens. The female plants have radial flowers with 4 or 5 greenish-white to purplish sepals, no petals, and 7-20 pistils. In every other way, the plants are the same. The inflorescence is a panicle that is leafy or bracted above. Basal leaves and alternate stem leaves are pinnately compound with lobed leaflets. These erect perennials grow 60-200 cm tall. You will find Fendler’s Meadow Rue in wet meadows, blooming in mid- to late-June, and going to fruit in late-June to early-July. Look for Mountain Sagebrush (Artemisia tridentata var. vaseyana) and Bigleaf Avens (Geum macrophyllum) blooming nearby. Thalictrum is a name given by Dioscorides, a Greek physician-botanist. Fendleri remembers Augustus Fendler (1813-1883), a German botanist who collected many plants that were later identified by Dr. Asa Gray. Buckthorn Family (Rhamnaceae) Mahala Mat Ceanothus prostratus Benth. Also called Squaw Carpet. Native. Buckthorn Family (Rhamnaceae ). This prostrate shrub is rather unique with its holly-like leaves. The opposite, evergreen leaves are coarsely toothed. The upper surface is green and glabrous, while the lower surface is paler green with hairy veins. The opposite twigs are round and red-brown becoming gray-brown. The radial flowers can be light blue, deep blue, lavender or purple. Five sepals and five petals are all colored the same. In the center are 5 stamens and 1 pistil with 3 styles. Look for Mahala Mat in forested areas at lower elevations. The seeds of this shrub were eaten by Native Americans, and the leaves were smoked or made into tea. Tonics and astringents were made from the bark and roots. Ceanothus is Greek for thorny plant, or a kind of thistle, because some members of this genus have thorns. The Ceanothus species on Mount Ashland do not have thorns. In Latin, prostratus means to lay flat, indicating that the branches of this shrub lay on or close to the ground. Tobacco Brush Ceanothus velutinus Dougl. var. velutinus Also called Mountain Balm, Sticky Laurel, and Snowbrush Ceanothus. Native. Buckthorn Family (Rhamnaceae ). An erect shrub, Tobacco Brush has sprays of many small, white, radial flowers with 5 petals. The leaves are alternate, evergreen, and aromatic when crushed. Perhaps the easiest way to identify this shrub is by looking at the shallowly toothed leaves – they have three prominent veins that are hard to miss. Look for Tobacco Brush in open areas on the south slopes. Growing nearby, you should see Greenleaf Manzanita (Arctostaphylos patula), and perhaps Bitter Cherry (Prunus emarginata). These shrubs are important in the poor, granitic soil on Mount Ashland because the roots have a mycorhizal fungi on them that makes nitrogen available to plants. This shrub is not shade tolerant, and so will only grow in open, sunny areas. Ceanothus is Greek for thorny plant, or a kind of thistle, because some members of this genus have thorns. The Ceanothus species on Mount Ashland do not have thorns. Velutinus is New Latin for velvety, probably referring to the twigs which have microscopic hairs that make the plant feel velvety. 202 Photo by Steven Thorsted Plant Descriptions Photo by Charles Webber, California Academy of Sciences Fendler’s Meadow Rue (Thalictrum fendleri var. fendleri) Mahala Mat (Ceanothus prostratus) Tobacco Brush (Ceanothus velutinus var. velutinus) 203 Plant Descriptions Rose Family (Rosaceae) Serviceberry Amelanchier alnifolia (Nutt.) Nutt. Also called Western Service Berry, and Saskatoon. Native. Rose Family (Rosaceae). This medium-sized deciduous shrub has grayish bark and showy white, radial flowers with five petals. The center of the flowers appears “bushy” because of the many stamens and styles. The leaves are alternate and are coarsely toothed along the upper 2/3 of the margins. There are two varieties of this flower on Mount Ashland. To distinguish between the two, you need to look at the top of the ovary with a hand lens. If the ovary top is glabrous, you are looking at Dwarf Serviceberry, variety pumila (Nutt.) Nelson. If the ovary top is covered with densely interwoven, matted hairs, you are looking at Saskatoon Serviceberry, variety semiintegrifolia (Hook.) C. Hitchc. Serviceberry blooms fairly early on Mount Ashland, in late May and early June. It is obvious along the side of USFS Road 20 between the picnic area and the Rabbit Ears, growing next to Bitter Cherry (Prunus emarginata) and Greenleaf Manzanita (Arctostaphylos patula). Some Native Americans dried the berries and used them in pemmican. Some groups used the wood to make arrows, digging sticks, and drying racks. This shrub is an important winter browse for the deer on Mount Ashland, and many birds rely on the berries in August. Stories say that early colonists named this shrub or small tree Serviceberry. When the flowers of this tree bloomed, they knew the soil was soft enough to bury those who had died in the winter. So, they held a service for their burial. Amelanchier is a Latin modification of the old French common name; alnifolia is Latin for alder leaf. The leaves must have reminded someone of alder trees. Pale Serviceberry Amelanchier pallida E. Greene Native. Rose Family (Rosaceae). Glabrous red-brown to gray twigs bear alternate, simple leaves that are entire to toothed. The white, radial flowers have 5 sepals, 5 petals, about 15 stamens, and 3-4 styles. The fruits are purplishblack pomes. Look for Pale Service Berry on dry, gravelly slopes. This deciduous shrub looks very similar to Serviceberry. Look closely at the leaves. If the underside is paler than the upper surface, it is Pale Serviceberry. Also look at the number of styles. If there are 5, it is Serviceberry (A. alnifolia). Amelanchier is a Latin modification of the old French common name; pallida is Latin for pale, possibly referring to the pale lower surface of the leaves, or referring to the white flowers. Wood Strawberry Fragaria vesca L. Also called Woodland Strawberry. Native. Rose Family (Rosaceae). These radial flowers have 5 green sepals, 5 white petals, 20-35 yellow stamens, and many yellow pistils. The flowers are usually taller than the compound basal leaves. Each leaf is made up of 3 toothed leaflets. The bright red fruit looks like a miniature store-bought strawberry. This perennial will be found in forested areas at lower elevations. There are two subspecies of Wood Strawberry on Mount Ashland: ssp. bracteata and ssp. californica (California Strawberry). Subspecies bracteata has a bract along the flowering stem that looks like one leaflet. Subspecies californica does not have this bract. The berries are prized by berry pickers today. Native Americans ate the berries fresh or mashed with other berries and dried. The runners were used as cordage (rope), and the leaves were dried and powdered as a disinfectant for open sores. Fragaria is Latin for fragrant. Vesca means little, slender or feeble, referring to the slender flowering stem. Bracteata is Latin for a thin, metal plate and refers to the bract on the flowering stem of this subspecies. Californica means of California and refers to the general range of this subspecies. 204 Plant Descriptions © Stanford University Press Serviceberry (Amelanchier alnifolnia) Photo by Beatrice F. Howitt, California Academy of Sciences Pale Serviceberry (Amelanchier pallida) Wood Strawberry (Fragaria vesca var. bracteata) 205 Plant Descriptions Bigleaf Avens Geum macrophyllum Willd. Also called Large-leaf Avens. Native. Rose Family (Rosaceae). Bigleaf Avens has yellow flowers with five petals surrounding many stamens with yellow anthers, and many pistils that are light green when immature and turning greenish-yellow when mature. The five small, green sepals are reflexed. When the fruits are mature, the flower looks like a spiny, red ball. The basal leaves have long stems and are compound with a large, kidney-shaped to heart-shaped terminal segment. The other leaflets are much smaller than the terminal leaflet. The alternate stem leaves are deeply lobed, and the three lobes are coarsely toothed. Lower surfaces of the leaves are often mottled reddish. At the base of the leaves, there are large, leaf-like stipules. The reddish-brown stems are hairy, and usually reach up to 60-90 cm. Bigleaf Avens blooms around mid-June in wet areas. Nearby, you may find Tall Phacelia (Phacelia procera) or Cow Parsnip (Heracleum lanatum). Geum is an ancient Latin name. In Greek, macrophyllum means large leaf, referring to the large terminal leaflet on the basal leaves. Henderson’s Horkelia Horkelia hendersonii Howell Native. Rose Family (Rosaceae). This low-lying, mat-forming plant has small, regular flowers with 5 whitish pink petals and 5 whitish pink sepals. It is a perennial that grows approximately 4-6 inches high. The alternate, pinnately compound leaves are silky with white hairs; each leaf has 11-19 leaflets. It occurs at elevations from 6,400 – 7,500 feet. Look for this rare plant around Rabbit Ears from early June through mid-July. This inconspicuous flower is a gem on Mt. Ashland. It is globally rare, but locally abundant, found only on the dry, granitic soils of four peaks in southern Oregon (Mount Ashland, McDonald Peak, Dutchman Peak, and Observation Peak) and in one small area in California (Dry Lake Lookout). From 1980 to 1996, Henderson’s Horkelia was a Category 2 candidate for listing under the U.S. Endangered Species Act. This category included species for which sufficient data was not available to confirm vulnerability. In 1996, this category of species was discontinued, and Henderson’s Horkelia was dropped. In September 1999, the U.S. Fish and Wildlife Service received a formal petition from the Rogue Group of the Sierra Club of Ashland to list Henderson’s Horkelia as endangered or threatened. In June 2000, the U.S. Fish and Wildlife Service announced that it will conduct a comprehensive review of Henderson’s Horkelia to determine if it should be protected under the ESA. Since 1995, the U.S. Forest Service has considered this plant a sensitive species, and the Oregon Natural Heritage Program considers it “Threatened Throughout Range.” The scientific name of this plant honors two individuals who have contributed to the study of botany. J. Horkel (1769-1846) was a German plant physiologist; Louis Henderson (1854-1942) was an American botanist who lived and botanized in the western United States. 206 Plant Descriptions © Stanford University Press Bigleaf Avens (Geum macrophyllum) Henderson’s Horkelia (Horkelia hendersonii) 207 Plant Descriptions Ocean Spray Holodiscus discolor (Pursh) Maxim. Also called Creambush Ocean Spray, Ironwood, Arrow Wood. Native. Rose Family (Rosaceae). Triangular sprays of white flowers make this shrub obvious when it blooms in July and early August. The small radial flowers have 5 petals and a bushy center with many pistils and stamens that are strongly exserted. The dry, withered flowers usually persist well into the following year. The simple, wedge-shaped leaves are alternate and toothed, and are borne on reddish-brown twigs that turn to gray with maturity. The twigs are short-hairy. Ocean Spray is found mostly around rocky outcrops and on rocky slopes, and is often surrounded by Greenleaf Manzanita (Arctostaphylos patula) and Cream Stonecrop (Sedum oregonense). A similar species, Rock Spiraea, Holodiscus microphyllus Rydb. var. microphyllus, is found in the rocks only at the summit of Mount Ashland. This species looks almost identical to Ocean Spray, but is characterized by very small, hairy leaves. The foliage is a food source for deer. Also called Ironwood by many tribesThe stems were used by Native Americans for making arrow shafts, bows, digging sticks, campfire tongs, spears, drum hoops, and canoe paddles. The bark and leaves were used to treat burns and sores. Infusions of seeds were used for prevention of contagious diseases. Holodiscus is Greek for whole disk. Discolor is derived from the Greek words for two colors, possible referring to the bark which changes from reddish-brown to gray. Sticky Cinquefoil Potentilla glandulosa Lindley Native. Rose Family (Rosaceae). Although the stem of Sticky Cinquefoil is always hairy, they are not always sticky. (Someone told me once that Mother Nature didn’t read the book.) The radial flowers vary from pale yellow to deep yellow and have 5 sepals, 5 petals, 20 stamens, and many pistils. The open cymes usually bear 2-8 flowers. The flowering stems are erect, but the compound basal leaves are spreading to erect. The basal leaves are pinnately compound with 7-11 toothed leaflets. The few stem leaves are also compound but look different from the basal leaves and have fewer leaflets. Three different subspecies have been recorded on Mount Ashland. I found ssp. glandulosa around the summit. The stem of this one is densely glandular-hairy, therefore, it is very sticky. This subspecies prefers open slopes. I recorded ssp. globosa Keck along County Road 1151 near the ski area parking lot. The stem is densely hairy, but completely non-glandular, and therefore, not sticky. The flowers of this subspecies do not fully open. Finally, ssp. ashlandica (E. Greene) Keck is usually found in moist places. The stem of this one is hairy, but not glandular. The common name, cinquefoil, means five leaves and refers to five leaflets that many of the plants in this genus possess. In Latin, potentilla means small but powerful, referring to the reputed medicinal value of some plants in this genus. Also from Latin, glandulosa means full of glands. Sulphur Cinquefoil Potentilla recta L. Non-native from Eurasia. Rose Family (Rosaceae). These very pale yellow radial flowers have 5 green sepals, 5 petals, 20 stamens, and many pistils. The palmately compound leaves have 6-7 leaflets that are toothed ½ way to the mid-vein. There are basal leaves and stem leaves, and the basal leaves are generally fallen or withered when the flowers bloom. The erect flowering stem is usually 10-30 cm tall. Sulphur Cinquefoil tends to grow in disturbed areas, so look for it along County Road 1151 at elevations below 6300 feet. It blooms in early to mid-August with Idaho Gum Weed (Grindelia nana) and Woolly Mullein (Verbascum thapsus). The common name, cinquefoil, means five leaves and refers to five leaflets that many of the plants in this genus possess. In Latin, potentilla means small but powerful, referring to the reputed medicinal value of some plants in this genus. Recta is Latin for straight or upright, possibly referring to the habit of this plant to grow erect. 208 Plant Descriptions Ocean Spray (Holodiscus discolor) Sticky Cinquefoil (Potentilla glandulosa) Sulphur Cinquefoil (Potentilla recta) 209 Plant Descriptions Bitter Cherry Prunus emarginata (Hook.) Walp. Also called Wild Cherry. Native. Rose Family (Rosaceae). This deciduous shrub has fairly small but showy flowers. Many stamens with white filaments and bright yellow anthers protrude from the center of the five pure white petals. The flowers are clustered along the stem in groups of 3 to 12. The elliptic leaves have many shallow teeth that can be either sharp or rounded. The bark is often reddish-brown with obvious horizontal rows of yellow lenticels. The oval to spherical fruits vary in color from red to purple, and are called drupe. The botanical definition of a drupe is a fleshy or pulpy fruit that superficially resembles a berry but has only one seed encased in a hardened stone. This shrub usually flowers in May and the fruits mature around August. It is abundant along USFS Road 20 between the picnic area and Rabbit Ears. The leaves, twigs, and pits of this plant contain amygdalin, a cyanogenetic glycoside. When cracked, the pits are extremely dangerous, but if intact, they are apparently harmless. The common name comes from the drupe, which is too bitter to eat. Prunus is derived from the Latin word pruninus which means plum blue, referring to the color of some of the fruits in this genus. Emarginata is derived from the Latin word for border or edge, referring to the finely toothed leaf margins. Wood Rose Rosa gymnocarpa Nutt. Also called Dwarf Rose, and Baldhip Rose. Native. Rose Family (Rosaceae). Wild roses are such a delight in the dark woods. The delicate, radial flowers have 5 green sepals with long, tapered tips, 5 pale pink to rosy pink petals that are white toward the center, and many pale yellow pistils and stamens. One to three flowers grow in clusters toward the ends of the upper branches. Very short, glandular hairs are on the pedicels and the outer surfaces of the sepals. The thin branches are covered with many slender, straight thorns. The alternate leaves are odd-pinnate with 3 to 9 leaflets with double-toothed, glandular margins. The stipules at the base of each petiole are also gland-margined. This deciduous shrub usually grows 30-90 cm tall. Wood Rose is found in wooded areas along the Pacific Crest Trail at elevations below 6400 feet. It blooms in early to mid-July with Spotted Coralroot (Corallorhiza maculata) and Sweet-scented Bedstraw (Gallium trillifolium). Although rose hips are considered edible and are a great source of vitamin C, the seeds of this species contain hairs that are irritating to the digestive tract, especially the very end of the digestive tract! Native peoples of the Pacific Northwest used Wood Rose as a medicine to cure several ailments, and as a ceremonial herb. One of the common names, Baldhip Rose, is descriptive of the fruit. The calyx drops from the fruit early, leaving the end of the hip exposed, or bald. Rosa is an ancient Latin name. Gymno is Greek for naked or bare, and carpa is Greek for fruit, again referring to the naked or bare fruit (hip) as a result of the calyx falling away. Rough Fruit Berry Rubus lasiococcus A. Gray Also called Dwarf Bramble. Native. Rose Family (Rosaceae). A prostrate, ground covering shrub, Rough Fruit Berry is a bramble with no prickles. One to two flowers grow in the leaf axils. Each radial flower has 5 green reflexed sepals, 5 white petals, many green pistils, and many stamens. The alternate leaves are palmately lobed, more or less compound, with 3 toothed leaflets. At the base of each petiole are two ovate stipules. The red raspberry-like fruits are covered with many minute hairs. Rough Fruit Berry is most often found in open forested areas. Rubus is an ancient Latin word for bramble. Lasiococcus is derived from the Greek words for hairy berry, referring to the dense tiny hairs on the fruits. 210 Plant Descriptions Bitter Cherry (Prunus emarginata) Wood Rose (Rosa gymnocarpa) Rough Fruit Berry (Rubus lasiococcus) 211 Plant Descriptions Black-cap Raspberry Rubus leucodermis Torrey & A. Gray Also called White-bark Raspberry, Black Raspberry, Western Raspberry, and Blackcap. Native. Rose Family (Rosaceae). An arched to erect shrub, Black-cap Raspberry has many prickles that are either straight or recurved. Few-flowered clusters are borne at the end of the upper branches. Each radial flower has 5 green sepals, 5 white petals, many pistils, and many stamens. The alternate compound leaves usually have 3 leaflets but can have as many as 7 leaflets. The leaflets are shallowly lobed, irregularly toothed, and white on the lower surface. Linear stipules are at the base of each petiole. The branches are round in cross section, and the fruits are red-purple to black with many tiny hairs. Look for this bramble in moist, rocky areas below 6200 feet. These berries can be used to make a purple stain, especially when combined with berries of other shrubs. Many Pacific Northwest native groups ate the berries. Rubus is an ancient Latin word for bramble. Leucodermis is derived from Greek and means white skin, referring to the white lower surface of the leaflets. Thimbleberry Rubus parviflorus Nutt. Native. Rose Family (Rosaceae). Thimbleberry is a tall, erect shrub at lower elevations, but on Mount Ashland, it is a low-growing, spreading shrub that reaches only 30-60 cm tall. The round branches have no prickles. Three to four white or pink flowers grow at the branch tips. Each flower has 5 green sepals, 5 white or pink petals, many pistils, and many stamens. The pedicels are densely glandular. Alternate, simple leaves are 5 lobed, finely toothed, and similar in shape to maple leaves. At the base of each leaf are two lanceolate stipules. The red fruits are raspberry-like and covered with hairs that can only be seen with a hand lens. Thimbleberry blooms in early to mid-July in wet, shaded areas along the Pacific Crest Trail at 6400 feet and lower. Look for Wiggin’s Lily (Lilium pardalinum ssp. wigginsii) and Douglas’s Knotweed (Polygonum douglasii) blooming nearby. Almost all coastal tribes in the Pacific Northwest ate the berries. Rubus is an ancient Latin word for bramble. In Latin, parviflorus means small flowered. As this shrub has some of the largest flowers in the Rubus genus, it is uncertain why this species was named parviflorus. Pennsylvania Blackberry Rubus pensilvanicus Poiret Non-native from eastern North America. Rose Family (Rosaceae). Several racemes bear 5-10 radial flowers. Each white flower has 5 sepals, 5 petals, many stamens, and many pistils. The compound leaves have 3-5 sharply toothed leaflets, and linear stipules at the base of the petiole. The 5-angled stems have many curved prickles with wide bases. The arched to erect bramble doesn’t grow very tall on Mount Ashland, only about 30-60 cm. The glabrous fruits are blackberry-like. Generally growing in disturbed areas, Pennsylvania Blackberry is found on Mount Ashland growing under the wooden fence along the ski area parking lot. Other non-natives grow nearby, including Virginia Pepperweed (Lepidium virginicum var. virginicum) and Shepherd’s Purse (Capsella bursapastoris). Rubus is an ancient Latin word for bramble. Pensilvanicus is New Latin meaning of Pennsylvania and refers to the native range of this species. 212 Plant Descriptions Photo by Charles Webber, California Academy of Sciences Black-cap Raspberry (Rubus leucodermis) © University of California Press Thimbleberry (Rubus parviflorus) Pennsylvania Blackberry (Rubus pensylvanicus) 213 Plant Descriptions Western Burnet Sanguisorba occidentalis Nutt. Native. Rose Family (Rosaceae). An innocuous plant with inconspicuous flowers, one has to look closely to find the beauty and mystery of Western Burnet. The erect, highly branched stems bear many terminal spikes with 10-50 tiny flowers per spike. Using your hand lens, you will discover that there are no petals on these greenish flowers, but there are 4 green sepals, 2 stamens, and 0-3 pistils. The alternate stem leaves, which I find sublimely beautiful, are pinnately compound with leaflets that are again pinnately lobed. The basal leaves are often withered by the time the flowers bloom. This annual or biennial grows 10-70 cm in disturbed areas. Look for Western Burnet along County Road 1151 up to the ski area parking lot. It blooms in mid- to late-July with Bird’s Foot Trefoil (Lotus purshianus var. purshianus) and Pearly Everlasting (Anaphalis margariticea). Latin for blood-absorbing, sanguisorba is so named for its tendency to halt bleeding by contracting the surrounding tissues or blood vessels. Occidentalis is Latin for western, referring to the range of this species. Cascade Mountain Ash Sorbus scopulina E. Greene var. cascadensis (G. Jones) C. Hitchc. Native. Rose Family (Rosaceae). A shrub with many flat-topped panicles of white flowers could be either Cascade Mountain Ash or Blue Elderberry. The flowers of Cascade Mountain Ash have five light green sepals and five white petals. Protruding from the center of the flower are many (15-20) stamens with reddish-brown anthers. The pinnately compound leaves are opposite with 9-11 leaflets that are toothed along the upper ½ to 2/3 of the margin. There are generally two green stipules at the base of each leaf. The bark is reddish-brown and ridged. I have seen this plant only on the south slope of the knoll at mile 8 along County Road 1151. It is growing along a rocky outcrop with Serviceberry (Amelanchier alnifolia). It flowers in late May or early June. The genus name, Sorbus, is an ancient Latin name. Scopulina Latin for rock or crag, referring to the rocky habitats where this species is found. Cascadensis is New Latin for of the Cascades, referring to the general range of this subspecies. Madder Family (Rubiaceae) Goosegrass Galium aparine L. Also called Bedstraw, Catchweed Bedstraw, Cleavers, Stickywilly, and Cheese Rennet Herb. Native. Madder Family (Rubiaceae). Children refer to this as nature’s velcro. The entire plant has small, hooked prickles that cause it to stick to clothing and animal fur. The flowers are so small and inconspicuous that you may not even notice them. If you look with your hand lens, however, you will see the whitish flowers in the axils. The flowers have four petals. The linear leaves are in whorls of 6-8; the lower leaves have petioles while the upper leaves are sessile. The stem is square. The fruits are nutlets and are also covered with many tiny hooks. This annual grows in grassy, shaded areas along the Pacific Crest trail and near wet meadows. As one of the common names suggests, this plant was used as a rennet to curdle milk for cheesemaking. It is said to impart a sweet flavor to cheese. True coffee is a member of this family, so it is not surprising that some sources claim that the seeds of this plant can be roasted and ground as a coffee substitute. Galium is Greek for Bedstraw. It originates from a word that means milk, referring to the use of some of the species in this genus in curdling milk. Aparine is the old, generic name for this species. 214 Plant Descriptions Western Burnet (Sanguisorba occidentalis) Cascade Mountain Ash (Sorbus scopulina var. cascadensis) Goosegrass (Galium aparine) 215 Plant Descriptions Low Mountain Bedstraw Galium bifolium S. Watson Also called Twinleaf Bedstraw. Native. Madder Family (Rubiaceae). This tiny plant grows at the base of California False Hellebore (Veratrum californicum) in wet meadows. You’ll probably have to get down on your belly to appreciate them. The tiny white flowers have three petals and bloom at the end of nodding, axillary pedicels. The linear leaves grow in whorls of 4 around the square stem. The plant is glabrous, but the fruits – nutlets – have short, hooked hairs that will adhere to clothing and animal fur. This delicate annual blooms in May and can be found in almost any wet meadow on Mount Ashland. It is often found growing next to Small-flower Nemophila (Nemophila parviflora) beneath the pleated leaves of California False Hellebore (Veratrum californicum var. californicum). Galium is Greek for Bedstraw. It originates from a word that means milk, referring to the use of some of the species in this genus in curdling milk. In Latin, bifolium means two leaves, referring to two sets of two leaves that grow from each node. Sweet-scented Bedstraw Galium triflorum Michaux Also called Fragrant Bedstraw. Native. Madder Family (Rubiaceae). The creamy white flowers of this perennial have four petals and are borne in clusters of 2-3 at the end of axillary stalks. The ovate to obovate leaves grow in whorls of 6 and are generally horizontal regardless of the stem’s orientation. The plant is mostly glabrous but may have some short hairs that can only be seen with a hand lens. The square stem can be either erect or decumbent, and grows to about 12 inches tall. Sweet-scented Bedstraw grows in the damp, shady forests along the Pacific Crest Trail. It blooms in July with Brewer’s Mitrewort (Mitella breweri) and Spotted Coralroot (Corallorhiza maculata). Galium is Greek for Bedstraw. It originates from a word that means milk, referring to the use of some of the species in this genus in curdling milk. Triflorum is Latin for three flowers, referring to this plant’s tendency to have a cluster of three flowers growing in the leaf axils. Milk Kelloggia Kelloggia galioides Torrey Native. Madder Family (Rubiaceae). The trumpet shaped corolla is made up of 4 or 5 petals that are mostly white and often tinged with pink. The flower buds are deep magenta. The flower seems to sit atop a short-bristly, ball-like calyx. The four stamens are exserted from the center of the flower. The opposite, entire leaves have tiny white stipules at the base. The slender stem is more or less four-angled, and grows erect, up to 20 cm tall. Milk Kelloggia prefers cool, partly shaded areas, along with White Hawkweed (Hieracium albiflorum) and Siskiyou Penstemon (Penstemon anguineus). Look for it in late June through July along wooded portions of the Pacific Crest Trail. There are only two species in this genus that honors Dr. Albert Kellogg (1813-1887), a pioneer California botanist. Galioides probably means galium-like, referring to the flowers which are similar to flowers in the genus, Galium. 216 Plant Descriptions Low Mountain Bedstraw (Galium bifolium) Sweet-scented Bedstraw (Galium triflorum) Milk Kelloggia (Kelloggia galioides) 217 Plant Descriptions Salicaceae (Willow Family) Sierra Willow (Salix eastwoodiae) A.A. Heller These familiar deciduous shrubs grow 2-7 feet tall on in the wet meadows on Mount Ashland. Like several other plants on the mountain, willows are dioecious, which means there are two kinds of flowers growing on the same shrub: flowers with only stamens, and flowers with only pistils. The dense, velvety catkins are packed with the tiny flowers, none of which have sepals or petals. The simple, alternate leaves vary in shape from linear to lanceolate to obovate. They can be entire or toothed, and are generally hairy. The flexible, glabrous twigs with smooth bark can be yellow, reddish, or brown. There is probably more than one species of willow on Mount Ashland. Willows have been vital to the lifestyle of many Native American groups. Willows have been used across the U.S. for utilitarian and medicinal purposes. The flexible twigs are perfect for constructing strong baskets, and their bitter bark contains salicylates, or salicin, which is used to make the acetylsalicylic acid found in aspirin. It is an ancient remedy for pain and inflammation. Willows are important on Mount Ashland for the animals. It is a choice browse food for grouse, deer and others. Salix is the ancient Latin name for these plants. Eastwoodiae honors Alice Eastwood (18591953) an American botanist raised in the western United States. Eastwood, who was listed in American Men of Science in 1906, authored three books on California flora, and one book on the flora of Denver, Colorado. Saxifrage Family (Saxifragaceae) Large-flowered Boykinia Boykinia major A. Gray Also called Mountain Boykinia, and Large Boykinia. Native. Saxifrage Family (Saxifragaceae). White, radial flowers with five petals help to distinguish this plant. Many flowers grow together in a flat-topped inflorescence at the top of the stem. The large basal leaves are maple-like with several lobes that have coarse teeth. Stem leaves are smaller than the basal leaves, but shaped similarly. Largeflowered Boykinia can grow up to 60 cm in moist areas. Look for this in the wet meadow above the road near the picnic area. It will be found growing near Tall Sneezeweed (Helenium bigelovii) and California False Hellebore (Veratrum californicum var. californicum). This genus was named after Dr. S. Boykin (1786-1848), a naturalist from Georgia. Major is Latin for larger, referring to the flowers which are large compared to other flowers in this genus. Small-flower Woodland Star Lithophragma parviflorum (Hook.) Torrey & A. Gray Native. Saxifrage Family (Saxifragaceae). There aren’t many of these flowers on Mount Ashland, so if you’ve found one, you must be very lucky, or a keen observer. One to six non-fragrant, radial flowers grow in a tight raceme. Each flower has 5 green sepals, 5 white petals with 3 lobes at the tip, 10 stamens, and 1 pistil. Alternate stem leaves and basal leaves are deeply 3 lobed to palmately compound. Each lobe or leaflet is lobed again. There are very few stem leaves. The erect stem of this perennial is usually 15-45 cm tall. Preferring open areas, Small-flower Woodland Star grows in open meadows. It blooms in midto late- June close to Oregon Sunshine (Eriophyllum lanatum) and Alice’s Daisy (Erigeron aliceae). Greek for rock hedge, lithophragma refers to the rocky habitats of many species in this genus. In Latin, parviflorum means small flowered. 218 © Stanford University Press Plant Descriptions Sierra Willow (Salix eastwoodiae) Photo by Bob Vos Large-flowered Boykinia (Boykinia major) Small-flower Woodland Star (Lithophragma parviflorum) 219 Plant Descriptions Brewer’s Mitrewort Mitella breweri A. Gray Native. Saxifrage Family (Saxifragaceae). Wow! Look at those amazing flowers! Use your hand lens, and get up close with this one. The 5 greenish-yellow petals are pinnately divided into thread-like lobes. The feather-like petals recurve away from the central disk to which 5 short, green sepals are attached. The 5 stamens are attached to the central disk and are visible between each petal. One pistil lies at the center. The tiny radial flowers are borne in a raceme atop a leafless stem. Brewer’s Mitrewort has only basal leaves that have 7-11 shallow lobes with rounded teeth. This perennial stands about 10-45 cm tall. Brewer’s Mitrewort is found on moist shady slopes along the Pacific Crest Trail below 6300 feet. Nearby, you may find Baneberry (Acteae rubra) or Starry False Solomon’s Seal (Smilacina stellata). Mitella is Latin and means a head-band or small cap. This refers to the fruit, which breaks open along a transverse line so that the top comes off like a lid. William Henry Brewer (1828-1910), an American botanist, was a pioneer botanical explorer in California, and California's first State Botanist. Five-Point Bishops Cap Mitella pentandra Hook. Also called Five-point Mitrewort, and Five-stamen Mitrewort. Native. Saxifrage Family (Saxifragaceae). Will ya look at that! You must use your hand lens to get a good look at these incredible flowers. The 5 green petals are pinnately dissected into 5-10 opposite, threadlike lobes. Unbelievable! The crazy petals recurve away from a central reddish-brown disk to which 5 short, green sepals are attached. Each small, radial flower has 5 stamens, one opposite each petal, and 1 pistil in the center. The terminal raceme rises above the basal leaves, which have 5-9 lobes with sharp teeth. The erect stem has many fine, white hairs. This perennial usually stands 15-30 cm tall. I found Five-point Bishops Cap on the north slopes, in a wooded area between the Windsor Chair Lift and Winter ski run. A small stream runs through the woods. Along the streambank, you should find Five-point Bishops Cap with Brook Saxifrage (Saxifraga odontoloma) and Hornemann’s Willow-herb (Epilobium hornemannii var. hornemannii). They all bloom in early to mid-July. The fruit and seeds of this little flower are interesting. The fruit breaks open along a transverse line so that the top comes off like a lid. Inside, there are many tiny, round, shiny black seeds. The seeds are then dispersed when animals or humans bump into the cups, spilling the seeds. Some sources call this type of seed dispersal a “splash cup dispersal” because rain drops that directly hit the seed cups eject the tiny, smooth seeds. Mitella is the Latin word for head-band or small cap. This refers to the manner in which the fruit breaks open, like someone lifting a cap from his head. Pentandra is Greek and literally means five men, referring to the five stamens. Oregon Saxifrage Saxifraga oregana J. Howell Native. Saxifrage Family (Saxifragaceae). A tall, sparsely woolly, leafless stem bears an elongated panicle of many small, white flowers. The radial flowers have 5 small green sepals, 5 small white petals, 10 stamens with yellow anthers that turn rusty-brown with maturity, and 1 pistil with 2 styles and a swollen, shiny green ovary. The basal leaves are somewhat fleshy, and vary from entire to toothed. The erect stem of this perennial usually stands 15-60 cm tall. You will find Oregon Saxifrage in wet meadows and boggy areas on the south slopes. It blooms in mid- to late-June and towers over the belly flowers, Primrose Monkey Flower (Mimulus primuloides) and Tinker’s Penny (Hypericum anagalloides). It also grows next to the large basal leaves and ripening fruits of White Marsh Marigold (Caltha leptosepala var. biflora). Saxifraga is Latin for rock-breaking, referring to the tendency of many flowers in this genus to grow on or near rocks, their roots eventually breaking the rocks apart. Oregana is New Latin for of Oregon, referring to the general range of this species. 220 Plant Descriptions Brewer’s Mitrewort (Mitella breweri) Five-Point Bishops Cap (Mitella pentandra) Oregon Saxifrage (Saxifraga oregana) 221 Plant Descriptions Brook Saxifrage Saxifraga odontoloma Piper Also called Stream Saxifrage. Native. Saxifrage Family (Saxifragaceae). Oh! Here’s another interesting flower growing right next to Five-point Bishops Cap (Mitella pentandra). Five white petals hide the five green, reflexed sepals. Each round or oval petal has two green spots toward the base. Ten stamens have club shaped filaments and anthers that look like small red balls, turning yellow with maturity. When the flower first opens, all the stamens lie flat atop the petals. Then 5 by 5, they stand erect. The erect purplish-red stem bears the panicle of flowers and is surrounded with basal leaves. The leaves are more or less round with coarse, sharp teeth and a kidney shaped base. This perennial usually grows 10-45 cm tall. I have seen Brook Saxifrage growing in two different areas, both off the beaten path. In some trees next to a stream between Windsor Chair Lift and Winter ski run, you will find it growing surrounded by Five-point Bishops Cap. The second area is found by parking your car at the confluence of USFS Road 20 and Road 40S30 and hiking in the opposite direction of the road. You will walk over a saddle and downhill into a hidden wet meadow in the Ashland watershed. Along the stream in the meadow, you will find it growing next to Hornemann’s Willow-herb (Epilobium hornemannii var. hornemannii). Saxifraga is Latin for rock-breaking, referring to the tendency of many flowers in this genus to grow on or near rocks, their roots eventually breaking the rocks apart. Derived from Greek, odontoloma means toothed fringe, and refers to the leaf margins. Lace Flower Tiarella trifoliata L. var. unifoliata (Hook.) Kurtz Also called One-leaf Foam Flower, False Mitrewort, Western Coolwort, and Single Sugar Scoop. Native. Saxifrage Family (Saxifragaceae). Tiny, delicate, white, radial flowers nod from an elongated raceme-like inflorescence. The 5 white sepals are much wider than the 5 white thread-like petals. Ten long, whisker-like stamens with yellow anthers are exserted beyond petals, and 1 pistil centers the flower. The inflorescence is covered with many white glandular hairs. The alternate leaves are mostly at the base of the stem. They are maple-like with 3-5 lobes; the lobes are toothed. This hairy perennial usually stands 20-40 cm. Moist, shady banks are perfect habitats for Lace Flower. The common name, Foam Flower, refers to the small flowers, which look like flecks of foam. Another common name, Single Sugar Scoop, refers to the seed capsules, which some people believe to resemble sugar scoops. Tiarella is Greek for small tiara, and refers to the beautiful little flowers. Trifoliata is Latin for three leaves, and refers to the compound leaf with three leaflets of the primary species. However, this subspecies has a simple maple-like leaf, hence, unifoliata which is Latin for one leaf. Figwort Family (Scrophulariaceae) Coast Indian Paintbrush Castilleja affinis Hook. & Arn. Native. Figwort Family (Scrophulariaceae). All Paintbrushes are very difficult to identify. The most colorful parts of these plants are the bracts, which are bright red to yellow in this species. Inside each bract, you will find a slender bilateral flower that has reddish or yellowish margins. This erect perennial grows 15-60 cm tall, and the stems below the flowers have alternate linear leaves. The stem is generally few-branched and yellow becoming purplish with age. Some of the leaves may have 2-5 lobes at the tip. Look for this one in dry places. The genus name comes from the 18th century Spanish botanist, Domingo Castillejo. Affinis is Latin for allied or related. It is uncertain to what this refers. 222 Plant Descriptions Photo by Charles Webber, California Academy of Sciences Brook Saxifrage (Saxifraga odontoloma) © Stanford University Press Photo by Br. Alfred Brousseau, St. Mary’s College Lace Flower (Tiarella trifoliata var. unifoliata) Coast Indian Paintbrush (Castilleja affinis) 223 Plant Descriptions Cobwebby Paintbrush Castilleja arachnoidea Greenman Also called Cotton Indian Paintbrush, and Yellow Paintbrush. Native. Figwort Family (Scrophulariaceae). These flowers are difficult to discern. What catches your attention first are the yellowish-green bracts that protect the flowers hidden beneath. The lower bracts usually turn pinkish to rusty red. If you can pull back the bracts, you will find long, bilateral flowers that are pale green to pale yellow. The entire inflorescence is wooly with white hairs. These erect perennials grow 10-15 cm tall, and the stems below the flowers have alternate leaves that are so deeply lobed that they may appear to be compound. There are 3 narrow leaflets on each leaf. Look for this flower around Rabbit Ears in late June and early July. It will be blooming with Sulfur Flower (Eriogonum umbellatum) and Henderson’s Horkelia (Horkelia hendersonii). The genus name comes from the 18th century Spanish botanist, Domingo Castillejo. In Greek, arachnoidea means spider web, which refers to the many white, cob-webby hairs on this plant. Giant Red Indian Paintbrush Castilleja miniata Hook. ssp. miniata Native. Figwort Family (Scrophulariaceae). This name is appropriate for the paintbrush that has the largest inflorescence on Mount Ashland. The flowers and bracts are bright red. Bracts may be entire, or can have three to five lobes. The elongated, bilateral flower is straight. The inflorescence is hairy, but the rest of the plant is glabrous. The stem is slightly ridged and can be simple or branched. Leaves are entire, and glabrous, except for the upper-most leaves just beneath the inflorescence, which are hairy. Giant Red Indian Paintbrush grows 20-45 cm. Look for this perennial in wet areas on the south slopes, growing next to Swamp White-heads (Sphenosciatum capitellatum) and Big-leaf Lupine (Lupinus polyphyllus var. pallidipes). They bloom through July and August. The genus name comes from the 18th century Spanish botanist, Domingo Castillejo. Miniata is Latin for red lead, perhaps referring to the red color of the bracts and flowers. Frosted Indian Paintbrush Castilleja pruinosa Fern. Native. Figwort Family (Scrophulariaceae). At first glance, you may confuse this plant with Giant Red Indian Paintbrush. The easiest way to tell these two apart is by looking at the stem. If the entire stem has short, white hairs, you are looking at Frosted Indian Paintbrush. If the stem (below the inflorescence) is glabrous, you probably have Giant Red Indian Paintbrush. Frosted Indian Paintbrush has red bracts and flowers, but the flowers are slightly smaller than those of Giant Red Indian Paintbrush. Bracts can vary from entire to lobed (three to five lobes). The alternate leaves can either be entire to lobed, with 2 or 3 lobes. Frosted Indian Paintbrush usually grows 20-30 cm. Look for this perennial in dry areas on the south slopes from June through August. Nearby, you may see Meadow Larkspur (Delphinium nuttallianum) and Torrey’s Blue-Eyed Mary (Collinsia torreyi var. latifolia). The genus name comes from the 18th century Spanish botanist, Domingo Castillejo. Pruinosa, in Latin, means full of hoar frost, indicating the frosted appearance of this plant that is the result of many white hairs. 224 Plant Descriptions Cobwebby Paintbrush (Castilleja arachnoidea) Giant Red Indian Paintbrush (Castilleja miniata var. miniata) Frosted Indian Paintbrush (Castilleja pruinosa) 225 Plant Descriptions Blue-eyed Mary Collinsia parviflora Lindley Also called Bluelips, Small Flowered Blue-eyed Mary, or Maiden Blue-Eyed Mary. Native. Figwort Family (Scrophulariaceae). These cheerful, bilateral flowers seem to laugh, perhaps because they could not decide whether they wanted to be white or purple. The upper lip is white and the lower lip is deep purple or blue. Most of the flowers grow in whorls above the leaves, but some of the flowers are in the upper leaf axils. The entire leaves are opposite. This plant usually grows from 15-20 cm tall. This annual blooms in late May and sets seed a few weeks later. The seeds lie dormant through the winter and germinate as soon as the snow melts the following spring. Look for Blue-Eyed Mary in shady places, including rocky outcrops that are shaded during the heat of the day. The genus name, Collinsia, honors the American botanist, Zacheus Collins (1764 – 1831). Parviflora means small flowered. Torrey’s Blue-Eyed Mary Collinsia torreyi A. Gray var. latifolia V. Newsom Also called Torrey’s Collinsia. Native. Figwort Family (Scrophulariaceae). This little annual is easily identified because of the bialteral, bicolored flowers: the upper lobes are white, and the lower lobes are bluish/lavender. The lower flowers are axillary and paired, while the upper flowers grow in open whorls, with about 3-5 flowers per whorl. The inflorescence is very sticky. The simple leaves are opposite, and shallowly/sparsely toothed. Some leaves may appear to be more or less entire. This is a diminutive plant, growing about 7-10 cm tall in sandy, granitic soil. It blooms from mid-June through mid-July. Look for Torrey’s Blue-Eyed Mary around Rabbit Ears, and along the road between Rabbit Ears and Grouse Gap shelter. This annual will be found growing along with Staining Collomia (Collomia tincturia) on loose, gravelly soils. Thomas Nuttall named this genus after the American botanist, Zacheus Collins (1764 – 1831). The specific epithet acknowledges Dr. John Torrey (1796-1879), a botanist of western North America. He reviewed and described the collections of many other noted botanists, including William Baldwin, and David Douglas. Dalmatian Toadflax Linaria genistifolia (L.) Miller ssp. dalmatica (L.) Maire & Petitm. Non-native from the Mediterranean. Figwort Family (Scrophulariaceae). Large, showy, bilateral flowers are tri-colored. The upper lip is pale yellow, the lower lip is a rich gold-yellow, and in the middle are orange hairs. The base of the lower lip forms a long spur. The 5 obvious sepals are glabrous, glaucous, long and pointed at the tip, with wavy margins. The alternate leaves are somewhat fleshy. They are entire with wavy margins, and the bases clasp the glabrous stem. The tall, branched stem is glabrous and glaucous. Dalmatian Toadflax is found at lower elevations along County Road 1151, blooming next to Bull Thistle (Cirsium vulgare) and Naked Buckwheat (Eriogonum nudum). This perennial blooms from late July through mid-August. Dalmatian Toadflax is considered a noxious weed in the western United States. The Greek word linon, and the Celtic word, lin, mean thread, and refer to the flax-like leaves of some of the plants in this genus. Genistifolia is Latin for leaves of broom plant. The reference for this is uncertain, but perhaps the leaves of Dalmation Toadflax are similar to the leaves of a broom plant. Dalmatica is the Latinized version of the city name of Dalmatia, Yugoslavia. The original name of this species was Linaria dalmatica, perhaps because it was first identified in Dalmatia, Yugoslavia. 226 Photo by Br. Alfred Brousseau, St. Mary’s College Plant Descriptions Blue-eyed Mary (Collinsia parviflora) Torrey’s Blue-eyed Mary (Collinsia torreyi var. latifolia) Dalmatian Toadflax (Linaria genistifolia ssp. dalmatica) 227 Plant Descriptions Seep Monkey Flower Mimulus guttatus DC. Also called Common Monkey Flower, Yellow Monkey Flower, and Seep-Spring Monkey Flower. Native. Figwort Family (Scrophulariaceae). Wet meadows brag of these wonderful yellow flowers. The bilateral flowers that grow in the axils of upper leaves have two lips. The upper lip has two lobes and the larger lower lip has three shallow lobes. On the lower lip, in the throat of the flower are red or maroon dots. The 5 sepals are fused into a ridged tube with hairs and red markings. Opposite simple leaves are sessile, and vary from toothed to entire. The pedicels and upper most leaves have fine, white hairs, but the remainder of the plant is glabrous. This species can be an annual or a perennial; it usually grows about 2-40 cm tall. As the name implies, you should look for Seep Monkey Flower next to running water in the wet meadows on the south slopes. It blooms from mid-June through mid-August with Musk Monkey Flower (Mimulus moschatus) and American Brooklime (Veronica americana). Mimulus is derived from Latin and means little mime or comic actor. This refers to the face-like corolla of some of the species in this genus. Guttatus is Latin for spotted, referring to the red dots on the petals. Musk Monkey Flower Mimulus moschatus Lindley Also called Musk Flower. Native. Figwort Family (Scrophulariaceae). Often growing right next to Seep Monkey Flower (Mimulus guttatus), you will find this close relative: another bright yellow bilateral flower with 5 petals. The flower appears almost radial, and there is one large red or purple splotch on the lower petal at the throat. The yellow petals rise out of the 5 sepals that are fused into a green tube. One of the most distinguishing characteristics of this species is the opposite, fuzzy leaves. Many white, fairly long hairs cover the calyx and the entire to shallowly toothed leaves. This perennial is often musk scented, and stands 5-30 cm tall. The stems are sometimes prostrate, sometimes ascending. Streambanks are the preferred habitat of Musk Monkey Flower. It blooms from early- to lateJuly next to White-flowered Bog Orchid (Platanthera leucostachys) and Arrowhead Butterweed (Senecio triangularis). Mimulus is derived from Latin and means little mime or comic actor. This refers to the face-like corolla of some of the species in this genus. Moschatus is Greek for musk, referring to the scent of this species. Dwarf Purple Monkey Flower Mimulus nanus Hook. & Arn. Also called Dwarf Monkey Flower. Native. Figwort Family (Scrophulariaceae). Gorgeous bilateral flowers have 5 green sepals that are fused at the base to form a tube. Five petals protrude from the calyx tube; the petals are also fused at the base forming a tube. Four stamens are hidden inside the corolla tube, and 1 pistil shows off an exserted stigma. The deep magenta/purple flowers have 2 gold stripes along the throat floor. The stripes are surrounded by and often dotted deeper magenta. The opposite, entire leaves are hairy, but you’ll probably need a hand lens to see the hairs. The lower surfaces of the leaves are purple, and on the upper surface, the margins are often colored purplered. This annual only grows 1-10 cm tall. Look for Dwarf Purple Monkey Flower in bare openings around shrubs or in sandy runoff areas above small streams. Mimulus is derived from Latin and means little mime or comic actor. This refers to the face-like corolla of some of the species in this genus. Nanus is Greek for dwarf, referring to the diminutive stature of this species. 228 Plant Descriptions Seep Monkey Flower (Mimulus guttatus) Photo by Bob Vos Musk Monkey Flower (Mimulus moschatus) Dwarf Purple Monkey Flower (Mimulus nanus) 229 Plant Descriptions Primrose Monkey Flower Mimulus primuloides Benth. ssp. primuloides Native. Figwort Family (Scrophulariaceae). This is one of my favorite flowers on Mount Ashland. It is unbelievable that something so tiny can be so perfect. One tiny little flower is borne at the end of a slender, thread-like, brownish-red stem. The bright yellow flower is bilateral, but looks almost radial. It blossoms from the fused, ridged calyx tube with brown-red markings on the ridges. Five yellow petals are notched at the tips, and two of the petals have one red dot each. The throat is open revealing 4 stamens and 1 pistil, and a few more very small red dots at the base of the petals. The entire basal leaves form a rosette around the flowering stem. The fuzzy leaves hold water droplets, making them appear shimmery silver. This little perennial grows no more than 4 cm tall. Primrose Monkey Flower is found in wet meadows and boggy areas. Look for spongy mats of moss; here you will find the little belly flower blooming in mid-July. Nearby, you will find Tinker’s Penny (Hypericum anagalloides) and Bolander’s Madia (Madia bolanderi). Mimulus is derived from Latin and means little mime or comic actor. This refers to the face-like corolla of some of the species in this genus. Primuloides is derives from the Latin word, primulinus, which means primrose-yellow, and the Greek suffix, -iod, which means like. This word refers to the yellow color of these flowers, which is similar to primroses. Woodland Penstemon Nothochelone nemorosus (Lindley) Straw Also called Woodland Beard-tongue. Native. Figwort Family (Scrophulariaceae). The common name, Woodland Penstemon, is a misnomer. Although in almost every way, this perennial looks like a Penstemon, it is not a Penstemon. The large (at least 1 inch long) flowers vary in color from white, to pink, to maroon, often two of these colors are mottled on one flower. The open, glandular panicle displays the bilateral flowers. Each flower has 5 sepals and 5 petals, both of which are fused at the base. Four stamens with obvious woolly anthers, 1 pistil, and 1 staminode are nestled inside the corolla throat. The large, opposite leaves are coarsely toothed. Erect stems stand 30-60 cm tall. Look for this unique flower on the north slopes, beyond the ski runs in rocky, open places in the forest. Woodland Penstemon blooms in early August. I found lots of it among Greenleaf Manzanita (Artcostaphylos patula), and on a rocky outcrop with One-sided Wintergreen (Orthilia secunda). Nothochelone is derived from Greek and means false turtle. This refers to the corolla, which is similar to that of Chelone, which means turtlehead. Chelone is another genus in the Figwort family. Derived from the Latin word for woodland, nemorus describes the general habitat of this species. Siskiyou Mountains Owl’s Clover Orthocarpus cuspidatus E. Greene ssp. cuspidatus Also called Broad-scaled Orthocarpus, and Siskiyou Mountains Orthocarpus. Native. Figwort Family (Scrophulariaceae). These beautiful plants have many bilateral flowers with a large, lower white balloon-like structure and a bright pink beak that protrudes above the white balloon. The flowers are protected by greenish-pink bracts. The branching stems can grow from 1-3 terminal spikes of flowers. The alternate leaves at first appear entire and grass-like, but upon closer examination you’ll see that these leaves are simple but deeply lobed with three pointed lobes. The erect reddish stems are sparsely hairy, and grow to heights of 10-20 cm. Look for these lovely annual flowers in dry soils around Rabbit Ears and along the Pacific Crest Trail from late June through July. It will be surrounded by Sulfur Flower (Eriogonum umbellatum) and Nettle-leaf Giant Hyssop (Agastache urticifolia). Derived from Greek, orthocarpus means straight fruit, referring to the fruit morphology. Cuspidatus is derived from a Latin word meaning a stiff point. This refers to the stiff leaves. 230 Plant Descriptions Primrose Monkey Flower (Mimulus primuloides ssp. primuloides) © University of California Press Woodland Penstemon (Nothochelone nemorosus) Siskiyou Mountains Owl’s Clover (Orthocarpus cuspidatus var. cuspidatus) 231 Plant Descriptions Mountain Owl’s Clover Orthocarpus imbricatus S. Watson Native. Figwort Family (Scrophulariaceae). Mountain Owl’s Clover is very similar to Siskiyou Mountains Owl’s Clover (Orthocarpus cuspidatus var. cuspidatus). The coloration and general characteristics of the two plants are the same. The difference is that the flowers of Mountain Owl’s Clover are much smaller, mostly hidden by the bracts, and do not have a large, white, balloon-like structure. Mountain Owl’s Clover prefers dry to moist soil. These flowers bloom from mid-July through August. Derived from Greek, orthocarpus means straight fruit, referring to the fruit morphology. Derived from Latin, imbricatus means a roof tile or a shingle, referring to the calyx lobes which are laterally imbricate when the fruits are ripe. Leafy Lousewort Pedicularis racemosa Hook. Also called Parrot’s Beak, Sickletop Lousewort, and Leafy Pedicularis. Native. Figwort Family (Scrophulariaceae). The bilateral flowers can be either white or pink and have an upper beak-like portion that looks a bit like a parrot’s beak or perhaps like an elephant trunk. The prominent lower petal is 3 lobed. The double-serrated leaves are alternate along the stems. Stems are often clustered, and if you look closely at the base of the stems, you will discover that they are slightly woody. Leafy Lousewort is usually 30-45 cm tall and grows in partly shaded, dry areas along the Pacific Crest Trail. It flowers in the first half of July. Growing nearby, you may find Silverleaf Phacelia (Phacelia hastata) or Milk Kelloggia (Kelloggia galioides). Pedicularis comes from the Latin word for lice. This name comes from an old belief that ingestion by stock promoted lice infestation. Racemosa is Latin for cluster, referring to the inflorescence. Siskiyou Penstemon Penstemon anguineus Eastw. Also called Siskiyou Beard-tongue. Native. Figwort Family (Scrophulariaceae). The tubular blue-purple flowers of Siskiyou Penstemon are made up of five petals that are fused at the base and free around the mouth. There are two lobes (or petals) on top and three lobes (or petals) on the bottom. The lower portion of the flower had three deep grooves that are highlighted in white. There are some hairs around the mouth of the flowers. The flowers occur in pairs or whorls along the upper portion of the stem with leaf-like bracts below each whorl or pair of flowers. The inflorescence (including the upper portion of the stem, the calyx and outer corolla) is sticky with many short hairs, while the lower portion of the stem is glabrous. The opposite leaves are more or less toothed, and the upper leaves sometimes clasp the stem, but are not fused around the stem. The green stem turns redbrown with maturity, and grows to heights of 30-60 cm. The stems are usually ascending to decumbent. Look for this one in shaded to partly shaded areas along the Pacific Crest Trail. Western Wallflower (Erysimum capitatum) and Leafy Lousewort (Pedicularis racemosa) may be growing nearby. In both Latin and Greek, penstemon means almost thread, or almost stamen, referring to the staminode, which is a sterile, stamen-like structure. Anguineus is derived from the Latin word for snake. It is uncertain to what this refers. 232 Plant Descriptions Photo by Bob Vos Mountain Owl’s Clover (Orthocarpus imbricatus) © University of California Press Leafy Lousewort (Pedicularis racemosa) Sisikiyou Penstemon (Penstemon anguineus) 233 Plant Descriptions Davidson’s Penstemon Penstemon davidsonii E. Greene var. davidsonii Native. Figwort Family (Scrophulariaceae). One to several large flowers (at least 1 inch long) bloom in a glandular raceme at the top of short, leafy stems. The pinkish-purple to deep blue, bilateral flowers have 5 fused sepals that are red-brown with short sticky hairs, 5 fused petals, 4 stamens with woolly anthers, 1 staminode, and 1 pistil. The corolla floor is white, and shaggy-hairy. The leaves are opposite, and entire to shallowly and irregularly toothed. The upper portion of the stem is red-brown, and lower portion of the stem is woody. This plant has a mat-like habit, growing no more than 10 cm tall. Davidson’s Penstemon can be found on the summit and on rocky outcrops around Rabbit Ears. It blooms in June with Cut-leaf Daisy (Erigeron compositus) and Cream Stonecrop (Sedum oregonense). In both Latin and Greek, penstemon means almost thread, or almost stamen, referring to the staminode, which is a sterile, stamen-like structure. This species is named after Dr. George Davidson (1825-1911), a geographer and astronomer who collected plants in Yosemite and throughout California. Azure Penstemon Penstemon parvulus (A. Gray) Krautter Native. Figwort Family (Scrophulariaceae). This erect to decumbent perennial has fairly large flowers that are bright, deep purple and blue. The tube (fused portion of the petals) is usually purple-pink, while the lobes are usually sky blue. The long, open raceme bears 4 – 24 flowers in pairs or whorls. Five sepals are fused at the base, 5 petals are also fused at the base, and 4 stamens, 1 staminode and 1 pistil are tucked inside the corolla throat. The stamen filaments are usually curved along the sides of the corolla throat so the anthers hug the top. The staminode lies on the corolla floor. The opposite, glabrous leaves have entire margins that are often wavy margins. The leaf base clasps the stem, which is glabrous, glaucous, and brownish-red. These plants usually grow in clumps, and are 15-20 cm tall. Azure Penstemon grows in dry, rocky soils along USFS Road 20. It blooms from late June through late July with Frosted Indian Paintbrush (Castilleja pruinosa), Palmer’s Catchfly (Silene bernardina), or Scarlet Gilia (Ipomopsis aggregata). In both Latin and Greek, penstemon means almost thread, or almost stamen, referring to the staminode, which is a sterile, stamen-like structure. Parvulus is derived from Latin and means very small or insignificant; it is uncertain to what this refers. Woolly Mullein Verbascum thapsus L. Also called Common Mullein. Non-native from Eurasia. Figwort Family (Scrophulariaceae). For a tactile delight, rub these woolly hairy leaves against your cheek. They’re just wonderful! No wonder there are stories of cowboys using these leaves for toilet paper. These are better than Charmin! The dense, spike-like raceme bears many bright yellow flowers that are more or less radial. Five sepals are hidden by the 5 petals, but the 5 stamens and 1 pistil are visible in the center. The basal leaves and alternate stem leaves are entire and densely woolly. The stout, simple stem grows 30-200 cm tall. Woolly Mullein, like many non-natives, grows in disturbed areas. Look for it along County Road 1151 at lower elevations, and around the ski lodge. This biennial blooms from late July through August, with Silverleaf Phacelia (Phacelia hastata) or Idaho Gum Weed (Grindelia nana). Verbascum is a Latin word that means Mullein. It comes from a Latin root word that means bearded, referring to the woolly leaves. Thapsus used to be the name of a peninsula on the island of Sicily that is now called Magnisi, probably where this species was first found. 234 Plant Descriptions © University of California Press Davidson’s Penstemon (Penstemon davidsonii var. davidsonii) © University of California Press Photo by Br. Alfred Brousseau, St. Mary’s College Azure Penstemon (Penstemon parvulus) Woolly Mullein (Verbascum thapsis) 235 Plant Descriptions American Brooklime Veronica americana (Raf.) Benth. Also called Brooklime, and American Speedwell. Native. Figwort Family (Scrophulariaceae). These clear blue flowers are very delicate, and the entire corolla drops at the slightest touch. Technically bilateral, but appearing radial, these flowers have 4 free sepals that are hidden by 4 petals. The petals are violet-blue with magenta or deep blue lines and bright magenta spots near the center of the flower. Two stamens and 1 pistil are exserted and, therefore, obvious in the center. One plant bears many flowers in several axillary racemes; each raceme has 10-25 flowers. The opposite leaves are entire to shallowly toothed. This perennial usually stands 10-25 cm. You will find American Brooklime in almost every wet meadow on the south slopes of Mount Ashland. It blooms from late June through mid-July amidst Common Chickweed (Stellaria media), Goosegrass (Galium aparine), or White-flowered Bog Orchid (Platanthera leucostachys). “God speed you well,” is the old English meaning of the common name Speedwell. The name, Brooklime, comes from “brook” because this flower is found in wet areas, and the verb “to lime,” a practice of trapping birds with a sticky substance. This was often done near water, so the little blue flower became associated with wet places and trapped birds. The genus name, Veronica, is from Saint Veronica, and through her, this flower is credited with miraculous healing powers, even the dread scrofula, or King’s Evil. Such powers are attributed to other species in this family, hence the family name, Scrophulariaceae. Americana means of America, referring to the range of this species. Common Speedwell Veronica arvensis L. Also called Corn Speedwell. Non-native from Eurasia. Figwort Family (Scrophulariaceae). This very small annual, no more than 5-10 cm tall, is usually hidden among other plants. If you find it, you will see a terminal raceme of small flowers, each flower growing in the upper leaf axils. Four petals that are fused at the base hide 4 free sepals. The petals are white to blue to violet with dark blue or violet lines. Two stamens and 1 pistil are obvious in the center. The opposite leaves are crenate to serrate. Common Speedwell generally grows in meadows, gardens, and lawns. I found this one blooming along USFS Road 20 in early June, hidden among some rushes. “God speed you well,” is the old English meaning of the common name. It is also called Corn Speedwell because it often grows in gardens. It is believed that the genus name comes from Saint Veronica. Arvensis is derived from the Latin word for field, referring to the meadow habitats of this species. Thyme-leaf Speedwell Veronica serpyllifolia L. Native. Figwort Family (Scrophulariaceae). A narrow inflorescence bears small flowers in the upper leaf axils. These flowers have 4 sepals, 4 petals, 2 exserted stamens, and 1 pistil. The upper petal is bright blue with cobalt blue lines, the two side petals are pale blue with cobalt blue lines, and the lower petal is white. The center of the corolla is yellow. The opposite leaves are entire to crenate. This perennial usually stands 5-15 cm tall. Look for Thyme-leaf Speedwell along the Pacific Crest Trail near wet meadows. It blooms in late June and early July with Hornemann’s Willow-herb (Epilobium hornemannii var. hornemannii) and Tiny Trumpet (Collomia linearis). “God speed you well,” is the old English meaning of the common name. The genus name, Veronica, probably comes from Saint Veronica. Serpyllifolia is Latin for thyme-shaped leaf. The leaves must have reminded someone of Thyme (Thymus serpyllum). 236 Plant Descriptions Photo by Bob Vos American Brooklime (Veronica americana) Photo by William R. Hewlett, California Academy of Sciences Common Speedwell (Veronica arvensis) Thyme-leaf Speedwell (Veronica serpyllifolia) 237 Plant Descriptions Nettle Family (Urticaceae) Stinging Nettle Urtica dioica L. ssp. holosericea (Nutt.) Thorne Also called Hoary Nettle. Native. Nettle Family (Urticaceae) Tiny greenish, unisexual flowers grow in drooping panicles in the leaf axils of this erect perennial. The staminate flowers have 4 sepals, 0 petals, and 4 stamens, while the pistillate flowers have 4 stamens, 0 petals, and 1 pistil. Both male and female flowers are borne on the same plant, making Stinging Nettle a monoecious plant. The inconspicuous flowers indicate that this plant is wind pollinated. The opposite leaves are narrowly lanceolate, and coarsely toothed. On Mount Ashland, Stinging Nettle grows 15- 90 cm tall. Look for this one in moist waste places and streambanks on the north slopes in the Ashland Watershed. It will be tangled in with Giant Red Indian Paintbrush (Castilleja miniata var. miniata) and willows (Salix sp.). Most people notice this plant only after brushing against it. Feel that burning sensation? Many of the hairs on this plant contain an acid that causes intense itching after contact. In spite of our aversion to Stinging Nettle, it is a very important medicinal plant with many uses, from curing tuberculosis, to beating dandruff. It is also a valuable food plant, and I can attest to the fact that nettle leaf soup is quite tasty. The acid in the hairs is destroyed when the leaves are boiled. The leaves are high in vitamins A and C, and protein. This plant was also an important source of fiber for making strong, fine rope. Urtica comes from the Latin word to burn, referring to the stinging hairs. Dioica is Greek for two houses, referring to one of the subspecies that is dioecious – staminate flowers on one plant and pistillate flowers on a separate plant. Holosericea means entirely silky to the touch, perhaps referring to the fact that not all the hairs on this plant contain the stinging acid. Valerian Family (Valerianaceae) California Valerian Valeriana californica A.A. Heller Native. Valerian Family (Valerianaceae). Cream white, funnel-shaped flowers are borne in terminal and axillary cymes. Each flower has 515 calyx lobes that are coiled inward, and become beautifully plumose in fruit. Five petals, 3 stamens, 1 stigma form a more or less radial flower. There are only a few stem leaves on California Valerian. They are opposite, sessile and lobed; the 3-9 lobes have margins that are entire or toothed. The basal leaves can vary from simple, to deeply 3 lobed to compound. The stems and leaves can be glabrous to short-hairy. This erect perennial grows 25-50 cm tall in moist places and coniferous forests. Look for it around the wet meadows on the north and south slopes. It blooms from mid-June through early July, and may grow right next to it’s very close relative, Sitka Valerian (Valeriana sitchensis), or California False Hellebore (Veratrum californicum). It is uncertain where this genus name comes from. In Latin, it means strength possibly referring to its use in folk medicine. Or it could be named after Valerian, a Roman emperor. Californica means of California, referring to the general range of this species. 238 Plant Descriptions © University of California Press Stinging Nettle (Urtica dioica ssp. holosericea) California Valerian (Valeriana californica) 239 Plant Descriptions Sitka Valerian Valeriana sitchensis Bong. Also called Mountain Valerian, and Northern Valerian. Native. Valerian Family (Valerianaceae). White to pink, flowers are made up of 5 petals, 5-15 sepals, 5 stamens and 1 pistil that are all strongly exserted. The flowers appear to be more or less radial, but they are actually bilateral. The sepals aren’t very noticeable until the plant is in fruit. At that time, you can easily see that they are coiled inward and they become beautifully feathery. The stem leaves are opposite and compound with 3-5 leaflets that are irregularly crenate to toothed. However, the uppermost leaves are often simple and entire. The swollen nodes have short, white hairs. The stem and leaves are generally glabrous. Sitka Valerian often stands 20-60 cm tall. This perennial blooms from mid-June through early July in wet meadows and shaded areas. Nearby, you may find Tall Phacelia (Phacelia procera) or Heart-leaf Spring Beauty (Claytonia cordifolia). It is uncertain where the genus name, Valerian, comes from. In Latin, it means strength possibly referring to its use in folk medicine. Or it could be named after Valerian, a Roman emperor. The derivation and meaning of the word sitchensis is unclear. Violet Family (Violaceae) Stream Violet Viola glabella Nutt. Also called Pioneer Violet, Smooth Violet, Yellow Violet, Smooth Yellow Violet, and Johnny Jump-Up. Native. Violet Family (Violaceae). Early in the subalpine season, these yellow bilateral flowers brighten the wet meadows on Mount Ashland. Five yellow petals with purple lines cover the 5 green sepals. Five stamens and 1 pistil are hidden in the throat of the flower. Basal and stem leaves are simple, heart-shaped, and crenate to toothed. Two alternate stem leaves only grow near the tip of the stem, just beneath the flower. The tallest violet on Mount Ashland, Stream Violet is usually 6-18 cm tall. The stem and leaves are mostly glabrous. The violet has purple nectar guides on the petals direct pollinators to the nectary of the flower. While probing for nectar, the bees bump the pollen-laden anthers. Hairs on the side petals and style comb the pollen on the bees’ bodies from the previous flower. The seeds that result from this grand pollination scheme are often dispersed by ants that feed on the seed appendages. Stream Violet blooms in May and early June with White Marsh Marigold (Caltha leptosepala var. biflora) and Plantain-leaf Buttercup (Ranunculus alismifolius var. alismifolius) Violet flowers can be eaten raw, and the leaves can be cooked and eaten. Viola is the ancient Latin name for these flowers. Glabella means smooth, referring to the hairless leaves of this species. Small White Violet Viola macloskeyi F. Lloyd Also called Macloskey’s Violet. Native. Violet Family (Violaceae). This small, low-lying violet has a white, bilateral flower. The five petals are white, and the lower petal has several purple veins leading to the center of the flower. The two lateral petals can be glabrous or hairy (bearded). The white stigma and four golden-yellow anthers peep from the center of the flower. The basal leaves vary from being ovate with a heart-shaped base or round and reniform (kidney-shaped). They are glabrous and can be entire to shallowly crenate. There are usually no stem leaves, but if present, they will be very small and on the lower half of the stem. Small White Violet grows no higher than 5-6 cm tall. Look for this small violet close to streams and wet places in May and early June. It will often grow near Alpine Shooting Stars (Dodecatheon alpinum). Viola is the ancient Latin name for these flowers. Someone by the name of Macloskey is honored by this specific epithet. 240 © University of California Press Plant Descriptions Sitka Valerian (Valeriana sitchensis) Stream Violet (Viola glabella) Small White Violet (Viola macloskeyi) 241 Plant Descriptions Mountain Violet Viola purpurea Kellogg var. purpurea Also called Goosefoot Violet. Native. Violet Family (Violaceae). Bilateral, deep golden-yellow flowers have purple lines on all petals. Five green sepals, 5 petals, 5 stamens, and 1 pistil make up these early bloomers. The basal leaves and alternate stem leaves are simple, and nearly triangular – shaped somewhat like a goose’s foot. The shiny green leaves are often tinted with purple, and glabrous above while hairy below. The margins vary from entire to toothed. Mountain Violet grows in small clumps, no more than 5-6 cm tall. Look for this perennial in dry forested areas through May. You might find Shelton’s Violet (Viola sheltonii) and Greenleaf Manzanita (Arctostaphylos patula) blooming nearby. Although many uses of violets can be found in the literature, I think the most interesting use was noted by Pliny, the Roman of long ago, who recommended a garland of violets as a cure for a hangover headache. The leaves of violets are high in Vitamin C, which might be why they have been used to treat a variety of illnesses since medieval times. Viola is the ancient Latin name for these flowers. Purpurea is derived from the Latin word for purple, referring to the purple tint on the leaves. Shelton’s Violet Viola sheltonii Torrey Native. Violet Family (Violaceae). This violet is unique because it has two types of flowers: those that open and are pollinated by insects, and those that never open and are self-pollinated. These tiny, self-pollinating flowers that never open are called cleistogamous. The flowers that open have 5 bright yellow petals with purple veins. The petals cover the 5 small, green sepals. Five stamens and 1 pistil remain hidden in the corolla throat. The basal leaves and alternate stem leaves are highly dissected with linear segments. These flowers grow in clumps, and stand 5-10 cm tall. Shelton’s Violet can be found in gravelly soil in open forested areas at 6200 feet and lower. Look for them bloming in the middle of and along the sides of the old logging roads. You may also see Mountain Violet (Viola purpurea var. purpurea) and Greenleaf Manzanita (Arctostaphylos patula) blooming nearby. Viola is the ancient Latin name for these flowers. The specific epithet possibly honors Christopher A. Shelton, an American botanist who is noted for introducing the honeybee in California in 1837. 242 Plant Descriptions Photo by Bob Vos Mountain Violet (Viola purpurea var. purpurea) Shelton’s Violet (Viola sheltonii) 243 Plant Descriptions 244 Appendix A - Glossary APPENDIX A GLOSSARY Accessory a fleshy fruit consisting of many achenes on the external surface of a swollen receptacle; Wood Strawberry (Fragaria vesca) has this type of fruit Achene a dry fruit containing 1 seed; common in the Sunflower Family (Asteraceae) Aggregate a fleshy fruit consisting of many small drupes; Thimbleberry (Rubus parviflorus) has this type of fruit Alternate arranged singly along the stem Annual a plant that completes its life cycle in one growing season, usually one year Anther pollen-baring portion of the stamen; atop the filament; often described as "canoeshaped" Appressed pressed tightly upward; often pressed against the stem or leaf surface Ascending curving or angling upward Axil the angle or area between the main stem and the leaf stem Basal found at the base of the stem; usually in reference to leaves Beak an elongated structure on a flower or fruit; as in Siskiyou Mountains Owl's Clover (Orthocarpus cuspidatus var. cuspidatus) Berry a fleshy fruit with several to many seeds scattered throughout; Western Blueberry (Vaccinium uliginosum) has this type of fruit Biennial a plant that completes its life cycle in two growing seasons, usually two years Bilabial a bilateral flower with an upper and a lower lip Bilateral divisible into two halves that are mirror images of each other; asymmetrical; irregular; usually in reference to flowers Binomial the two word scientific name for a species, consisting of the genus name and specific epithet Blade the flat, extended part of the leaf 245 Appendix A - Glossary Bloom (pronounced bio me) a white powdery substance on the surface of the plant that easily rubs off Bract a small, leaf-like structure, usually growing at the base of a flower or flower cluster Calyx collective term for all of the sepals; the outer-most circle of flower parts Capsule a dry fruit that encloses many small seeds; breaks open to release the seeds; many orchids have this type of fruit Cathartic purging; usually referring to a medicinal plant that stimulates the bowels Cauline found on the stem Chaff dry, papery bracts on the receptacle in the Sunflower Family (Asteraceae) Chlorophyll green pigment in plant cells; essential to photosynthesis Clasping the base partially surrounds the stem; usually in reference to leaves Compound composed of two or more parts; usually in reference to leaves; compound leaves consist of several to many leaflets Composite a flowering head that is composed of many small, individual flowers; in reference to flowering heads in the Sunflower Family (Asteraceae) Coniferous cone-bearing; generally referring to evergreen, gymnospermous trees, including pine, fir, cedar, etc. Cordate heart-shaped; usually in reference to leaves Corolla collective term for all of the petals; whorl of flower parts immediately inside or above calyx Corymb a flat-topped inflorescence in which the outer flower stalks are long and the center flower stalks are short; similar in appearance to a candelabra; Yarrow (Achillea millefolium) has a corymb of heads Cyme a more or less flat-topped inflorescence in which flowers are oppositely branched Deciduous shedding leaves annually; Mountain Maple (Acer glabrum) is a deciduous shrub Decumbent lying flat on the ground with tips curving upward Disk Flower in the Sunflower Family (Asteraceae), a radial flower with a 4 or 5 lobed corolla 246 Appendix A - Glossary Drupe a fleshy fruit containing one seed that has a very hard seed cover; Bitter Cherry (Prunus emarginata) has this type of fruit Elliptic a leaf or leaflet that has the general outline of an ellipse Emetic causing vomiting Endemic native to a particular country, nation, or region Entire with the margin unbroken by teeth or lobes; in reference to leaves or leaflets Erect standing straight without support Evergreen having leaves that are green all year Exserted protruding; thrusting out; usually in reference to stamens or pistil Filament the stalk of the stamen that bears the anther at the top; usually thread-like Follicle a dry fruit that splits along one suture; species in the Stonecrop Family (Crassulaceae) have this type of fruit Glabrous without hairs Glandular having glands that exude a sticky substance; as in Bolander's Madia (Madia bolanderi) Glaucous covered with a white or blue powdery film that is easily rubbed off Head dense inflorescence of sessile flowers; refers to clovers and all members of the Sunflower family (Asteraceae) Herbaceous lacking wood Hip a false fruit; a fleshy container holding many achenes; species in the Rose Family (Rosaceae) have this type of "fruit" Imperfect a unisexual flower Inflorescence the flowering part of the plant; the cluster of flowers Irregular divisible into two halves that are mirror images of each other; asymmetrical; bilateral; usually in reference to flowers 247 Appendix A - Glossary Lanceolate broad at the base and tapering to a point at the tip; usually in reference to leaf shape Leaflet one of the separate leaf-like parts of a compound leaf Legume a dry fruit that splits along two sutures to release the seeds; a true pod; species in the Legume Family (Fabaceae) have this type of fruit Ligulate in Sunflower Family (Asteraceae), a bilateral flower with a 5-lobed strap-shaped structure that is often mistaken for one petal Ligule in Sunflower Family (Asteraceae), the strap-shaped structure on a ligulate flower Linear long and narrow with nearly parallel sides; usually in reference to leaf shape Lobed an expansion or rounded bulge of a leaf margin Node the point on a stem from which a leaf grows Nutlet a small, dry, nut-like fruit; species in the Borage Family (Boraginaceae) have this type of fruit Oblanceolate narrow at the base, broad and rounded at the tip; usually in reference to leaf shape Opposite arranged in pairs on the stem; generally in reference to leaves Organic derived from living organisms; often referring to the upper layer of soil that is made up of decaying plant matter Ovary the base of the pistil that contains ovules/seeds; this becomes the fruit as seeds mature Ovule the structure containing the egg; will become the seed after fertilization Palmate radiating from a common point; usually in reference to compound leaves Panicle a branched inflorescence in which the lower or outer flowers bloom before the upper or inner flowers; a compound raceme Pedicel stem of an individual flower or fruit Perennial a plant that completes its life cycle in three or more growing seasons, usually three or more years 248 Appendix A - Glossary Perfoliate leaf tissue completely surrounds the stem; stem appears to pierce the leaf Petal a segment of the corolla; often brightly colored Perfect a flower that has pistil(s) and stamen(s); containing male and female reproductive organs Perianth the calyx and corolla collectively Petiole stem of an individual leaf Pinnate having two rows of structures on opposite sides of an axis; feather-like; usually in reference to compound leaves Pistil the female productive organ of the flower; consists of ovary, stigma and style Pistillate a flower that lacks stamens Phyllaries In Sunflower Family (Asteraceae), bracts that subtend the flowering head; often mistaken for sepals Pollen the fine, dust-like grains produced by the anther and containing the male sexual cells Pome a fleshy fruit containing seeds in united compartments; apples and pears have this type of fruit Prostrate lying completely flat on the ground Raceme un-branched inflorescence of stalked flowers along the central stem; flowers usually bloom from bottom to top Radial branching out in all directions from a central point; divisible into many parts all of which look similar; regular; usually in reference to the flower Radiate in Sunflower Family (Asteraceae), a flowering head made up of central disk flowers surrounded by ray flowers; often mistaken for one radial flower Ray Flowers in Sunflower Family (Asteraceae), similar to ligulate flowers, but the ligule has only 3 lobes instead of 5 Receptacle the structure to which flower parts are attached; the "base" or "foundation" of a flower Recurved gradually curved outward 249 Appendix A - Glossary Reflexed abruptly bent outward or downward Regular branching out in all directions from a central point; divisible into many parts all of which look similar; radial; usually in reference to the flower Reniform kidney-shaped; usually in reference to leaf shape Rhizome underground, horizontal stem; often mistaken for a root Sagittate arrow-shaped with two lobes at the base; usually in reference to leaf shape Saprophyte an organism that obtains its nutrients from dead or decaying organic matter Scale a small, membranous structure; often similar to reduced leaves Schizocarp a dry fruit that splits into two or more one-seeded containers; Richardson's Geranium (Geranium richardsonii) has this type of fruit Sclerophyllous stiff and usually waxy; generally in reference to leaves Sepal a segment of the calyx; usually green Sessile without a stalk, pedicel, petiole or peduncle; directly attached to the stem or flower receptacle Sheathing surrounding or partially surrounding a structure; usually in reference to a leaf base surrounding or partly surrounding the stem Shrub a woody plant much branched from the base; usually no taller than 5 feet Silicle a short, broad silique; Shepherd's Purse (Capsella bursa-pastoris) has this type of fruit Silique a dry fruit; a long, narrow seed container made up of two valves that are divided by a thin membrane; species in the Mustard Family (Brassicaceae) have this type of fruit Simple leaf an undivided leaf composed of one single part Specific epithet Spike 250 the second half of the scientific name of a species an elongated inflorescence of sessile flowers along the main stem; flowers usually bloom from bottom to top Appendix A - Glossary Spine a plant structure, often a stipule or leaf, that is modified into a sharp-pointed projection Spur a hollow, tubular or conic project of a petal or sepal; as in Mountain Larkspur {Delphinium glaucum) Stamen the male reproductive organ of a flower; consists of filament and anther Staminate a flower with no pistils Stigma the tip of the pistil that receives the pollen; usually sticky Stipule a usually green, usually leaf-like structure at the base of the leaf stem; usually paired Style the tube that directs the pollen from the stigma to the ovary Subtend to grow beneath Taproot a main, tapered root that grows straight down into the soil Tendril a slender, coiling extension of a stem used for support or climbing; a modified stem Tepal a structure that could be a petal or a sepal; usually found in the Lily family on flowers in which both petals and sepals look identical or nearly the same Terminal at the end Ternate made up of three parts; usually in reference to a compound leaf Toothed having several to many sharp indentations along the margin of a leaf Tuber short, fleshy underground stem that stores nutrients for the plant Umbel an inflorescence in which all flower stalks radiate from the same point; similar to an upturned umbrella Vein transport tissue within a leaf; one of a network of tiny channels that transport nutrients and sugars within the leaf Vine a trailing or climbing plant; often using tendrils for support Whorl group of three or more leaves arranged in a circle around the stem 251 Appendix B - Illustrated Glossary APPENDIX B ILLUSTRATED GLOSSARY Pistil Stamen Basic Flower Parts / Bilateral Flowers Radial Flowers Umbel Raceme Paired in the axils Corymb Head Panicle Inflorescences 252 Appendix B - Illustrated Glossary Beak Special Flower Parts Blade Axil Petiole Stem Stipule Basic Leaf Parts Basal Opposite Alternate Whorled Leaf Arrangements Entire Toothed Lobed Wavy Leaf Margins 253 Appendix B - Illustrated Glossary Palmately compound leaf with entire leaflets Pinnately compound leaf with toothed leaflets Compound Leaves Sagittate Leaf Shapes 254 Appendix C - Bibliography APPENDIX C BIBLIOGRAPHY These sources were quoted directly or indirectly in this field guide: Bernstein, Art. 1986. Trees of Southern Oregon. Grants Pass, Oregon: New Leaf Books. Billings, W.D. 1974. Arctic and Alpine Vegetation: Plant Adaptations to cold summer climates. Arctic and Alpine Enviroments, ed. London: Ives, J.D. and Barry, R.G. William Clowes and Sons Limited. Billings, W.D. 1988. Alpine Vegetation. North American Terrestrial Vegetation, Second Edition. Barbour, M.G. and W.D. Billings, ed. New York: Cambridge University Press, pp. 537-570. Blunt, W. 1971. The Complete Naturalist: A life of Linnaeus. Borror, Donald J. 1988. Dictionary of Word Roots and Combining Forms. Mountain View, California: Mayfield Publishing Company. Chipman, A. 1970. Wildflower Trails of the Pacific Northwest. Medford, Oregon: Pine Cone Publishers. Coon, Nelson. 1974. The Dictionary of Useful Plants. Emmaus, Pennsylvania: A Rodale Press. Dennis, La Rea J. 1959. A Taxonomic Study of the Vascular Flora on Ashland Peak, Jackson County, Oregon. Thesis submitted to Oregon State College. Dennis, La Rea J. 1972. Name Your Poison: A Guide to Cultivated and Native Oregon Plants Toxic to Humans. Published by the author. EisleyL. 1957. The Immense Journey. New York: Vintage Books. 210 p. Elliott, Douglas B. 1976. Roots: An Underground Botany and Forager's Guide. Old Greenwich, Connecticut: The Chatham Press. G, Chuck. 1996. Dictionary of Botanical Epithets. http://www.winternet.com/~chuckg/dictionary.html Hickman, J.C. (ed.) 1993. The Jepson Manual: Higher Plants of California. Berkeley: University of California Press. Horn, Elixabeth L. 1994. Coastal Wildflowers of the Pacific Northwest: Wildflowers and 255 Appendix C - Bibliography Flowering Shrubs from British Columbia to Northern California. Missoula, Montana: Mountain Press Publishing Company. Houk, R. 1987. Wildflowers of the American West. San Francisco: Chronicle Books, pp. 47,57,62,63,71,75,93 Lang, Frank. 1991. Nature Notes, 1989-1991, Volume I. Ashland, Oregon: Southern Oregon State College Library. Lang, Frank. 1993. Nature Notes, 1991-1993, Volume II. Ashland, Oregon: Southern Oregon State College Library. Lang, Frank. 2000. A Nature Notes Sampler. Ashland, Oregon: Jefferson Public Radio, p. 4344. Mason, F. (editor). 1935. The Great Design: Order and Intelligence in Nature. New York: MacMillan Company, pp. 161-188. McKelvey, Susan Delano. 1956. Botanical Exploration of the Trans-Mississippi West, 17901850. Corvalis, Oregon: Oregon State University Press. Munz, P. A. 1959. A California Flora. Los Angeles: University of California Press. Papanicolaou K. and Kokkini S. 1982. Adaptive Plant Evolution in the Alpine Environment of the Greek Mountains. Being Alive on Land: Proceedings of the International Symposium on Adaptations to the Terrestrial Environment Held in Halkidiki, Greece, 1982. Margaris, N.S., M. Arianoustou-Faraggitaki, and W.C. Oechel ed. Boston: Dr. W. Junk Publishers, pp. 21-31 Peck, M.E. 1961. A Manual of the Higher Plants of Oregon. Portland, Oregon: Binfords and Mort. Petrides, G.A. and O. Petrides. 1992. Peterson Field Guides: Western Trees. New York: Houghton Mifflin Company, p. 43. Radford, Albert E„ William C. Dickison, Jimmy R. Massey, and C. Ritchie Bell. 1974. Vascular Plant Systematics. New York: Harper and Row Publishers. Robinson, P. 1977. Profiles of Northwest Plants: Food Uses, Medicinal Uses, Legends. Portland, Oregon: Green World Press, pp. 48, 114 Robinson, A.F., Jr. 2000. Federal Register Document for Horkelia hendersonii and Lupinus aridus ashlandensis. Federal Register. 65 (114): 37108-37111. Ross, Michael Elsohn. 1997. Flower Watching with Alice Eastwood. Minneapolis: Carolrhoda Books, Inc. 256 Appendix C - Bibliography Seda, A. 1989. Key Species for Plant Associations on the Rogue River, Siskiyou and Umpqua National Forests. USDA Forest Service. Schaffer, Jeffrey P. and Andy Selters. 1990. The Pacific Crest Trail Volume 2: Oregon and Washington. Berkeley, California: Wilderness Press. Ski Ashland. 2000. http://www.mtashland.com. Stearn, William T. 1966. Botanical Latin: History, Grammar, Syntax, Terminology and Vocabulary. Great Britain: David and Charles. 566 p. Sweetser, A.R. and M.E. Kent. 1908. Key and Flora: Some of the Common Flowers of Oregon. Chicago, Illinois: Ginn and Company. USFS, Ashland Ranger District. 2001. Final Environmental Impact Statement: Ashland Watershed Protection Project, pp. 1-6, III-1 to III-9. USFS Rogue River National Forest. 1991 .Final Environmental Impact Statement: Mount Ashland Ski Area. Jackson County, Oregon: USDA, Forest Service, Pacific Northwest Region. 243 p. Wiley, Leonard. 1968. Rare Wild Flowers of North America. Portland, Oregon: Published by the author. Ulrich, Larry. 1999. Wildflowers of the Pacific Northwest. Santa Barbara, California: Companion Press. United States Department of Agriculture (USDA) Natural Resources Conservation Service (NRCS). PLANTS Database. 2001. http://plants.usda.gov. University of California, Berkeley. Digital Library Project: CalPhotos. http://elib.cs.berkeley.edu/photos/ [WRCC] Western Regional Climate Center. 1997. website: www.wrcc.dri.edu. Line drawings are from... Abrams L. 1940. An Illustrated Flora of the Pacific States: Washington, Oregon and California. Volume 1, Ophioglossaceae to Aristolochiaceae. Stanford University, California: Stanford University Press. 538 p. Abrams L. 1944. An Illustrated Flora of the Pacific States: Washington, Oregon and California. Volume 2, Polygonaceae to Krameriaceae. Stanford University, California: Stanford University Press. 635 p. 257 Appendix C - Bibliography Abrams L. 1951. An Illustrated Flora of the Pacific States: Washington, Oregon and California. Volume 3, Geraniaceae to Scrophulariaceae. Stanford University, California: Stanford University Press. 866 p. Abrams L, Ferris RS. 1960. An Illustrated Flora of the Pacific States: Washington, Oregon and California. Volume 4, Bignoniaceae to Compositae. Stanford University, California: Stanford University Press. 732 p. Hickman JC editor. 1993. The Jepson Manual: Higher Plants of California. Berkeley: University of California Press. 1400 p. Hitchcock CL, Cronquist A, Ownbey M, Thompson JW. 1973. Vascular Plants of the Pacific Northwest. Seattle: University of Washington Press. Part 1, Vascular Cryptogams, Gymnosperms, and Monocotyledons; 914 p. Hitchcock CL, Cronquist A, Ownbey M, Thompson JW. 1973. Vascular Plants of the Pacific Northwest. Seattle: University of Washington Press. Part 2, Salicaceae to Saxifragaceae; 597 p. Hitchcock CL, Cronquist A, Ownbey M, Thompson JW. 1973. Vascular Plants of the Pacific Northwest. Seattle: University of Washington Press. Part 3, Saxifragaceae to Ericaceae; 614 p. Hitchcock CL, Cronquist A, Ownbey M, Thompson JW. 1973. Vascular Plants of the Pacific Northwest. Seattle: University of Washington Press. Part 4, Ericaceae to Campanulaceae; 510 PHitchcock CL, Cronquist A, Ownbey M, Thompson JW. 1973. Vascular Plants of the Pacific Northwest. Seattle: University of Washington Press. Part 5, Compositae; 343 p. Munz PA. 1961. California Spring Wildflowers. Berkeley: University of California Press. 122 p. Quotes were found in the following references: Kramer, Jack. 1996. Women of Flowers: A Tribute to Victorian Women Illustrators. New York: Stewart, Tabori & Chang. Terrill, Steve. 1995. Wildflowers of Oregon. Englewood, Colorado: Westcliffe Publishers, Inc. If you would like to study the wildflowers of southern Oregon in greater depth, try one of the following technical identification manuals. 258 Appendix C - Bibliography Hickman, J.C. (ed.) 1993. The Jepson Manual: Higher Plants of California. Berkeley: University of California Press. Hitchcock, C.L., A. Cronquist, M. Ownbey, and J.W. Thompson. 1973. Vascular Plants of the Pacific Northwest. 5 Parts. Seattle: University of Washington Press. Munz, P. A. 1959. A California Flora. Los Angeles: University of California Press. Peck, M.E. 1961. A Manual of the Higher Plants of Oregon. Portland, Oregon: Binfords and Mort Publishers. To learn more about botanical Latin, try these: Borror, Donald J. 1988. Dictionary of Word Roots and Combining Forms. Mountain View, California: Mayfield Publishing Company. Stearn, William T. 1966. Botanical Latin: History, Grammar, Syntax, Terminology and Vocabulary. Great Britain: David and Charles. 566 p. http ://www. calflora. net/botanicalnames 259 Index - Common Dames Agoseris Large-flowered, 64 Orange, 62 Pale, 66 Tall, 62 Anemone Drummond's, 194 Lyall's, 196 Angelica,52 Arnica, Heartleaf, 66 Aster Leafy, 68 Shasta Tansy, 80 Avens, Bigleaf, 206 Baneberry, 192 Barberry Family, 42, 90, 91 Bedstraw Low Mountain, 216 Sweet Scented, 216 Bell flower Family, 43, 100-103 Pale, 102 Birthwort Family, 41, 60, 61 Bistort, Western, 182 Blackberry, Pennsylvania, 212 Blazing Star, Bushy, 154 Blueberry, Western, 122 Blue-eyed Grass, 140 Blue-Eyed Mary, 226 Blue-Eyed Mary, Torrey's, 226 Borage Family, 42, 92-95 Boxwood, Oregon, 112 Boykinia, Large-flowered, 218 Bride's Bonnet, 144 Bristleweed, Whitney's, 78 Brooklime, American, 236 Broomrape Clustered, 170 Family, 47, 170-173 Naked, 172 Buckthorn Family, 49,202-203 Buckwheat Family, 48, 180-187 Jayne's Canyon, 180 Naked, 180* Burnet, Western, 214 Buttercup Family, 49, 190-203 Plantain-leaf, 200 Popular, 200 Butterweed, Arrowhead 88 Western, 86 Candy Flower, 188 Carrot Family, 41, 52-59 Cat's Ear, Hairy, 80 Cacthfly, Palmer's, 108 Checkerbloom, Oregon, 156 Cherry, Bitter, 210 Chickweed, Common, 110 Chicory, 70 Chinquapin 260 Bush, 128 Golden, 128 Clover Long-Stalk, 126 White 128 Collomia Large-flowered, 174 Staining, 176 Columbine, Western, 196 Coneflower, Western, 84 Coral root Spotted, 162 Striped, 164 Western, 164 Cinquefoil Sticky, 208 Sulphur 208 Cryptantha Pinewoods, 92 Torrey's, 92 Weak-stem, 92 Cudweed, Wright's, 76 Currant Red Flowering, 132 Sticky, 132 Swamp, 130 Wax, 130 Daisy Alice's, 72 Cut-leaf; 74 Dandelion, 90 Dandelion, Alpine Lake Prairie, 84 Delphinium, Anderson's, 198 Desert Parsley, Piper's, 54 Dock, Willow, 186 Dusty Maidens, 68 Elderberry Blue, 104 Red, 106 Evening Primrose Fam, 47,156-161 Faiiybells, Hooker's, 144 False Hellebore California, 152 Green, 152 Siskiyou, 152 Figwort Family, 50, 222-237 Fireweed, 156 Five-point Bishop's Cap, 220 Flax Common Blue, 154 Family, 46, 154-155 Fleabane Scabland, 74 Subalpine, 74 Fruit Berry, Rough, 210 Geranium Family, 45, 130-131 Richardson's, 130 Gilia, Scarlet. 176 Ginger, Wild, 60 Goldenbush, Greene's, 72 Goldenrod, Canada, 88 Gooseberry Family, 45, 130-133 Gummy, 132 Goosegrass, 214 Grape, Oregon, 90 Ground Cone, California, 170 Gum Weed, Idaho, 76 Harebell, California, 100 Hawksbeard, Naked-stem, 72 Hawkweed Scouler's, 80 White 78 Heath Family, 44, 114-123 Hedge Nettle, Rough, 142 Honeysuckle Family, 43, 102-107 Orange, 104 Wedded, 104 Huckleberry, Thinleaf, 120 Hyssop, Nettle-leaf Giant, 140 Indian Potato, California, 56 Ipomopsis, Ballhead, 178 Iris Family, 45, 140-141 Yellow-flowered, 140 Jacob's Ladder, Showy, 180 Kelloggia, Milk, 216 Klamath Weed, 138 Knotweed Davis's, 182 Douglas's, 184 Lace Flower, 222 Ladies' Tresses, Hooded, 168 Larkspur Meadow, 200 Mountain, 198 Yellowtinge 198 Laurel, Bog, 118 Legume Family, 44, 122-129 Lily Chocolate, 146 Family, 46, 142-153 Glacier, 146 Washington, 148 Wiggin's, 146 Loasa Family, 46, 154-155 Lomatium, California, 54 Louse wort, Leafy, 232 Lovage, Gray's, 54 Lupine Big-leaf, 126 Brewer's, 124 Mount Ashland, 124 Sickle-keel Lupine, 124 Madder Family, 49, 214-217 Madia, Bolander's, 82 Mahal a Mat, 202 Mallow Family, 46, 156-157 Manzanita Greenleaf, 116 Pinemat, 114 Maple Index - Common flames Family, 41,52, 53 Mountain, 52 Marigold, White Marsh, 196 Meadow Rue, Fendler's, 202 Miner's Lettuce, 188 Mint Coyote, 142 Family, 46, 140-143 Mitrewort, Brewer's, 220 Monkey Flower Dwarf Purple, 228 Musk, 228 Primrose, 230 Seep, 228 Monkshood, Columbian, 190 Mountain Ash, Cascade, 214 Mugwort, 66 Mullein, Woolly, 234 Mustard Family, 43, 94-101 Nemophila, Small-flower, 134 Nettle Family, 50, 238-239 Oak Family, 44, 128-129 Ocean Spray, 208 Onion, Pacific, 142 Orchid Calypso, 162 Family, 47, 162-169 Green-flowered Bog, 166 Phantom, 162 Sparse-flowered Bog, 168 White-flowered Bog, 168 Oregon Sunshine, 76 Owl's Clover Mountain, 232 Siskiyou Mtns, 230 Oxalis Family, 47, 172-173 Paintbrush Coast Indian, 222 Cobwebby, 224 Frosted Indian, 224 Giant Red Indian, 224 Parsley Family, 41, 52-59 Parsnip, Cow, 52 Pasque Flower, White 194 Pea Family. 44, 122-129 Leafy, 122 Pearly Everlasting, 64 Penstemon Azure, 234 Davidson's, 234 Siskiyou. 232 Woodland 230 Pepperweed, Virginia, 100 Phacelia Changeable, 136 Silverleaf. 134 Tall, 136 Vari-Leaf. 136 Phlox Family, 48, 174-181 Slender. 78 Spreading, 178 Pineapple Weed, 70 Pinedrops, Woodland, 120 Pinesap, 118 Pink Family, 43, 108-111 Pipsissewa, 116 Plantain English, 174 Family, 48, 174-175 Rattlesnake, 166 Poppy Family, 47, 172-173 Primrose Family, 48, 190-191 Prince's Pine, Little, 116 Purslane Family, 48, 186-189 Pussy Ears, 144 Pussy Paws, 186 Pussytoes, Rosy. 64 Raillardella, Silky, 86 Raspberry, Black-cap, 212 Rockcress Holboell's. 96 Lyall's, 96 Pioneer, 96 Spreading Pod, 94 Tower, 94 Rose Family, 49, 204-215 Wood, 210 Sagebrush, Mountain, 68 Salsify, Yellow, 88 Sand Spurry, Red, 110 Sandwort, Ballhead 108 Large-leaf, 108 Sanicle, Northern, 58 Sawwort. American, 84 Saxifrage Brook, 222 Family, 50, 218-223 Oregon. 220 Serviceberry, 204 Shepherd's Purse, 98 Shooting Star, Alpine, 190 Sneezeweed, Tall, 78 Snowberry, 106 Snowberry, Creeping, 106 Solomon's Seal False, 148 Starry False, 150 Sorrel, Sheep, 184 Speedwell Common, 236 Thyme-leaf, 236 Spreading Ground Smoke, 160 Spring Beauty Heart-leaf, 186 Red-stem, 188 St. John's Wort Family, 45, 138-139 Staff-Tree Family, 43, 112-113 Starflower, Western, 190 Starwort, Curled, 110 Steer's Head, 172 Stickseed, Jessica's, 94 Stinging Nettle, 238 Stonecrop Cream, 114 Family, 44, 112-115 Spear-leaf, 112 Worm-leaf, 114 Strawberry, Wood, 204 Sulfur Flower, 182 Sunflower Family, 42, 60-91 Swamp White Heads, 58 Sweet Cicely, Western, 56 Sweet Root, Western, 56 Tansy Mustard, Mountain, 98 Tarweed Mountain, 82 Slender, 82 Thimbleberry, 212 Thistle, Bull 70 Tinker's Penny, 138 Tiny Trumpet, 174 Toadflax, Dalmatian, 226 Tobacco Brush, 202 Tofieldia, Western, 150 Toothwort, Nuttall's, 98 Trail Plant, 62 Trefoil, Bird's Foot, 122 Trillium, Western, 150 Twayblade, Broad-leaved 166 Twinflower, 102 Valerian California, 238 Family, 50, 238-241 Sitka, 240 Violet Family, 50, 240-243 Macloskey's, 240 Mountain, 242 Shelton's, 242 Stream, 240 Wallflower, Western, 100 Waterleaf Family, 45, 134-137 White, 134 Willow, 218 Willow Family, 49, 218-219 Willow-herb Fringed, 158 Hornemann's, 158 Milkflower, 160 Oregon, 160 Tall Annual, 158 Windflower, Columbia, 192 Wintergreen Family, 44, 114-123 One-sided, 118 White-veined, 120 Wood Sorrel, Three-leafed, 172 Woodland Star, Small-flower, 218 Yampah, 58 Yarrow, 60 261 Index - Scientific flames Acer glabrum, 52 Aceraceae, 41,52-53 Achillea millefolium, 60 Aconitum columbianum, 190 Acteae rubra, 192 Adenocaulon bicolor, 62 Agastache urticifolia, 140 Agoseris aurantiaca, 62 elata, 62 glauca, 66 grandiflora, 64 Allium validum, 142 Amelanchier alnifolia, 204 Anaphalis margariticea, 64 Anemone delto idea, 192 drummondii, 194 lyallii, 196 occidentalis, 194 Angelica arguta, 52 Antennaria rosea, 64 Apiaceae, 41, 51-59 Aquilegia formosa, 196 Arabis xdivaricarpa, 94 glabra, 94 holboellii, 96 lyallii, 96 platysperma, 96 Arctostaphylos nevadensis, 114 patula, 116 Arenaria congesta, 108 Aristolochiaceae, 41, 60-61 Arnica cordifolia, 66 Artemisia douglasiana, 66 tridentata, 68 Asarum caudatum, 60 Aster foliaceus, 68 Asteraceae, 42, 60-91 Berberidaceae, 42, 90-91 Berberis nervosa, 90 Boraginaceae, 42, 92-95 Boschniaka strobilacea, 170 Boykinia major, 218 Brassicaceae, 43, 94-101 Calochortus tolmiei, 144 Caltha leptosepala var. biflora, 196 Calypso bulbosa, 162 Calyptridium umbellatum, 186 Campanula prenanthoides, 100 scouleri, 102 Campanulaceae, 43, 100-103 Caprifoliaceae, 43, 102-107 Capsella bursa-pastoris, 98 Cardamine nuttallii, 98 Caryophyllaceae, 43, 108-111 Castilleja affinis, 222 262 arachnoidea, 224 miniata, 224 pruinosa, 224 Ceanothus prostratus, 202 velutinus, 202 Celastraceae, 43, 112-113 Cephalanthera austinae, 162 Chaenactis douglasii, 68 Chamomilla suaveolens, 70 Chimaphila menziesii, 116 umbellata, 116 Chrysolepis chrysophyllus, 128 sempervirens, 128 Cichorium intybus, 70 Cirsium vulgare, 70 Claytonia cordifolia, 186 perfoliata, 188 rubra, 188 sibirica, 188 Clintonia uniflora, 144 Collinsia parviflora, 226 torreyi, 226 Collomia grandiflora, 174 linearis, 174 tinctoria, 176 Corallorhiza maculata. 162 mertensiana, 164 striata, 164 Crassulaceae, 44, 112-115 Crepis pleurocarpa, 72 Cryptantha flaccida, 92 simulans, 92 torreyana, 92 Delphinium andersonii, 198 decorum, 198 glaucum, 198 nuttalianum, 200 Descurainea incisa, 98 Dicentra uniflora, 172 Disporum hookeri, 144 Dodecatheon alpinum, 190 Epilobium angustifolium, 156 brachycarpum, 158 ciliatum, 158 hornemannii, 158 lactiflorum, 160 oregonense, 160 Ericaceae, 44, 114-123 Ericameria greenei, 72 Erigeron alicea, 72 bloomeri, 74 compositus, 74 peregrinus, 74 Eriogonum diclinum, 180 nudum, 180 umbellatum, 182 Eriophyllum lanatum, 76 Erysimum capitatum, 100 Erythronium grandiflorum, 146 Fabaceae, 44, 122-129 Fagaceae, 44, 128-129 Fragaria vesca, 204 Fritillaria atropurpurea, 146 Galium aparine, 214 bifolium, 216 triflorum, 216 Gayophytum diffusum, 160 Geraniaceae, 45, 103-131 Geranium richardsonii, 130 Geum macrophyllum, 206 Gnaphalium canescens, 76 Goodyera oblongifolia, 166 Grindelia nana, 76 Grossulariaceae, 45, 130-133 Hackelia micrantha, 94 Hazardia whitneyi, 78 Helenium bigelovii, 78 Heracleum lanatum, 52 Hieracium albiflorum, 78 scouleri, 80 Holodiscus discolor, 208 Hydrophyllaceae, 45, 134-137 Hydrophyllum fendleri, 134 Hypericaceae, 45, 138-139 Hypericum anagalloides, 138 perforatum, 138 Hypochaeris radicata, 80 Ipomopsis aggregata, 176 congesta, 178 Iridaceae, 45, 140-141 Iris chrysophylla, 140 Kalmia polifolia, 118 Kelloggia galioides, 216 Lamiaceae, 46, 140-143 Lathyrus polyphyllus, 122 Lepidium virginicum, 100 Ligusticum grayi, 54 Liliaceae, 46, 142-153 Lilium pardalinum, 146 washingtonianum, 148 Linaceae, 46, 154-155 Linaria genistifolia, 226 Linnaea borealis, 102 Linum lewisii, 154 Listera convallarioides, 166 Lithophragma parviflorum, 218 Loasaceae, 46, 154-155 Index - Scientific names Lomatium califomicum, 54 piperi, 54 Lonicera ciliosa, 104 conjugialis, 104 Lotus purshianus, 122 Lupinus albicaulis, 124 aridus var. ashlandensis, 124 breweri, 124 polyphyllus, 126 Machaeranthera canescens, 80 Madia bolanderi, 82 glomerata, 82 gracilis, 82 Mahonia nervosa, 90 Maianthemum racemosum, 148 stellatum, 150 Malvaceae, 46, 156-157 Mentzelia dispersa, 154 Mimulus guttatus, 228 moschatus, 228 nanus, 228 primuloides, 230 Mitella breweri, 220 pentandra, 220 Moehringia macrophylla, 108 Monardella odoratissima, 142 Monotropa hypopytis, 118 Nemophila parviflora, 134 Nothocalais alpestris, 84 Nothochelone nemorosus, 230 Onagraceae, 47, 156-161 Orchidaceae, 47, 162-169 Orobanchaceae, 47, 170-173 Orobanche fasciculata, 170 uniflora, 172 Orogenia fusiformis, 56 Orthilia secunda, 118 Orthocarpus cuspidatus, 230 imbricatus, 232 Osmorhiza chilensis, 56 occidentalis, 56 Oxalidaceae, 47, 172-173 Oxalis trilliifolia, 172 Papaveraceae, 47. 172-173 Paxistima myrsiniles, 112 Pedicularis racemosa, 232 Penstemon anguineus, 232 davidsonii, 234 parvulus, 234 Perideridia oregana, 58 Phacelia hastata, 134 heterophylla, 136 mutabilis, 136 procera, 136 Phlox diffusa, 178 gracilis, 178 Plantaginaceae, 48, 174-175 Plantago lanceolata, 174 Platanthera hyperborea, 166 leucostachys, 168 sparsiflora, 168 Polemoniaceae, 48, 174-181 Polemonium califomicum, 180 Polygonaceae, 48, 180-187 Polygonum bistortoides, 182 davisiae, 182 douglasii, 184 Portulacaceae, 48, 186-189 Potentilla glandulosa, 208 recta, 208 Primulaceae, 48. 190-191 Prunus emarginata, 210 Pterospora andromedea, 120 Pyrola picta. 120 Raillardella argentea, 86 Ranunculaceae, 49, 190-203 Ranunculus alismifolius, 200 populago, 200 Rhamnaceae, 49, 202-203 Ribes cereum, 130 lacustre, 130 lobbii, 132 sanguineum, 132 viscosissimum, 132 Rosa gymnocarpa, 210 Rosaceae, 49, 204-215 Rubiaceae, 49, 214-217 Rubus lasiococcus, 210 leucodermis, 212 parviflorus, 212 pensilvanicus, 212 Rudbeckia occidentalis, 84 Rumex acetosella, 184 salicifolius, 186 Salicaceae, 49, 218-219 Salix eastwoodiae, 218 Sambucus mexicana, 104 racemosa, 106 Sanguisorba occidentalis, 214 Sanicula graveolens, 58 Saussurea Americana, 84 Saxifraga odontoloma, 222 oregana. 220 Saxifragaceae, 50, 218-223 Scrophulariaceae, 50, 222-237 Sedum lanceolatum. 112 oregonense, 114 stenopetalum, 114 Senecio integerrimus, 86 triangularis, 88 Sidalcea oregana, 156 Silene bernardina, 108 Sisyrinchium bellum, 140 Smilacina racemosa, 148 stellata, 150 Solidago canadensis, 88 Sorbus scopulina, 214 Spergularia rubra, 110 Sphenosciadium capitellatum, 58 Spiranthes romanzoffiana, 168 Stachys ajugoides var. rigida, 142 Stellaria crispa, 110 media, 110 Symphoricarpos albus, 106 mollis, 106 Taraxacum officinale, 90 Thalictrum fendleri, 202 Tiarella trifoliata v. unifoliata, 222 Tofieldia occidentalis, 150 Tragopogon dubius, 88 Trientalis latifolia, 190 Trifolium longipes, 126 repens, 128 Trillium ovatum, 150 Urtica dioica, 238 LJrticaceae, 50, 238-239 Vaccinium membranaceum, 120 uliginosum, 122 Valeriana californica, 238 sitchensis, 240 Valerianaceae, 50, 238-241 Veratrum califomicum, 152 insolitum, 152 viride, 152 Verbascum thapsus, 234 Veronica americana, 236 arvensis, 236 serpyllifolia, 236 Viola glabella, 240 macloskey, 240 purpurea, 242 sheltonii, 242 Violaceae, 50, 240-243 263